# EDGAR Filing Document

**Accession Number:** 0001097519
**File Stem:** 0001145549-25-047809
**Filing Date:** 2025-7
**Character Count:** 64521
**Document Hash:** 5de1d7e1213c5815880b8965eddc0a82
**Contains OCR:** False
**Source Format:** 

## Filing Content

## Filing Summary
**0001145549-25-047809.hdr.sgml**: 20250725

**ACCESSION NUMBER**: 0001145549-25-047809

**CONFORMED SUBMISSION TYPE**: NPORT-P

**PUBLIC DOCUMENT COUNT**: 3

**CONFORMED PERIOD OF REPORT**: 20250531

**FILED AS OF DATE**: 20250725

**DATE AS OF CHANGE**: 20250725

**PERIOD START**: 20260228

**FILER**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** COLUMBIA FUNDS SERIES TRUST
- **CENTRAL INDEX KEY:** 0001097519

**ORGANIZATION NAME:**
- **EIN:** 000000000
- **STATE OF INCORPORATION:** DE
- **FISCAL YEAR END:** 0331

**FILING VALUES:**
- **FORM TYPE:** NPORT-P
- **SEC ACT:** 1940 Act
- **SEC FILE NUMBER:** 811-09645
- **FILM NUMBER:** 251148808

**BUSINESS ADDRESS:**
- **STREET 1:** 290 CONGRESS STREET
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02210
- **BUSINESS PHONE:** 800-345-6611

**MAIL ADDRESS:**
- **STREET 1:** 290 CONGRESS STREET
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02210

**FORMER COMPANY:**
- **FORMER CONFORMED NAME:** NATIONS FUNDS TRUST
- **DATE OF NAME CHANGE:** 19991022

## Series and Classes Contracts Data

### Columbia Overseas Value Fund (Series ID: S000021046)

| Class ID   | Class Name                                         | Ticker Symbol   |
|:---|:---|:---|
| C000059843 | Columbia Overseas Value Fund Class A               | COAVX           |
| C000059844 | Columbia Overseas Value Fund Class C               | COCVX           |
| C000059845 | Columbia Overseas Value Fund Class R               | COVUX           |
| C000059846 | Columbia Overseas Value Fund Institutional Class   | COSZX           |
| C000159975 | Columbia Overseas Value Fund Institutional 2 Class | COSSX           |
| C000159976 | Columbia Overseas Value Fund Institutional 3 Class | COSYX           |
| C000253709 | Columbia Overseas Value Fund Class S               | COSBX           |

## Nport-Ex

![](img28c52f501.jpg)

Columbia Overseas Value Fund

**First Quarter Report**

May 31, 2025 (Unaudited)

---

| | | |
|:---|:---|:---|
| **Not FDIC or NCUA Insured** | **No Financial Institution Guarantee** | **May Lose Value** |

---

------

Portfolio of Investments

Columbia Overseas Value Fund, May 31, 2025 (Unaudited)

(Percentages represent value of investments compared to net assets)

**Investments in securities**

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Common Stocks 96.4%** | **Common Stocks 96.4%** | **Common Stocks 96.4%** |
| **Issuer** | **Shares** | **Value ($)** |
| **Australia 2.4%** | **Australia 2.4%** | **Australia 2.4%** |
| Northern Star Resources Ltd. | &nbsp;&nbsp; 5549965 | &nbsp;&nbsp; 75472420 |
| Paladin Energy Ltd.<sup>(a)</sup> <br>| &nbsp;&nbsp; 5339166 | &nbsp;&nbsp; 21471619 |
| **Total** | **Total** | &nbsp;&nbsp; **96944039** |
| **Brazil 2.1%** | **Brazil 2.1%** | **Brazil 2.1%** |
| Banco do Brasil SA | &nbsp;&nbsp; 11986817 | &nbsp;&nbsp; 49083181 |
| JBS S/A | &nbsp;&nbsp; 5018356 | &nbsp;&nbsp; 35184208 |
| **Total** | **Total** | &nbsp;&nbsp; **84267389** |
| **Canada 3.1%** | **Canada 3.1%** | **Canada 3.1%** |
| Cameco Corp.<sup>(b)</sup> <br>| &nbsp;&nbsp; 1017601 | &nbsp;&nbsp; 59560186 |
| Pan American Silver Corp.<sup>(b)</sup> <br>| &nbsp;&nbsp; 1567084 | &nbsp;&nbsp; 38221179 |
| Teck Resources Ltd., Class B<sup>(b)</sup> <br>| &nbsp;&nbsp; 223744 | &nbsp;&nbsp; 8289715 |
| Vermilion Energy, Inc. | &nbsp;&nbsp; 3383353 | &nbsp;&nbsp; 22025628 |
| **Total** | **Total** | &nbsp;&nbsp; **128096708** |
| **Finland 1.1%** | **Finland 1.1%** | **Finland 1.1%** |
| UPM-Kymmene OYJ | &nbsp;&nbsp; 1706796 | &nbsp;&nbsp; 47239410 |
| **France 12.8%** | **France 12.8%** | **France 12.8%** |
| AXA SA | &nbsp;&nbsp; 2192035 | &nbsp;&nbsp; 103322830 |
| BNP Paribas SA | &nbsp;&nbsp; 883541 | &nbsp;&nbsp; 77375664 |
| Cie de Saint-Gobain SA | &nbsp;&nbsp; 177623 | &nbsp;&nbsp; 19997501 |
| Eiffage SA | &nbsp;&nbsp; 256407 | &nbsp;&nbsp; 35267706 |
| Engie SA | &nbsp;&nbsp; 4547991 | &nbsp;&nbsp; 98159348 |
| Sanofi SA | &nbsp;&nbsp; 827807 | &nbsp;&nbsp; 81986795 |
| TotalEnergies SE | &nbsp;&nbsp; 1850408 | &nbsp;&nbsp; 108930117 |
| **Total** | **Total** | &nbsp;&nbsp; **525039961** |
| **Germany 4.7%** | **Germany 4.7%** | **Germany 4.7%** |
| Duerr AG | &nbsp;&nbsp; 760394 | &nbsp;&nbsp; 19801626 |
| E.ON SE | &nbsp;&nbsp; 4548080 | &nbsp;&nbsp; 79710126 |
| Fresenius Medical Care AG | &nbsp;&nbsp; 913248 | &nbsp;&nbsp; 51876700 |
| KION Group AG | &nbsp;&nbsp; 533276 | &nbsp;&nbsp; 24828015 |
| Merck KGaA | &nbsp;&nbsp; 119587 | &nbsp;&nbsp; 15670546 |
| **Total** | **Total** | &nbsp;&nbsp; **191887013** |
| **Greece 2.6%** | **Greece 2.6%** | **Greece 2.6%** |
| Piraeus Financial Holdings SA | &nbsp;&nbsp; 12925906 | &nbsp;&nbsp; 86110753 |
| Star Bulk Carriers Corp. | &nbsp;&nbsp; 1133518 | &nbsp;&nbsp; 18714382 |
| **Total** | **Total** | &nbsp;&nbsp; **104825135** |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Hong Kong 2.7%** | **Hong Kong 2.7%** | **Hong Kong 2.7%** |
| BOC Hong Kong Holdings Ltd. | &nbsp;&nbsp; 12353500 | &nbsp;&nbsp; 51952811 |
| WH Group Ltd. | &nbsp;&nbsp; 64397330 | &nbsp;&nbsp; 59229931 |
| **Total** | **Total** | &nbsp;&nbsp; **111182742** |
| **Ireland 3.1%** | **Ireland 3.1%** | **Ireland 3.1%** |
| Bank of Ireland Group PLC | &nbsp;&nbsp; 6162999 | &nbsp;&nbsp; 84610768 |
| Flutter Entertainment PLC<sup>(a)</sup> <br>| &nbsp;&nbsp; 71984 | &nbsp;&nbsp; 18190357 |
| Smurfit WestRock PLC | &nbsp;&nbsp; 567507 | &nbsp;&nbsp; 24590078 |
| **Total** | **Total** | &nbsp;&nbsp; **127391203** |
| **Israel 2.2%** | **Israel 2.2%** | **Israel 2.2%** |
| Bezeq Israeli Telecommunication Corp., Ltd. | &nbsp;&nbsp; 4606069 | &nbsp;&nbsp; 7195659 |
| Check Point Software Technologies Ltd.<sup>(a)</sup> <br>| &nbsp;&nbsp; 365378 | &nbsp;&nbsp; 83627717 |
| **Total** | **Total** | &nbsp;&nbsp; **90823376** |
| **Italy 0.5%** | **Italy 0.5%** | **Italy 0.5%** |
| Buzzi SpA | &nbsp;&nbsp; 416337 | &nbsp;&nbsp; 21419424 |
| **Japan 24.1%** | **Japan 24.1%** | **Japan 24.1%** |
| Dai-ichi Life Holdings, Inc. | &nbsp;&nbsp; 6649700 | &nbsp;&nbsp; 51904204 |
| Daiwabo Holdings Co., Ltd. | &nbsp;&nbsp; 2535700 | &nbsp;&nbsp; 42718116 |
| ITOCHU Corp. | &nbsp;&nbsp; 463100 | &nbsp;&nbsp; 24579597 |
| Iyogin Holdings, Inc. | &nbsp;&nbsp; 1675100 | &nbsp;&nbsp; 18247247 |
| Kinden Corp. | &nbsp;&nbsp; 1137600 | &nbsp;&nbsp; 30768725 |
| Macnica Holdings, Inc. | &nbsp;&nbsp; 2573800 | &nbsp;&nbsp; 34021192 |
| Marubeni Corp. | &nbsp;&nbsp; 2399400 | &nbsp;&nbsp; 48634147 |
| MatsukiyoCocokara & Co. | &nbsp;&nbsp; 1814000 | &nbsp;&nbsp; 36584454 |
| Mebuki Financial Group, Inc. | &nbsp;&nbsp; 11994100 | &nbsp;&nbsp; 59095322 |
| Medipal Holdings Corp. | &nbsp;&nbsp; 2132700 | &nbsp;&nbsp; 33353525 |
| Nippon Television Holdings, Inc. | &nbsp;&nbsp; 1567400 | &nbsp;&nbsp; 35470743 |
| ORIX Corp. | &nbsp;&nbsp; 2950400 | &nbsp;&nbsp; 62524441 |
| Sankyo Co., Ltd. | &nbsp;&nbsp; 4394300 | &nbsp;&nbsp; 76641181 |
| Sanwa Holdings Corp. | &nbsp;&nbsp; 874800 | &nbsp;&nbsp; 30116977 |
| Shimamura Co., Ltd. | &nbsp;&nbsp; 938800 | &nbsp;&nbsp; 65056940 |
| Ship Healthcare Holdings, Inc. | &nbsp;&nbsp; 1153500 | &nbsp;&nbsp; 14902927 |
| Subaru Corp. | &nbsp;&nbsp; 2849900 | &nbsp;&nbsp; 52315812 |
| Sumitomo Corp. | &nbsp;&nbsp; 1812800 | &nbsp;&nbsp; 46165803 |
| Sumitomo Mitsui Financial Group, Inc. | &nbsp;&nbsp; 3324000 | &nbsp;&nbsp; 85396681 |
| Sundrug Co., Ltd. | &nbsp;&nbsp; 712700 | &nbsp;&nbsp; 22193447 |

---

Columbia Overseas Value Fund \| 2025

------

Portfolio of Investments (continued)

Columbia Overseas Value Fund, May 31, 2025 (Unaudited)

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| Tokyo Tatemono Co., Ltd. | &nbsp;&nbsp; 2513100 | &nbsp;&nbsp; 44511236 |
| TOPPAN Holdings, Inc. | &nbsp;&nbsp; 2020100 | &nbsp;&nbsp; 53641364 |
| Toyota Motor Corp. | &nbsp;&nbsp; 1087500 | &nbsp;&nbsp; 20704051 |
| **Total** | **Total** | &nbsp;&nbsp; **989548132** |
| **Netherlands 8.9%** | **Netherlands 8.9%** | **Netherlands 8.9%** |
| ASR Nederland NV | &nbsp;&nbsp; 1345823 | &nbsp;&nbsp; 86298572 |
| ING Groep NV | &nbsp;&nbsp; 3193646 | &nbsp;&nbsp; 67839924 |
| Koninklijke Ahold Delhaize NV | &nbsp;&nbsp; 938636 | &nbsp;&nbsp; 39614213 |
| Prosus NV, Class N<sup>(a)</sup> <br>| &nbsp;&nbsp; 819380 | &nbsp;&nbsp; 42090078 |
| Shell PLC | &nbsp;&nbsp; 3991296 | &nbsp;&nbsp; 131692530 |
| **Total** | **Total** | &nbsp;&nbsp; **367535317** |
| **Norway 1.0%** | **Norway 1.0%** | **Norway 1.0%** |
| Leroy Seafood Group ASA | &nbsp;&nbsp; 9459306 | &nbsp;&nbsp; 41307765 |
| **Portugal 2.1%** | **Portugal 2.1%** | **Portugal 2.1%** |
| Banco Comercial Portugues SA | &nbsp;&nbsp; 109622373 | &nbsp;&nbsp; 85511263 |
| **Russian Federation —%** | **Russian Federation —%** | **Russian Federation —%** |
| Lukoil PJSC<sup>(a),(c),(d),(e),(f)</sup> <br>| &nbsp;&nbsp; 106132 | &nbsp;&nbsp; — |
| **Singapore 1.4%** | **Singapore 1.4%** | **Singapore 1.4%** |
| BW LPG Ltd. | &nbsp;&nbsp; 545083 | &nbsp;&nbsp; 5576646 |
| BW LPG Ltd. | &nbsp;&nbsp; 1460136 | &nbsp;&nbsp; 15273023 |
| Venture Corp., Ltd. | &nbsp;&nbsp; 4393300 | &nbsp;&nbsp; 37657613 |
| **Total** | **Total** | &nbsp;&nbsp; **58507282** |
| **South Africa 0.5%** | **South Africa 0.5%** | **South Africa 0.5%** |
| Impala Platinum Holdings Ltd.<sup>(a)</sup> <br>| &nbsp;&nbsp; 2973229 | &nbsp;&nbsp; 21373985 |
| **South Korea 0.5%** | **South Korea 0.5%** | **South Korea 0.5%** |
| Youngone Corp. | &nbsp;&nbsp; 500217 | &nbsp;&nbsp; 21978207 |
| **Spain 2.9%** | **Spain 2.9%** | **Spain 2.9%** |
| Banco Santander SA | &nbsp;&nbsp; 14684631 | &nbsp;&nbsp; 117148915 |
| **Switzerland 1.4%** | **Switzerland 1.4%** | **Switzerland 1.4%** |
| Nestlé SA, Registered Shares | &nbsp;&nbsp; 193979 | &nbsp;&nbsp; 20665968 |
| Novartis AG, Registered Shares | &nbsp;&nbsp; 326277 | &nbsp;&nbsp; 37661868 |
| **Total** | **Total** | &nbsp;&nbsp; **58327836** |
| **United Arab Emirates 1.5%** | **United Arab Emirates 1.5%** | **United Arab Emirates 1.5%** |
| Emaar Properties PJSC | &nbsp;&nbsp; 17746630 | &nbsp;&nbsp; 63549875 |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **United Kingdom 10.8%** | **United Kingdom 10.8%** | **United Kingdom 10.8%** |
| AstraZeneca PLC, ADR | &nbsp;&nbsp; 432871 | &nbsp;&nbsp; 31525995 |
| BT Group PLC | &nbsp;&nbsp; 23441737 | &nbsp;&nbsp; 56756409 |
| DCC PLC | &nbsp;&nbsp; 380740 | &nbsp;&nbsp; 23802371 |
| Imperial Brands PLC | &nbsp;&nbsp; 2464833 | &nbsp;&nbsp; 93478045 |
| JD Sports Fashion PLC | &nbsp;&nbsp; 21405050 | &nbsp;&nbsp; 24276178 |
| Just Group PLC | &nbsp;&nbsp; 23921170 | &nbsp;&nbsp; 48289524 |
| Marks & Spencer Group PLC | &nbsp;&nbsp; 9775189 | &nbsp;&nbsp; 49342761 |
| TP Icap Group PLC | &nbsp;&nbsp; 15301606 | &nbsp;&nbsp; 54084174 |
| Vodafone Group PLC | &nbsp;&nbsp; 29508563 | &nbsp;&nbsp; 30602290 |
| WPP PLC | &nbsp;&nbsp; 3785217 | &nbsp;&nbsp; 30569585 |
| **Total** | **Total** | &nbsp;&nbsp; **442727332** |
| **United States 4.0%** | **United States 4.0%** | **United States 4.0%** |
| Burford Capital Ltd. | &nbsp;&nbsp; 2815496 | &nbsp;&nbsp; 36207279 |
| Diversified Energy Co. PLC | &nbsp;&nbsp; 2644314 | &nbsp;&nbsp; 36941067 |
| Energy Fuels, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 3869972 | &nbsp;&nbsp; 18924163 |
| Insmed, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 111474 | &nbsp;&nbsp; 7773082 |
| Jazz Pharmaceuticals PLC<sup>(a)</sup> <br>| &nbsp;&nbsp; 288458 | &nbsp;&nbsp; 31173656 |
| Roche Holding AG, Genusschein Shares | &nbsp;&nbsp; 103980 | &nbsp;&nbsp; 33684039 |
| **Total** | **Total** | &nbsp;&nbsp; **164703286** |
| &nbsp;&nbsp; Total Common Stocks <br>(Cost $3,171,400,354) | &nbsp;&nbsp; Total Common Stocks <br>(Cost $3,171,400,354) | &nbsp;&nbsp; **3961335595** |
| **Exchange-Traded Equity Funds 2.4%** | **Exchange-Traded Equity Funds 2.4%** | **Exchange-Traded Equity Funds 2.4%** |
|  | **Shares** | **Value ($)** |
| **United States 2.4%** | **United States 2.4%** | **United States 2.4%** |
| iShares MSCI EAFE Value ETF | &nbsp;&nbsp; 1533063 | &nbsp;&nbsp; 97564130 |
| &nbsp;&nbsp; Total Exchange-Traded Equity Funds <br>(Cost $88,720,935) | &nbsp;&nbsp; Total Exchange-Traded Equity Funds <br>(Cost $88,720,935) | &nbsp;&nbsp; **97564130** |
| **Money Market Funds 0.5%** | **Money Market Funds 0.5%** | **Money Market Funds 0.5%** |
| Columbia Short-Term Cash Fund, 4.495%<sup>(g),(h)</sup> <br>| &nbsp;&nbsp; 19355677 | &nbsp;&nbsp; 19349870 |
| &nbsp;&nbsp; Total Money Market Funds <br>(Cost $19,348,568) | &nbsp;&nbsp; Total Money Market Funds <br>(Cost $19,348,568) | &nbsp;&nbsp; **19349870** |
| **Total Investments in Securities** <br>**(Cost $3,279,469,857)** | **Total Investments in Securities** <br>**(Cost $3,279,469,857)** | &nbsp;&nbsp; **4078249595** |
| **Other Assets & Liabilities, Net** |  | &nbsp;&nbsp; **30406185** |
| **Net Assets** | **Net Assets** | &nbsp;&nbsp; **$4108655780** |

---

At May 31, 2025, securities and/or cash totaling $55,955,926 were pledged as collateral.

Columbia Overseas Value Fund \| 2025

------

Portfolio of Investments (continued)

Columbia Overseas Value Fund, May 31, 2025 (Unaudited)

**Investments in derivatives** 

---

| | | | | | |
|:---|:---|:---|:---|:---|:---|
| **Forward foreign currency exchange contracts** | **Forward foreign currency exchange contracts** | **Forward foreign currency exchange contracts** | **Forward foreign currency exchange contracts** | **Forward foreign currency exchange contracts** | **Forward foreign currency exchange contracts** |
| **Currency to** <br>**be sold**<br>| &nbsp;&nbsp;&nbsp; **Currency to** <br>**be purchased**<br>| **Counterparty** | **Settlement** <br>**date**<br>| &nbsp;&nbsp;&nbsp; **Unrealized** <br>**appreciation ($)**<br>| &nbsp;&nbsp;&nbsp; **Unrealized** <br>**depreciation ($)**<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 78,654,000 NOK | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 7,591,645 USD | Barclays | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (113876)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 14,475,640 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 94,934,000 DKK | Barclays | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (17798)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 61,527,518 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 46,497,000 GBP | Barclays | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 1122571 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| 14,376,153,000 JPY | &nbsp;&nbsp;&nbsp;&nbsp; 101,379,876 USD | Citi | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 1382029 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 7,597,909 USD | &nbsp;&nbsp;&nbsp;&nbsp; 1,089,544,000 JPY | Citi | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (19244)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp; 128,009,627 USD | &nbsp;&nbsp;&nbsp;&nbsp; 1,236,605,000 SEK | Citi | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 1040704 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 8,138,041 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 6,693,000 CHF | Goldman Sachs International | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 2442 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp; 108,111,031 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 87,984,000 CHF | Goldman Sachs International | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (1098896)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 25,486,078 USD | &nbsp;&nbsp;&nbsp;&nbsp; 35,876,752,000 KRW | Goldman Sachs International | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 522737 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp; 119,380,978 USD | &nbsp;&nbsp;&nbsp;&nbsp; 187,640,000 AUD | HSBC | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 1585777 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp; 200,960,000 CAD | &nbsp;&nbsp;&nbsp;&nbsp; 144,986,582 USD | Morgan Stanley | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (1507070)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 8,208,332 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 6,784,000 CHF | Morgan Stanley | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 42831 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 7,608,028 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 6,244,000 CHF | Morgan Stanley | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (13650)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp; 293,026,000 SEK | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 30,418,029 USD | State Street | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (161745)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 52,225,000 SGD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 39,896,033 USD | State Street | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (616693)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 14,490,757 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 12,719,000 EUR | State Street | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; — | &nbsp;&nbsp;&nbsp;&nbsp; (41540)<br>|
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 11,800,908 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 8,892,000 GBP | State Street | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 180179 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 11,382,380 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 17,698,000 AUD | UBS | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 27072 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 26,274,979 USD | &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 23,178,000 EUR | UBS | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 56017 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| &nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp; 36,475,372 USD | &nbsp;&nbsp;&nbsp;&nbsp; 383,838,000 NOK | UBS | 06/11/2025 | &nbsp;&nbsp;&nbsp;&nbsp; 1128209 | &nbsp;&nbsp;&nbsp;&nbsp; — |
| Total |  |  |  | &nbsp;&nbsp;&nbsp;&nbsp; 7090568 | &nbsp;&nbsp;&nbsp;&nbsp; (3590512)<br>|

---

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Call option contracts written** | **Call option contracts written** | **Call option contracts written** | **Call option contracts written** | **Call option contracts written** | **Call option contracts written** | **Call option contracts written** | **Call option contracts written** | **Call option contracts written** |
| **Description** | **Counterparty** | &nbsp;&nbsp; **Trading** <br>**currency**<br>| **Notional** <br>**amount**<br>| **Number of** <br>**contracts**<br>| **Exercise** <br>**price/Rate**<br>| **Expiration** <br>**date**<br>| **Premium** <br>**received ($)**<br>| **Value ($)** |
| Cameco Corp. | Morgan Stanley | USD | &nbsp;&nbsp; (23821710)<br>| &nbsp;&nbsp; (4070)<br>| 70.00 | &nbsp;&nbsp; 06/20/2025 | &nbsp;&nbsp; (258284)<br>| &nbsp;&nbsp; (56980)<br>|
| Cameco Corp. | Morgan Stanley | USD | &nbsp;&nbsp; (11910855)<br>| &nbsp;&nbsp; (2035)<br>| 65.00 | &nbsp;&nbsp; 06/20/2025 | &nbsp;&nbsp; (239073)<br>| &nbsp;&nbsp; (96663)<br>|
| Total |  |  |  |  |  |  | &nbsp;&nbsp; (497357)<br>| &nbsp;&nbsp; (153643)<br>|

---

**Notes to Portfolio of Investments** 

&nbsp;&nbsp;&nbsp;&nbsp;(a) Non-income producing investment.

&nbsp;&nbsp;&nbsp;&nbsp;(b) This security or a portion of this security has been pledged as collateral in connection
 with derivative contracts.

&nbsp;&nbsp;&nbsp;&nbsp;(c) Represents fair value as determined in good faith under procedures approved by the
 Board of Trustees. At May 31, 2025, the total value of these securities amounted to $0, which represents less than 0.01% of total net assets.

&nbsp;&nbsp;&nbsp;&nbsp;(d) Denotes a restricted security, which is subject to legal or contractual restrictions
 on resale under federal securities laws. Disposal of a restricted investment may involve time-consuming negotiations and expenses, and prompt sale at an acceptable
 price may be difficult to achieve. Private placement securities are generally considered to be restricted, although certain of those securities may be traded between
 qualified institutional investors under the provisions of Section 4(a)(2) and Rule 144A. The Fund will not incur any registration costs upon such a trade. These
 securities are valued at fair value determined in good faith under consistently applied procedures approved by the Fund's Board of Trustees. At May 31, 2025, the total market value of these securities amounted to $0, which represents less than 0.01% of total net assets. Additional information on these securities is as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | |
|:---|:---|:---|:---|:---|
| **Security** | **Acquisition** <br>**Dates**<br>| **Shares** | **Cost ($)** | **Value ($)** |
| Lukoil PJSC | 01/25/2022-01/26/2022 | &nbsp;&nbsp; 106132 | &nbsp;&nbsp; 8693218 | &nbsp;&nbsp; — |

---

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;(e) As a result of sanctions and restricted cross-border payments, certain payments have
 not been recognized by the Fund. The Fund will continue to monitor the net realizable value and record payments when it is considered collectible.

&nbsp;&nbsp;&nbsp;&nbsp;(f) Valuation based on significant unobservable inputs.

&nbsp;&nbsp;&nbsp;&nbsp;(g) The rate shown is the seven-day current annualized yield at May 31, 2025.

Columbia Overseas Value Fund \| 2025

------

Portfolio of Investments (continued)

Columbia Overseas Value Fund, May 31, 2025 (Unaudited)

**Notes to Portfolio of Investments (continued)**

&nbsp;&nbsp;&nbsp;&nbsp;(h) Under Section 2(a)(3) of the Investment Company Act of 1940, an affiliated company is one in which the Fund owns 5% or more of the company's outstanding voting securities, or a company which is under common ownership or control with the Fund.
 The value of the holdings and transactions in these affiliated companies during the period ended May 31, 2025 are as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Affiliated issuers** | **Beginning** <br>**of period($)**<br>| **Purchases($)** | **Sales($)** | **Net change in** <br>**unrealized** <br>**appreciation** <br>**(depreciation)($)**<br>| **End of** <br>**period($)**<br>| **Realized gain** <br>**(loss)($)**<br>| **Dividends($)** | **End of** <br>**period shares**<br>|
| Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% | Columbia Short-Term Cash Fund, 4.495% |
|  | 6022145 | &nbsp;&nbsp; 602429730 | &nbsp;&nbsp; (589102694)<br>| &nbsp;&nbsp; 689 | &nbsp;&nbsp; 19349870 | &nbsp;&nbsp; (13602)<br>| &nbsp;&nbsp; 418280 | &nbsp;&nbsp; 19355677 |

---

**Abbreviation Legend** 

ADR American Depositary Receipt

**Currency Legend** 

---

| | |
|:---|:---|
| AUD | Australian Dollar |
| CAD | Canada Dollar |
| CHF | Swiss Franc |
| DKK | Danish Krone |
| EUR | Euro |
| GBP | British Pound |
| JPY | Japanese Yen |
| KRW | South Korean Won |
| NOK | Norwegian Krone |
| SEK | Swedish Krona |
| SGD | Singapore Dollar |
| USD | US Dollar |

---

Investments are valued using policies described in the Notes to Financial Statements in the most recent shareholder report.

Columbia Overseas Value Fund \| 2025

------

You may at any time request, free of charge, to receive a paper copy of this report by calling 800.345.6611.

1QT208_02_R01_(07/25)

------

## Form NPORT-P: Monthly Portfolio Investments Report

### NPORT-P: Part A: General Information

**Item A.1. Information about the Registrant.**

- **a. Name of Registrant:** Columbia Funds Series Trust

- **b. Investment Company Act file number:** 811-09645

- **c. CIK number of Registrant:** 0001097519

- **d. LEI of Registrant:** 549300712V4UL3XCAB43

- **e. Address and telephone number of Registrant.**

  - **Street Address 1:** 290 Congress Street

  - **City:** Boston

  - **State:** MA

  - **Foreign country:** US

  - **Zip / Postal Code:** 02110

  - **Telephone number:** 800-345-6611

**Item A.2. Information about the Series.**

- **a. Name of Series:** Columbia Overseas Value Fund

- **b. EDGAR series identifier (if any):** S000021046

- **c. LEI of Series:** UO5UKRJ52BO37ERP8Y54

**Item A.3. Reporting period.**

- **a. Date of fiscal year-end:** 2026-02-28

- **b. Date as of which information is reported:** 2025-05-31

**Item A.4. Final filing**

Does the Fund anticipate that this will be its final filing on Form N-PORT? **No**

### Fund Information

**Total Assets:** $4116424538.55

**Total Liabilities:** $10730369.00

**Net Assets:** $4105694169.55

**Cash Not Reported:** $0.08

**Monthly Return Information**

| Class               | Month 1 Return (%)   | Month 2 Return (%)   | Month 3 Return (%)   |
|:---|:---|:---|:---|
| Class ID C000059843 | 4.40%                | 3.04%                | 4.58%                |
| Class ID C000059844 | 4.36%                | 2.98%                | 4.47%                |
| Class ID C000059845 | 4.35%                | 3.04%                | 4.55%                |
| Class ID C000059846 | 4.48%                | 3.03%                | 4.57%                |
| Class ID C000159975 | 4.42%                | 3.05%                | 4.60%                |
| Class ID C000159976 | 4.41%                | 3.04%                | 4.59%                |
| Class ID C000253709 | 4.48%                | 3.03%                | 4.57%                |

**Monthly Gains & Losses**

| Period   | Net Realized Gain/Loss   | Net Unrealized Appreciation/Depreciation   |
|:---|:---|:---|
| Month 1  | $31586588.11             | $74796032.76                               |
| Month 2  | $-814057.74              | $91370268.19                               |
| Month 3  | $15031378.52             | $128216838.88                              |

**Designated Index Information**

- **Index Name:** MSCI EAFE Value Open USD NET TR Index

- **Index Identifier:** MSCIEAFE

### Schedule of Portfolio Investments

| Name                                     | Title                                        | Identifiers                                   | Payoff Profile   | Asset Category   | Issuer Category   | Country   |      Balance | Units   | Value (USD)   | % of Net Assets   | Maturity Date   | Coupon Type   | Annualized Rate (%)   | Restricted?   |   Fair Value Level | Lending Status   |
|:---|:---|:---|:---|:---|:---|:---|---:|:---|:---|:---|:---|:---|:---|:---|---:|:---|
| DIVERSIFIED ENERGY COMPANY PLC           | Diversified Energy Co PLC                    | CUSIP: G2891G204<br>LEI: 213800YR9TFRVHPGOS67 | Long             | EC               | CORP              | GB        |      2644314 | NS      | $36941066.58  | 0.90%             |  |  |  | No            |                  1 | On Loan: No      |
| FLUTTER ENTERTAINMENT PLC                | Flutter Entertainment PLC                    | CUSIP: G3643J108<br>LEI: 635400EG4YIJLJMZJ782 | Long             | EC               | CORP              | IE        |        71984 | NS      | $18190356.80  | 0.44%             |  |  |  | No            |                  1 | On Loan: No      |
| JAZZ PHARMACEUTICALS PLC                 | Jazz Pharmaceuticals PLC                     | CUSIP: G50871105<br>LEI: 635400GAUMJCLEZRRV50 | Long             | EC               | CORP              | IE        |       288458 | NS      | $31173656.06  | 0.76%             |  |  |  | No            |                  1 | On Loan: No      |
| SMURFIT WESTROCK PLC                     | Smurfit WestRock PLC                         | CUSIP: G8267P108<br>LEI: 635400EGSEOPDW2EXS14 | Long             | EC               | CORP              | IE        |       567507 | NS      | $24590078.31  | 0.60%             |  |  |  | No            |                  1 | On Loan: No      |
| Goldman Sachs International              | SWISS FRANC                                  | CUSIP: 000000000<br>LEI: W22LROWP2IHZNBB6K528 | N/A              | DFE              | CORP              | CH        |      6693000 | OU      | $2441.54      | 0.00%             |  |  |  | No            |                  2 | On Loan: No      |
| Goldman Sachs International              | SWISS FRANC                                  | CUSIP: 000000000<br>LEI: W22LROWP2IHZNBB6K528 | N/A              | DFE              | CORP              | CH        |     87984000 | OU      | $-1098895.87  | -0.03%            |  |  |  | No            |                  2 | On Loan: No      |
| Goldman Sachs International              | KOREAN WON                                   | CUSIP: 000000000<br>LEI: W22LROWP2IHZNBB6K528 | N/A              | DFE              | CORP              | KW        |  35876752000 | OU      | $522737.49    | 0.01%             |  |  |  | No            |                  2 | On Loan: No      |
| HSBC                                     | AUSTRALIA DOLLAR                             | CUSIP: 000000000<br>LEI: CYYGQCGNHMHPSMRL3R97 | N/A              | DFE              | CORP              | AU        |    187640000 | OU      | $1585776.98   | 0.04%             |  |  |  | No            |                  2 | On Loan: No      |
| CHECK POINT SOFTWARE TECH LTD.           | Check Point Software Technologies Ltd        | CUSIP: M22465104<br>LEI: 2549003ZVBH73EPNS513 | Long             | EC               | CORP              | IL        |       365378 | NS      | $83627716.64  | 2.04%             |  |  |  | No            |                  1 | On Loan: No      |
| Morgan Stanley                           | US DOLLARS                                   | CUSIP: 000000000<br>LEI: 9R7GPTSO7KV3UQJZQ078 | N/A              | DFE              | CORP              | CA        |   -200960000 | OU      | $-1507069.71  | -0.04%            |  |  |  | No            |                  2 | On Loan: No      |
| Morgan Stanley                           | SWISS FRANC                                  | CUSIP: 000000000<br>LEI: 9R7GPTSO7KV3UQJZQ078 | N/A              | DFE              | CORP              | CH        |      6784000 | OU      | $42831.04     | 0.00%             |  |  |  | No            |                  2 | On Loan: No      |
| DCC PLC                                  | DCC PLC                                      | CUSIP: 000000000<br>LEI: 213800UPS1QEE4TD3T45 | Long             | EC               | CORP              | IE        |       380740 | NS      | $23802371.03  | 0.58%             |  |  |  | No            |                  2 | On Loan: No      |
| ASTRAZENECA PLC                          | AstraZeneca PLC                              | CUSIP: 046353108<br>LEI: PY6ZZQWO2IZFZC3IOL08 | Long             | EC               | CORP              | GB        |       316648 | NS      | $23061473.84  | 0.56%             |  |  |  | No            |                  1 | On Loan: No      |
| AXA SA                                   | AXA SA                                       | CUSIP: 000000000<br>LEI: F5WCUMTUM4RKZ1MAIE39 | Long             | EC               | CORP              | FR        |      2192035 | NS      | $103322829.69 | 2.52%             |  |  |  | No            |                  2 | On Loan: No      |
| BOC HONG KONG HOLDINGS LTD               | BOC Hong Kong Holdings Ltd                   | CUSIP: 000000000<br>LEI: 254900JB98W77JDJMW15 | Long             | EC               | CORP              | HK        |     12353500 | NS      | $51952811.17  | 1.27%             |  |  |  | No            |                  2 | On Loan: No      |
| Barclays                                 | DANISH KRONA                                 | CUSIP: 000000000<br>LEI: AC28XWWI3WIBK2824319 | N/A              | DFE              | CORP              | DK        |     94934000 | OU      | $-17797.89    | -0.00%            |  |  |  | No            |                  2 | On Loan: No      |
| Barclays                                 | BRITISH POUND                                | CUSIP: 000000000<br>LEI: AC28XWWI3WIBK2824319 | N/A              | DFE              | CORP              | GB        |     46497000 | OU      | $1122570.88   | 0.03%             |  |  |  | No            |                  2 | On Loan: No      |
| JUST GROUP PLC                           | Just Group PLC                               | CUSIP: 000000000<br>LEI: 5493006456YEZEELRR90 | Long             | EC               | CORP              | GB        |     23921170 | NS      | $48289524.39  | 1.18%             |  |  |  | No            |                  2 | On Loan: No      |
| KONINKLIJKE AHOLD DELHAIZE NV            | Koninklijke Ahold Delhaize NV                | CUSIP: 000000000<br>LEI: 724500C9GNBV20UYRX36 | Long             | EC               | CORP              | NL        |       938636 | NS      | $39614212.73  | 0.96%             |  |  |  | No            |                  2 | On Loan: No      |
| BANK OF IRELAND GROUP PLC                | Bank of Ireland Group PLC                    | CUSIP: 000000000<br>LEI: 635400C8EK6DRI12LJ39 | Long             | EC               | CORP              | IE        |      6162999 | NS      | $84610768.12  | 2.06%             |  |  |  | No            |                  2 | On Loan: No      |
| ASR NEDERLAND NV                         | ASR Nederland NV                             | CUSIP: 000000000<br>LEI: 7245000G0HS48PZWUD53 | Long             | EC               | CORP              | NL        |      1345823 | NS      | $86298572.24  | 2.10%             |  |  |  | No            |                  2 | On Loan: No      |
| VERMILION ENERGY INC                     | Vermilion Energy Inc                         | CUSIP: 923725105<br>LEI: WT03B8BB1IX8WI9ZGV02 | Long             | EC               | CORP              | CA        |      3383353 | NS      | $22025628.03  | 0.54%             |  |  |  | No            |                  1 | On Loan: No      |
| VODAFONE GROUP PLC                       | Vodafone Group PLC                           | CUSIP: 000000000<br>LEI: 213800TB53ELEUKM7Q61 | Long             | EC               | CORP              | GB        |     29508563 | NS      | $30602289.82  | 0.75%             |  |  |  | No            |                  2 | On Loan: No      |
| WPP PLC                                  | WPP PLC                                      | CUSIP: 000000000<br>LEI: 549300LSGBXPYHXGDT93 | Long             | EC               | CORP              | JE        |      3785217 | NS      | $30569584.88  | 0.74%             |  |  |  | No            |                  2 | On Loan: No      |
| YOUNGONE CORP                            | Youngone Corp                                | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | KR        |       500217 | NS      | $21978207.13  | 0.54%             |  |  |  | No            |                  2 | On Loan: No      |
| EMAAR PROPERTIES PJSC                    | Emaar Properties PJSC                        | CUSIP: 000000000<br>LEI: 254900YWYEXYXK1BMP81 | Long             | EC               | CORP              | AE        |     17746630 | NS      | $63549875.15  | 1.55%             |  |  |  | No            |                  2 | On Loan: No      |
| SHIP HEALTHCARE HOLDINGS INC             | Ship Healthcare Holdings Inc                 | CUSIP: 000000000<br>LEI: 353800BIFUE1L527TH13 | Long             | EC               | CORP              | JP        |      1225100 | NS      | $15827980.98  | 0.39%             |  |  |  | No            |                  2 | On Loan: No      |
| NK LUKOIL PAO                            | LUKOIL PJSC                                  | CUSIP: 000000000<br>LEI: 549300LCJ1UJXHYBWI24 | Long             | EC               | CORP              | RU        |       106132 | NS      | $0.00         | 0.00%             |  |  |  | Yes           |                  3 | On Loan: No      |
| DAI-ICHI LIFE HOLDINGS INC               | Dai-ichi Life Holdings Inc                   | CUSIP: 000000000<br>LEI: 549300ZUFXI7JXZVTZ25 | Long             | EC               | CORP              | JP        |      6649700 | NS      | $51904203.98  | 1.26%             |  |  |  | No            |                  2 | On Loan: No      |
| KION GROUP AG                            | KION Group AG                                | CUSIP: 000000000<br>LEI: 5299005KY91C4C6U9H17 | Long             | EC               | CORP              | DE        |       533276 | NS      | $24828015.02  | 0.60%             |  |  |  | No            |                  2 | On Loan: No      |
| Barclays                                 | US DOLLARS                                   | CUSIP: 000000000<br>LEI: AC28XWWI3WIBK2824319 | N/A              | DFE              | CORP              | NO        |    -78654000 | OU      | $-113876.47   | -0.00%            |  |  |  | No            |                  2 | On Loan: No      |
| MEBUKI FINANCIAL GROUP INC               | Mebuki Financial Group Inc                   | CUSIP: 000000000<br>LEI: 353800K1RXV3E3JAE812 | Long             | EC               | CORP              | JP        |     11994100 | NS      | $59095322.38  | 1.44%             |  |  |  | No            |                  2 | On Loan: No      |
| PROSUS NV                                | Prosus NV                                    | CUSIP: 000000000<br>LEI: 635400Z5LQ5F9OLVT688 | Long             | EC               | CORP              | NL        |       819380 | NS      | $42090077.34  | 1.03%             |  |  |  | No            |                  2 | On Loan: No      |
| WH GROUP LTD                             | WH Group Ltd                                 | CUSIP: 000000000<br>LEI: 254900A4AYEDRWYSE928 | Long             | EC               | CORP              | KY        |     64397330 | NS      | $59229931.08  | 1.44%             |  |  |  | No            |                  2 | On Loan: No      |
| JD SPORTS FASHION PLC                    | JD Sports Fashion PLC                        | CUSIP: 000000000<br>LEI: 213800HROV6Y9MUU8375 | Long             | EC               | CORP              | GB        |     21405050 | NS      | $24276178.07  | 0.59%             |  |  |  | No            |                  2 | On Loan: No      |
| TP ICAP GROUP PLC                        | TP ICAP Group PLC                            | CUSIP: 000000000<br>LEI: 2138006YAA7IRVKKGE63 | Long             | EC               | CORP              | JE        |     15301606 | NS      | $54084174.12  | 1.32%             |  |  |  | No            |                  2 | On Loan: No      |
| PIRAEUS FINANCIAL HOLDINGS SA            | Piraeus Financial Holdings SA                | CUSIP: 000000000<br>LEI: M6AD1Y1KW32H8THQ6F76 | Long             | EC               | CORP              | GR        |     12925906 | NS      | $86110752.41  | 2.10%             |  |  |  | No            |                  2 | On Loan: No      |
| IYOGIN HOLDINGS INC                      | Iyogin Holdings Inc                          | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | JP        |      1675100 | NS      | $18247247.14  | 0.44%             |  |  |  | No            |                  2 | On Loan: No      |
| SHELL PLC                                | Shell PLC                                    | CUSIP: 000000000<br>LEI: 21380068P1DRHMJ8KU70 | Long             | EC               | CORP              | GB        |      3991296 | NS      | $131692529.93 | 3.21%             |  |  |  | No            |                  2 | On Loan: No      |
| BW LPG LTD                               | BW LPG Ltd                                   | CUSIP: 000000000<br>LEI: 5493006WBEME88YFDW23 | Long             | EC               | CORP              | SG        |       545083 | NS      | $5576645.79   | 0.14%             |  |  |  | No            |                  2 | On Loan: No      |
| MACNICA HOLDINGS INC                     | Macnica Holdings Inc                         | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | JP        |      2573800 | NS      | $34021191.51  | 0.83%             |  |  |  | No            |                  2 | On Loan: No      |
| ING GROEP NV                             | ING Groep NV                                 | CUSIP: 000000000<br>LEI: 549300NYKK9MWM7GGW15 | Long             | EC               | CORP              | NL        |      3193646 | NS      | $67839924.30  | 1.65%             |  |  |  | No            |                  2 | On Loan: No      |
| Citi                                     | US DOLLARS                                   | CUSIP: 000000000<br>LEI: MBNUM2BPBDO7JBLYG310 | N/A              | DFE              | CORP              | JP        | -14376153000 | OU      | $1382029.42   | 0.03%             |  |  |  | No            |                  2 | On Loan: No      |
| Citi                                     | JAPANESE YEN                                 | CUSIP: 000000000<br>LEI: MBNUM2BPBDO7JBLYG310 | N/A              | DFE              | CORP              | JP        |   1089544000 | OU      | $-19243.94    | -0.00%            |  |  |  | No            |                  2 | On Loan: No      |
| Citi                                     | SWEDISH KRONA                                | CUSIP: 000000000<br>LEI: MBNUM2BPBDO7JBLYG310 | N/A              | DFE              | CORP              | SE        |   1236605000 | OU      | $1040703.61   | 0.03%             |  |  |  | No            |                  2 | On Loan: No      |
| BURFORD CAPITAL LTD                      | Burford Capital Ltd                          | CUSIP: G17977110<br>LEI: 549300FUKUWFYJMT2277 | Long             | EC               | CORP              | GG        |      2815496 | NS      | $36207278.56  | 0.88%             |  |  |  | No            |                  1 | On Loan: No      |
| BT GROUP PLC                             | BT Group PLC                                 | CUSIP: 000000000<br>LEI: 213800LRO7NS5CYQMN21 | Long             | EC               | CORP              | GB        |     23441737 | NS      | $56756408.98  | 1.38%             |  |  |  | No            |                  2 | On Loan: No      |
| BANCO DO BRASIL SA                       | Banco do Brasil SA                           | CUSIP: 000000000<br>LEI: QE0Q0253K88YAGNPE356 | Long             | EC               | NUSS              | BR        |     11986817 | NS      | $49083181.07  | 1.20%             |  |  |  | No            |                  1 | On Loan: No      |
| BANCO COMERCIAL PORTUGUES SA             | Banco Comercial Portugues SA                 | CUSIP: 000000000<br>LEI: JU1U6S0DG9YLT7N8ZV32 | Long             | EC               | CORP              | PT        |    109622373 | NS      | $85511262.55  | 2.08%             |  |  |  | No            |                  2 | On Loan: No      |
| BANCO SANTANDER SA                       | Banco Santander SA                           | CUSIP: 000000000<br>LEI: 5493006QMFDDMYWIAM13 | Long             | EC               | CORP              | ES        |     16735130 | NS      | $133507088.01 | 3.25%             |  |  |  | No            |                  2 | On Loan: No      |
| BNP PARIBAS SA                           | BNP Paribas SA                               | CUSIP: 000000000<br>LEI: R0MUWSFPU8MPRO8K5P83 | Long             | EC               | CORP              | FR        |       883541 | NS      | $77375663.78  | 1.88%             |  |  |  | No            |                  2 | On Loan: No      |
| BEZEQ ISRAELI TELECOMMUNICATION CORP LTD | Bezeq The Israeli Telecommunication Corp Ltd | CUSIP: 000000000<br>LEI: 213800XCGZCFSWY9U863 | Long             | EC               | CORP              | IL        |      4606069 | NS      | $7195659.07   | 0.18%             |  |  |  | No            |                  2 | On Loan: No      |
| COMPAGNIE DE SAINT GOBAIN SA             | Cie de Saint-Gobain SA                       | CUSIP: 000000000<br>LEI: NFONVGN05Z0FMN5PEC35 | Long             | EC               | CORP              | FR        |       177623 | NS      | $19997501.68  | 0.49%             |  |  |  | No            |                  2 | On Loan: No      |
| CAMECO CORP                              | Cameco Corp                                  | CUSIP: 13321L108<br>LEI: N4BHALMXL3GMZKCUON55 | Long             | EC               | CORP              | CA        |      1017601 | NS      | $59560186.53  | 1.45%             |  |  |  | No            |                  1 | On Loan: No      |
| TOTALENERGIES SE                         | TotalEnergies SE                             | CUSIP: 000000000<br>LEI: 529900S21EQ1BO4ESM68 | Long             | EC               | CORP              | FR        |      1850408 | NS      | $108930116.95 | 2.65%             |  |  |  | No            |                  2 | On Loan: No      |
| TOYOTA MTR CORP                          | Toyota Motor Corp                            | CUSIP: 000000000<br>LEI: 5493006W3QUS5LMH6R84 | Long             | EC               | CORP              | JP        |      1087500 | NS      | $20704051.28  | 0.50%             |  |  |  | No            |                  2 | On Loan: No      |
| UPM-KYMMENE OYJ                          | UPM-Kymmene Oyj                              | CUSIP: 000000000<br>LEI: 213800EC6PW5VU4J9U64 | Long             | EC               | CORP              | FI        |      1706796 | NS      | $47239409.98  | 1.15%             |  |  |  | No            |                  2 | On Loan: No      |
| VENTURE CORPORATION LTD                  | Venture Corp Ltd                             | CUSIP: 000000000<br>LEI: 254900B1NLZC3J1MAE72 | Long             | EC               | CORP              | SG        |      4393300 | NS      | $37657613.34  | 0.92%             |  |  |  | No            |                  2 | On Loan: No      |
| Morgan Stanley                           | SWISS FRANC                                  | CUSIP: 000000000<br>LEI: 9R7GPTSO7KV3UQJZQ078 | N/A              | DFE              | CORP              | CH        |      6244000 | OU      | $-13649.98    | -0.00%            |  |  |  | No            |                  2 | On Loan: No      |
| State Street                             | US DOLLARS                                   | CUSIP: 000000000<br>LEI: 571474TGEMMWANRLN572 | N/A              | DFE              | CORP              | SE        |   -293026000 | OU      | $-161745.50   | -0.00%            |  |  |  | No            |                  2 | On Loan: No      |
| State Street                             | US DOLLARS                                   | CUSIP: 000000000<br>LEI: 571474TGEMMWANRLN572 | N/A              | DFE              | CORP              | SG        |    -52225000 | OU      | $-616693.45   | -0.02%            |  |  |  | No            |                  2 | On Loan: No      |
| State Street                             | EURO COUNTRIES                               | CUSIP: 000000000<br>LEI: 571474TGEMMWANRLN572 | N/A              | DFE              | CORP              | XX        |     12719000 | OU      | $-41540.25    | -0.00%            |  |  |  | No            |                  2 | On Loan: No      |
| State Street                             | BRITISH POUND                                | CUSIP: 000000000<br>LEI: 571474TGEMMWANRLN572 | N/A              | DFE              | CORP              | GB        |      8892000 | OU      | $180179.19    | 0.00%             |  |  |  | No            |                  2 | On Loan: No      |
| UBS                                      | AUSTRALIA DOLLAR                             | CUSIP: 000000000<br>LEI: T6FIZBDPKLYJKFCRVK44 | N/A              | DFE              | CORP              | AU        |     17698000 | OU      | $27072.05     | 0.00%             |  |  |  | No            |                  2 | On Loan: No      |
| UBS                                      | EURO COUNTRIES                               | CUSIP: 000000000<br>LEI: T6FIZBDPKLYJKFCRVK44 | N/A              | DFE              | CORP              | XX        |     23178000 | OU      | $56017.40     | 0.00%             |  |  |  | No            |                  2 | On Loan: No      |
| UBS                                      | NORWAY KRONA                                 | CUSIP: 000000000<br>LEI: T6FIZBDPKLYJKFCRVK44 | N/A              | DFE              | CORP              | NO        |    383838000 | OU      | $1128208.88   | 0.03%             |  |  |  | No            |                  2 | On Loan: No      |
| BW LPG LTD                               | BW LPG Ltd                                   | CUSIP: Y10230103<br>LEI: 5493006WBEME88YFDW23 | Long             | EC               | CORP              | SG        |      1460136 | NS      | $15273022.56  | 0.37%             |  |  |  | No            |                  1 | On Loan: No      |
| STAR BULK CARRIERS CORP                  | Star Bulk Carriers Corp                      | CUSIP: Y8162K204<br>LEI: 549300SO3FMK7H6DDY54 | Long             | EC               | CORP              | MH        |      1133518 | NS      | $18714382.18  | 0.46%             |  |  |  | No            |                  1 | On Loan: No      |
| Morgan Stanley                           | Cameco Corp                                  | CUSIP: 000000000<br>LEI: 9R7GPTSO7KV3UQJZQ078 | N/A              | DE               | CORP              | CA        |        -2035 | NC      | $-96662.50    | -0.00%            |  |  |  | No            |                  1 | On Loan: No      |
| Morgan Stanley                           | Cameco Corp                                  | CUSIP: 000000000<br>LEI: 9R7GPTSO7KV3UQJZQ078 | N/A              | DE               | CORP              | CA        |        -4070 | NC      | $-56980.00    | -0.00%            |  |  |  | No            |                  1 | On Loan: No      |
| BUZZI SPA                                | Buzzi SpA                                    | CUSIP: 000000000<br>LEI: 5299003DX5YLKSVJ6K59 | Long             | EC               | CORP              | IT        |       416337 | NS      | $21419424.34  | 0.52%             |  |  |  | No            |                  2 | On Loan: No      |
| COLUMBIA SHORT TERM CASH FUND            | COLUMBIA SHORT TERM CASH FUND                | CUSIP: 19766H239<br>LEI: N/A                  | Long             | STIV             | CORP              | US        |     19355677 | NS      | $19349870.15  | 0.47%             |  |  |  | No            |                  1 | On Loan: No      |
| DAIWABO HOLDINGS CO LTD                  | Daiwabo Holdings Co Ltd                      | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | JP        |      2535700 | NS      | $42718115.65  | 1.04%             |  |  |  | No            |                  2 | On Loan: No      |
| DUERR AG                                 | Duerr AG                                     | CUSIP: 000000000<br>LEI: 5299003APZ54EDTSRO20 | Long             | EC               | CORP              | DE        |       760394 | NS      | $19801626.20  | 0.48%             |  |  |  | No            |                  2 | On Loan: No      |
| E.ON SE                                  | E.ON SE                                      | CUSIP: 000000000<br>LEI: Q9MAIUP40P25UFBFG033 | Long             | EC               | CORP              | DE        |      5246907 | NS      | $91957840.37  | 2.24%             |  |  |  | No            |                  2 | On Loan: No      |
| EIFFAGE SA                               | Eiffage SA                                   | CUSIP: 000000000<br>LEI: 969500OQXKE5WDM9M994 | Long             | EC               | CORP              | FR        |       226888 | NS      | $31207491.29  | 0.76%             |  |  |  | No            |                  2 | On Loan: No      |
| ENERGY FUELS INC                         | Energy Fuels Inc/Canada                      | CUSIP: 292671708<br>LEI: 529900P0GCATXMGK8I62 | Long             | EC               | CORP              | CA        |      3869972 | NS      | $18924163.08  | 0.46%             |  |  |  | No            |                  1 | On Loan: No      |
| FRESENIUS MEDICAL CARE AG                | Fresenius Medical Care AG                    | CUSIP: 000000000<br>LEI: 549300CP8NY40UP89Q40 | Long             | EC               | CORP              | DE        |       913248 | NS      | $51876700.08  | 1.26%             |  |  |  | No            |                  2 | On Loan: No      |
| SUBARU CORP                              | Subaru Corp                                  | CUSIP: 000000000<br>LEI: 549300N244BVAEE6HH86 | Long             | EC               | CORP              | JP        |      2849900 | NS      | $52315812.39  | 1.27%             |  |  |  | No            |                  2 | On Loan: No      |
| ENGIE SA                                 | Engie SA                                     | CUSIP: 000000000<br>LEI: LAXUQCHT4FH58LRZDY46 | Long             | EC               | CORP              | FR        |      4547991 | NS      | $98159348.19  | 2.39%             |  |  |  | No            |                  2 | On Loan: No      |
| ROCHE HOLDING AG                         | Roche Holding AG                             | CUSIP: 000000000<br>LEI: 549300U41AUUVOAAOB37 | Long             | EC               | CORP              | CH        |       103980 | NS      | $33684039.51  | 0.82%             |  |  |  | No            |                  2 | On Loan: No      |
| IMPALA PLATINUM HOLDINGS LTD             | Impala Platinum Holdings Ltd                 | CUSIP: 000000000<br>LEI: 254900BZFIFB4PD2Z778 | Long             | EC               | CORP              | ZA        |      2703646 | NS      | $19436003.14  | 0.47%             |  |  |  | No            |                  2 | On Loan: No      |
| IMPERIAL BRANDS PLC                      | Imperial Brands PLC                          | CUSIP: 000000000<br>LEI: 549300DFVPOB67JL3A42 | Long             | EC               | CORP              | GB        |      2464833 | NS      | $93478045.09  | 2.28%             |  |  |  | No            |                  2 | On Loan: No      |
| INSMED                                   | Insmed Inc                                   | CUSIP: 457669307<br>LEI: 529900Q55QAG41CCAC11 | Long             | EC               | CORP              | US        |       111474 | NS      | $7773082.02   | 0.19%             |  |  |  | No            |                  1 | On Loan: No      |
| ISHARES MSCI VALUE INDEX FUND            | iShares MSCI EAFE Value ETF                  | CUSIP: 464288877<br>LEI: 549300AN74D4XJZB8Y40 | Long             | EC               | RF                | US        |      1533063 | NS      | $97564129.32  | 2.38%             |  |  |  | No            |                  1 | On Loan: No      |
| ITOCHU CORPORATION                       | ITOCHU Corp                                  | CUSIP: 000000000<br>LEI: J48DJYXDTLHM30UMYI18 | Long             | EC               | CORP              | JP        |       463100 | NS      | $24579596.69  | 0.60%             |  |  |  | No            |                  2 | On Loan: No      |
| JBS SA                                   | JBS S/A                                      | CUSIP: 000000000<br>LEI: 529900RM0MTMZL9O5050 | Long             | EC               | CORP              | BR        |      5018356 | NS      | $35184207.64  | 0.86%             |  |  |  | No            |                  1 | On Loan: No      |
| KINDEN CORP                              | Kinden Corp                                  | CUSIP: 000000000<br>LEI: 35380008T6JUN5TU9T34 | Long             | EC               | CORP              | JP        |      1137600 | NS      | $30768725.17  | 0.75%             |  |  |  | No            |                  2 | On Loan: No      |
| LEROY SEAFOOD GROUP ASA                  | Leroy Seafood Group ASA                      | CUSIP: 000000000<br>LEI: 5967007LIEEXZXJ2JK50 | Long             | EC               | CORP              | NO        |      9459306 | NS      | $41307765.47  | 1.01%             |  |  |  | No            |                  2 | On Loan: No      |
| MARKS AND SPENCER GROUP PLC              | Marks & Spencer Group PLC                    | CUSIP: 000000000<br>LEI: 213800EJP14A79ZG1X44 | Long             | EC               | CORP              | GB        |      9775189 | NS      | $49342760.69  | 1.20%             |  |  |  | No            |                  2 | On Loan: No      |
| MARUBENI CORP                            | Marubeni Corp                                | CUSIP: 000000000<br>LEI: 4P4N3ORD02UGQT1T1W12 | Long             | EC               | CORP              | JP        |      2399400 | NS      | $48634147.22  | 1.18%             |  |  |  | No            |                  2 | On Loan: No      |
| MATSUKIYOCOCOKARA & CO                   | MatsukiyoCocokara & Co                       | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | JP        |      1814000 | NS      | $36584454.12  | 0.89%             |  |  |  | No            |                  2 | On Loan: No      |
| MEDIPAL HOLDINGS CORP                    | Medipal Holdings Corp                        | CUSIP: 000000000<br>LEI: 353800PJDODZEA4NP730 | Long             | EC               | CORP              | JP        |      2132700 | NS      | $33353524.93  | 0.81%             |  |  |  | No            |                  2 | On Loan: No      |
| MERCK KGAA                               | Merck KGaA                                   | CUSIP: 000000000<br>LEI: 529900OAREIS0MOPTW25 | Long             | EC               | CORP              | DE        |       119587 | NS      | $15670546.19  | 0.38%             |  |  |  | No            |                  2 | On Loan: No      |
| NESTLE SA                                | Nestle SA                                    | CUSIP: 000000000<br>LEI: KY37LUS27QQX7BB93L28 | Long             | EC               | CORP              | CH        |       193979 | NS      | $20665967.83  | 0.50%             |  |  |  | No            |                  2 | On Loan: No      |
| NIPPON TELEVISION HOLDINGS INC.          | Nippon Television Holdings Inc               | CUSIP: 000000000<br>LEI: 353800QI43C3H97E4W63 | Long             | EC               | CORP              | JP        |      1567400 | NS      | $35470742.89  | 0.86%             |  |  |  | No            |                  2 | On Loan: No      |
| NOVARTIS AG                              | Novartis AG                                  | CUSIP: 000000000<br>LEI: 5493007HIVTX6SY6XD66 | Long             | EC               | CORP              | CH        |       326277 | NS      | $37661868.66  | 0.92%             |  |  |  | No            |                  2 | On Loan: No      |
| NORTHERN STAR RESOURCES LTD              | Northern Star Resources Ltd                  | CUSIP: 000000000<br>LEI: 254900XFXCRYYZAX6M78 | Long             | EC               | CORP              | AU        |      5549965 | NS      | $75472420.62  | 1.84%             |  |  |  | No            |                  2 | On Loan: No      |
| ORIX CORP                                | ORIX Corp                                    | CUSIP: 000000000<br>LEI: 35380016EMHNBOVYE123 | Long             | EC               | CORP              | JP        |      2950400 | NS      | $62524441.22  | 1.52%             |  |  |  | No            |                  2 | On Loan: No      |
| PALADIN ENERGY LTD                       | Paladin Energy Ltd                           | CUSIP: 000000000<br>LEI: 549300M44UTJAP8J6B37 | Long             | EC               | CORP              | AU        |      5339166 | NS      | $21471618.81  | 0.52%             |  |  |  | No            |                  2 | On Loan: No      |
| PAN AMERICAN SILVER CORP                 | Pan American Silver Corp                     | CUSIP: 697900108<br>LEI: 549300MKTF5851TYIB83 | Long             | EC               | CORP              | CA        |      1567084 | NS      | $38221178.76  | 0.93%             |  |  |  | No            |                  1 | On Loan: No      |
| SANWA HOLDINGS CORP                      | Sanwa Holdings Corp                          | CUSIP: 000000000<br>LEI: 5299003X6BCA4ROO5A70 | Long             | EC               | CORP              | JP        |       874800 | NS      | $30116977.23  | 0.73%             |  |  |  | No            |                  2 | On Loan: No      |
| SANKYO CO LTD                            | Sankyo Co Ltd                                | CUSIP: 000000000<br>LEI: 3538009H2R9ZBZAOAK48 | Long             | EC               | CORP              | JP        |      4394300 | NS      | $76641180.49  | 1.87%             |  |  |  | No            |                  2 | On Loan: No      |
| SANOFI SA                                | Sanofi SA                                    | CUSIP: 000000000<br>LEI: 549300E9PC51EN656011 | Long             | EC               | CORP              | FR        |       659684 | NS      | $65335732.45  | 1.59%             |  |  |  | No            |                  2 | On Loan: No      |
| SHIMAMURA CO LTD                         | Shimamura Co Ltd                             | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | JP        |       938800 | NS      | $65056940.32  | 1.58%             |  |  |  | No            |                  2 | On Loan: No      |
| SUMITOMO CORPORATION                     | Sumitomo Corp                                | CUSIP: 000000000<br>LEI: V82KK8NH1P0JS71FJC05 | Long             | EC               | CORP              | JP        |      1812800 | NS      | $46165802.47  | 1.12%             |  |  |  | No            |                  2 | On Loan: No      |
| SUMITOMO MITSUI FIN GRP INC              | Sumitomo Mitsui Financial Group Inc          | CUSIP: 000000000<br>LEI: 35380028MYWPB6AUO129 | Long             | EC               | CORP              | JP        |      3324000 | NS      | $85396680.83  | 2.08%             |  |  |  | No            |                  2 | On Loan: No      |
| SUNDRUG CO LTD                           | Sundrug Co Ltd                               | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | JP        |       712700 | NS      | $22193446.91  | 0.54%             |  |  |  | No            |                  2 | On Loan: No      |
| TECK RESOURCES LTD                       | Teck Resources Ltd                           | CUSIP: 878742204<br>LEI: T8KI5SQ2JRWHL7XY0E11 | Long             | EC               | CORP              | CA        |       223744 | NS      | $8289715.20   | 0.20%             |  |  |  | No            |                  1 | On Loan: No      |
| TOKYO TATEMONO CO LTD                    | Tokyo Tatemono Co Ltd                        | CUSIP: 000000000<br>LEI: 529900RQA4JTW1869J86 | Long             | EC               | CORP              | JP        |      2513100 | NS      | $44511236.20  | 1.08%             |  |  |  | No            |                  2 | On Loan: No      |
| TOPPAN HOLDINGS INC                      | TOPPAN Holdings Inc                          | CUSIP: 000000000<br>LEI: 3538003H9W4KIDODQY20 | Long             | EC               | CORP              | JP        |      2020100 | NS      | $53641364.01  | 1.31%             |  |  |  | No            |                  2 | On Loan: No      |

### Signature

**Date Signed:** 2025-06-20

**Name of Applicant:** Columbia Funds Series Trust

**Signature:** Martha Skinner

**Name of Signer:** Martha Skinner

**Title:** Assistant Treasurer