# EDGAR Filing Document

**Accession Number:** 0000002110
**File Stem:** 0001410368-26-055089
**Filing Date:** 2026-5
**Character Count:** 105723
**Document Hash:** fcb736e607f15b05b570343384402548
**Contains OCR:** False
**Source Format:** 

## Filing Content

## Filing Summary
**0001410368-26-055089.hdr.sgml**: 20260528

**ACCESSION NUMBER**: 0001410368-26-055089

**CONFORMED SUBMISSION TYPE**: NPORT-P

**PUBLIC DOCUMENT COUNT**: 3

**CONFORMED PERIOD OF REPORT**: 20260331

**FILED AS OF DATE**: 20260528

**DATE AS OF CHANGE**: 20260528

**PERIOD START**: 20261231

**FILER**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** COLUMBIA ACORN TRUST
- **CENTRAL INDEX KEY:** 0000002110

**ORGANIZATION NAME:**
- **EIN:** 362692100
- **STATE OF INCORPORATION:** MA
- **FISCAL YEAR END:** 1231

**FILING VALUES:**
- **FORM TYPE:** NPORT-P
- **SEC ACT:** 1940 Act
- **SEC FILE NUMBER:** 811-01829
- **FILM NUMBER:** 261033094

**BUSINESS ADDRESS:**
- **STREET 1:** 290 CONGRESS STREET
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02210
- **BUSINESS PHONE:** 8003456611

**MAIL ADDRESS:**
- **STREET 1:** 290 CONGRESS STREET
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02210

**FORMER COMPANY:**
- **FORMER CONFORMED NAME:** LIBERTY ACORN TRUST
- **DATE OF NAME CHANGE:** 20010424

**FORMER COMPANY:**
- **FORMER CONFORMED NAME:** ACORN INVESTMENT TRUST
- **DATE OF NAME CHANGE:** 19940204

**FORMER COMPANY:**
- **FORMER CONFORMED NAME:** ACORN FUND INC
- **DATE OF NAME CHANGE:** 19920703

## Series and Classes Contracts Data

### Columbia Acorn Fund (Series ID: S000009184)

| Class ID   | Class Name                                | Ticker Symbol   |
|:---|:---|:---|
| C000024954 | Columbia Acorn Fund Class A               | LACAX           |
| C000024956 | Columbia Acorn Fund Class C               | LIACX           |
| C000024957 | Columbia Acorn Fund Institutional Class   | ACRNX           |
| C000122736 | Columbia Acorn Fund Institutional 2 Class | CRBRX           |
| C000122737 | Columbia Acorn Fund Institutional 3 Class | CRBYX           |
| C000251059 | Columbia Acorn Fund Class S               | ACRSX           |

## Nport-Ex

![](img3c841fa41.jpg)

**First Quarter Report**

March 31, 2026 (Unaudited)

Columbia Acorn<sup>®</sup> Fund

Columbia Acorn International<sup>®</sup>

Columbia Acorn International Select<sup>SM</sup>

Columbia Thermostat Fund<sup>SM</sup>

Columbia Acorn European Fund<sup>SM</sup>

---

| | | |
|:---|:---|:---|
| **Not FDIC or NCUA Insured** | **No Financial Institution Guarantee** | **May Lose Value** |

---

------

Portfolio of Investments

Columbia Acorn<sup>®</sup> Fund, March 31, 2026 (Unaudited)

(Percentages represent value of investments compared to net assets)

**Investments in securities**

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Common Stocks 97.8%** | **Common Stocks 97.8%** | **Common Stocks 97.8%** |
| **Issuer** | **Shares** | **Value ($)** |
| **Communication Services 0.6%** | **Communication Services 0.6%** | **Communication Services 0.6%** |
| **Media 0.6%** | **Media 0.6%** | **Media 0.6%** |
| NIQ Global Intelligence PLC<sup>(a)</sup> <br>| &nbsp;&nbsp; 1232928 | &nbsp;&nbsp; 14018391 |
| **Total Communication Services** | **Total Communication Services** | &nbsp;&nbsp; **14018391** |
| **Consumer Discretionary 10.4%** | **Consumer Discretionary 10.4%** | **Consumer Discretionary 10.4%** |
| **Automobile Components 1.6%** | **Automobile Components 1.6%** | **Automobile Components 1.6%** |
| Dorman Products, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 344653 | &nbsp;&nbsp; 35967987 |
| **Hotels, Restaurants & Leisure 4.4%** | **Hotels, Restaurants & Leisure 4.4%** | **Hotels, Restaurants & Leisure 4.4%** |
| Churchill Downs, Inc. | &nbsp;&nbsp; 539022 | &nbsp;&nbsp; 48420346 |
| Rush Street Interactive, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 631555 | &nbsp;&nbsp; 13736321 |
| Wingstop, Inc. | &nbsp;&nbsp; 242143 | &nbsp;&nbsp; 37524901 |
| Total |  | &nbsp;&nbsp; 99681568 |
| **Household Durables 4.4%** | **Household Durables 4.4%** | **Household Durables 4.4%** |
| Cavco Industries, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 83379 | &nbsp;&nbsp; 40379616 |
| SharkNinja, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 238583 | &nbsp;&nbsp; 25265940 |
| TopBuild Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 92442 | &nbsp;&nbsp; 32474874 |
| Total |  | &nbsp;&nbsp; 98120430 |
| **Total Consumer Discretionary** | **Total Consumer Discretionary** | &nbsp;&nbsp; **233769985** |
| **Consumer Staples 4.9%** | **Consumer Staples 4.9%** | **Consumer Staples 4.9%** |
| **Consumer Staples Distribution & Retail 2.4%** | **Consumer Staples Distribution & Retail 2.4%** | **Consumer Staples Distribution & Retail 2.4%** |
| BJ's Wholesale Club Holdings, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 350621 | &nbsp;&nbsp; 34508119 |
| Sprouts Farmers Market, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 248151 | &nbsp;&nbsp; 19139886 |
| Total |  | &nbsp;&nbsp; 53648005 |
| **Household Products 1.3%** | **Household Products 1.3%** | **Household Products 1.3%** |
| WD-40 Co. | &nbsp;&nbsp; 146352 | &nbsp;&nbsp; 29847027 |
| **Personal Care Products 1.2%** | **Personal Care Products 1.2%** | **Personal Care Products 1.2%** |
| elf Beauty, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 444462 | &nbsp;&nbsp; 26938842 |
| **Total Consumer Staples** | **Total Consumer Staples** | &nbsp;&nbsp; **110433874** |
| **Energy 2.5%** | **Energy 2.5%** | **Energy 2.5%** |
| **Oil, Gas & Consumable Fuels 2.5%** | **Oil, Gas & Consumable Fuels 2.5%** | **Oil, Gas & Consumable Fuels 2.5%** |
| Antero Resources Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 355938 | &nbsp;&nbsp; 15106009 |
| Devon Energy Corp. | &nbsp;&nbsp; 421587 | &nbsp;&nbsp; 21214258 |
| EQT Corp. | &nbsp;&nbsp; 311439 | &nbsp;&nbsp; 19819978 |
| Total |  | &nbsp;&nbsp; 56140245 |
| **Total Energy** | **Total Energy** | &nbsp;&nbsp; **56140245** |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Financials 10.4%** | **Financials 10.4%** | **Financials 10.4%** |
| **Banks 1.9%** | **Banks 1.9%** | **Banks 1.9%** |
| Lakeland Financial Corp. | &nbsp;&nbsp; 214229 | &nbsp;&nbsp; 12292460 |
| Western Alliance Bancorp | &nbsp;&nbsp; 419941 | &nbsp;&nbsp; 29752820 |
| Total |  | &nbsp;&nbsp; 42045280 |
| **Capital Markets 7.5%** | **Capital Markets 7.5%** | **Capital Markets 7.5%** |
| Blue Owl Capital, Inc. | &nbsp;&nbsp; 1391042 | &nbsp;&nbsp; 12700213 |
| Carlyle Group, Inc. (The) | &nbsp;&nbsp; 950835 | &nbsp;&nbsp; 46010906 |
| GCM Grosvenor, Inc., Class A<sup>(b)</sup> <br>| &nbsp;&nbsp; 4428750 | &nbsp;&nbsp; 43401750 |
| Hamilton Lane, Inc., Class A | &nbsp;&nbsp; 169441 | &nbsp;&nbsp; 16842435 |
| Miami International Holdings, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 426477 | &nbsp;&nbsp; 16598485 |
| StoneX Group, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 413430 | &nbsp;&nbsp; 33343130 |
| Total |  | &nbsp;&nbsp; 168896919 |
| **Insurance 1.0%** | **Insurance 1.0%** | **Insurance 1.0%** |
| Ryan Specialty Holdings, Inc., Class A | &nbsp;&nbsp; 647394 | &nbsp;&nbsp; 21843073 |
| **Total Financials** | **Total Financials** | &nbsp;&nbsp; **232785272** |
| **Health Care 21.8%** | **Health Care 21.8%** | **Health Care 21.8%** |
| **Biotechnology 8.3%** | **Biotechnology 8.3%** | **Biotechnology 8.3%** |
| Absci Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 2566842 | &nbsp;&nbsp; 7700526 |
| Annexon, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 975637 | &nbsp;&nbsp; 5405029 |
| BridgeBio Pharma, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 316495 | &nbsp;&nbsp; 23502919 |
| Insmed, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 336411 | &nbsp;&nbsp; 55009927 |
| Ionis Pharmaceuticals, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 148670 | &nbsp;&nbsp; 11163630 |
| Madrigal Pharmaceuticals, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 26454 | &nbsp;&nbsp; 13847875 |
| Neurocrine Biosciences, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 116190 | &nbsp;&nbsp; 15306871 |
| NewAmsterdam Pharma Co. NV<sup>(a)</sup> <br>| &nbsp;&nbsp; 363308 | &nbsp;&nbsp; 11629489 |
| Nuvalent, Inc., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 174598 | &nbsp;&nbsp; 17887565 |
| Revolution Medicines, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 135489 | &nbsp;&nbsp; 13176305 |
| Rhythm Pharmaceuticals, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 124824 | &nbsp;&nbsp; 10855943 |
| Total |  | &nbsp;&nbsp; 185486079 |
| **Health Care Equipment & Supplies 2.4%** | **Health Care Equipment & Supplies 2.4%** | **Health Care Equipment & Supplies 2.4%** |
| Glaukos Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 311474 | &nbsp;&nbsp; 33533291 |
| iRhythm Holdings, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 169788 | &nbsp;&nbsp; 20038380 |
| Total |  | &nbsp;&nbsp; 53571671 |

---

Columbia Acorn<sup>®</sup> Fund \| 2026

------

Portfolio of Investments (continued)

Columbia Acorn<sup>®</sup> Fund, March 31, 2026 (Unaudited)

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Health Care Providers & Services 6.6%** | **Health Care Providers & Services 6.6%** | **Health Care Providers & Services 6.6%** |
| Alignment Healthcare, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 1618094 | &nbsp;&nbsp; 28510816 |
| Billiontoone, Inc., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 166205 | &nbsp;&nbsp; 13120223 |
| Encompass Health Corp. | &nbsp;&nbsp; 230244 | &nbsp;&nbsp; 22271502 |
| Guardant Health, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 303784 | &nbsp;&nbsp; 28060528 |
| HealthEquity, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 422315 | &nbsp;&nbsp; 35292865 |
| Pennant Group, Inc. (The)<sup>(a)</sup> <br>| &nbsp;&nbsp; 699307 | &nbsp;&nbsp; 21314877 |
| Total |  | &nbsp;&nbsp; 148570811 |
| **Health Care Technology 0.5%** | **Health Care Technology 0.5%** | **Health Care Technology 0.5%** |
| Doximity, Inc., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 432308 | &nbsp;&nbsp; 10072777 |
| **Life Sciences Tools & Services 3.2%** | **Life Sciences Tools & Services 3.2%** | **Life Sciences Tools & Services 3.2%** |
| Adaptive Biotechnologies Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 1148404 | &nbsp;&nbsp; 15939847 |
| BioLife Solutions, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 696462 | &nbsp;&nbsp; 13288495 |
| Charles River Laboratories International, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 70538 | &nbsp;&nbsp; 12167805 |
| DNA Script<sup>(a),(c),(d),(e)</sup> <br>| &nbsp;&nbsp; 1139 | &nbsp;&nbsp; 94710 |
| Fortrea Holdings, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 1173131 | &nbsp;&nbsp; 11050894 |
| Repligen Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 159623 | &nbsp;&nbsp; 18806782 |
| Total |  | &nbsp;&nbsp; 71348533 |
| **Pharmaceuticals 0.8%** | **Pharmaceuticals 0.8%** | **Pharmaceuticals 0.8%** |
| Crinetics Pharmaceuticals, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 252431 | &nbsp;&nbsp; 9168294 |
| Definium Therapeutics, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 512390 | &nbsp;&nbsp; 9684171 |
| Total |  | &nbsp;&nbsp; 18852465 |
| **Total Health Care** | **Total Health Care** | &nbsp;&nbsp; **487902336** |
| **Industrials 29.4%** | **Industrials 29.4%** | **Industrials 29.4%** |
| **Aerospace & Defense 10.0%** | **Aerospace & Defense 10.0%** | **Aerospace & Defense 10.0%** |
| Aerovironment, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 153583 | &nbsp;&nbsp; 28113368 |
| Astronics Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 340163 | &nbsp;&nbsp; 22699077 |
| Axon Enterprise, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 61466 | &nbsp;&nbsp; 26103995 |
| Curtiss-Wright Corp. | &nbsp;&nbsp; 65825 | &nbsp;&nbsp; 44834724 |
| FTAI Aviation Ltd. | &nbsp;&nbsp; 109872 | &nbsp;&nbsp; 26918640 |
| Karman Holdings, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 220172 | &nbsp;&nbsp; 17624769 |
| Kratos Defense & Security Solutions, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 233250 | &nbsp;&nbsp; 16446457 |
| VSE Corp. | &nbsp;&nbsp; 223919 | &nbsp;&nbsp; 41290664 |
| Total |  | &nbsp;&nbsp; 224031694 |
| **Building Products 2.6%** | **Building Products 2.6%** | **Building Products 2.6%** |
| Janus International Group, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 3345300 | &nbsp;&nbsp; 17228295 |
| Modine Manufacturing Co.<sup>(a)</sup> <br>| &nbsp;&nbsp; 189531 | &nbsp;&nbsp; 41073263 |
| Total |  | &nbsp;&nbsp; 58301558 |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Commercial Services & Supplies 0.7%** | **Commercial Services & Supplies 0.7%** | **Commercial Services & Supplies 0.7%** |
| Healthcare Services Group, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 784900 | &nbsp;&nbsp; 14559895 |
| **Construction & Engineering 3.8%** | **Construction & Engineering 3.8%** | **Construction & Engineering 3.8%** |
| MasTec, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 131179 | &nbsp;&nbsp; 42205532 |
| Sterling Infrastructure, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 107856 | &nbsp;&nbsp; 43926513 |
| Total |  | &nbsp;&nbsp; 86132045 |
| **Electrical Equipment 2.1%** | **Electrical Equipment 2.1%** | **Electrical Equipment 2.1%** |
| Bloom Energy Corp., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 288864 | &nbsp;&nbsp; 39138183 |
| Nextpower, Inc., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 71632 | &nbsp;&nbsp; 8635238 |
| Total |  | &nbsp;&nbsp; 47773421 |
| **Ground Transportation 1.1%** | **Ground Transportation 1.1%** | **Ground Transportation 1.1%** |
| Knight-Swift Transportation Holdings, Inc. | &nbsp;&nbsp; 437705 | &nbsp;&nbsp; 25203054 |
| **Machinery 4.7%** | **Machinery 4.7%** | **Machinery 4.7%** |
| Esab Corp. | &nbsp;&nbsp; 372810 | &nbsp;&nbsp; 36035815 |
| Mueller Water Products, Inc., Class A | &nbsp;&nbsp; 483160 | &nbsp;&nbsp; 13282068 |
| SPX Technologies, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 279748 | &nbsp;&nbsp; 55932815 |
| Total |  | &nbsp;&nbsp; 105250698 |
| **Professional Services 2.7%** | **Professional Services 2.7%** | **Professional Services 2.7%** |
| Parsons Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 486820 | &nbsp;&nbsp; 26371039 |
| Paylocity Holding Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 167076 | &nbsp;&nbsp; 18050891 |
| Willdan Group, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 206119 | &nbsp;&nbsp; 15780471 |
| Total |  | &nbsp;&nbsp; 60202401 |
| **Trading Companies & Distributors 1.7%** | **Trading Companies & Distributors 1.7%** | **Trading Companies & Distributors 1.7%** |
| Applied Industrial Technologies, Inc. | &nbsp;&nbsp; 84867 | &nbsp;&nbsp; 22516913 |
| McGrath Rentcorp | &nbsp;&nbsp; 136694 | &nbsp;&nbsp; 15074614 |
| Total |  | &nbsp;&nbsp; 37591527 |
| **Total Industrials** | **Total Industrials** | &nbsp;&nbsp; **659046293** |
| **Information Technology 17.8%** | **Information Technology 17.8%** | **Information Technology 17.8%** |
| **Communications Equipment 3.9%** | **Communications Equipment 3.9%** | **Communications Equipment 3.9%** |
| InterDigital, Inc. | &nbsp;&nbsp; 140996 | &nbsp;&nbsp; 42580792 |
| Viavi Solutions, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 1333081 | &nbsp;&nbsp; 44364936 |
| Total |  | &nbsp;&nbsp; 86945728 |
| **Electronic Equipment, Instruments & Components 4.4%** | **Electronic Equipment, Instruments & Components 4.4%** | **Electronic Equipment, Instruments & Components 4.4%** |
| Advanced Energy Industries, Inc. | &nbsp;&nbsp; 28789 | &nbsp;&nbsp; 9290498 |
| Celestica, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 172300 | &nbsp;&nbsp; 48533464 |
| Coherent Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 172879 | &nbsp;&nbsp; 41181507 |
| Total |  | &nbsp;&nbsp; 99005469 |

---

Columbia Acorn<sup>®</sup> Fund \| 2026

------

Portfolio of Investments (continued)

Columbia Acorn<sup>®</sup> Fund, March 31, 2026 (Unaudited)

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **IT Services 0.4%** | **IT Services 0.4%** | **IT Services 0.4%** |
| Applied Digital Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 439896 | &nbsp;&nbsp; 10443131 |
| **Semiconductors & Semiconductor Equipment 4.9%** | **Semiconductors & Semiconductor Equipment 4.9%** | **Semiconductors & Semiconductor Equipment 4.9%** |
| Credo Technology Group Holding Ltd.<sup>(a)</sup> <br>| &nbsp;&nbsp; 147591 | &nbsp;&nbsp; 13854367 |
| Lattice Semiconductor Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 184881 | &nbsp;&nbsp; 17149562 |
| Marvell Technology, Inc. | &nbsp;&nbsp; 298398 | &nbsp;&nbsp; 29556322 |
| Semtech Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 421296 | &nbsp;&nbsp; 32393449 |
| SiTime Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 48846 | &nbsp;&nbsp; 16868966 |
| Total |  | &nbsp;&nbsp; 109822666 |
| **Software 4.2%** | **Software 4.2%** | **Software 4.2%** |
| Dynatrace, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 695469 | &nbsp;&nbsp; 25718444 |
| Hut 8 Corp.<sup>(a)</sup> <br>| &nbsp;&nbsp; 240717 | &nbsp;&nbsp; 11292034 |
| Monday.com Ltd.<sup>(a)</sup> <br>| &nbsp;&nbsp; 106304 | &nbsp;&nbsp; 7346669 |
| SailPoint, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 944155 | &nbsp;&nbsp; 12500612 |
| Workiva, Inc., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 184993 | &nbsp;&nbsp; 11031133 |
| Zeta Global Holdings Corp., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 895992 | &nbsp;&nbsp; 14264193 |
| Zscaler, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 83570 | &nbsp;&nbsp; 11724035 |
| Total |  | &nbsp;&nbsp; 93877120 |
| **Total Information Technology** | **Total Information Technology** | &nbsp;&nbsp; **400094114** |
| &nbsp;&nbsp; Total Common Stocks <br>(Cost $1,925,239,900) | &nbsp;&nbsp; Total Common Stocks <br>(Cost $1,925,239,900) | &nbsp;&nbsp; **2194190510** |

---

---

| | | |
|:---|:---|:---|
| **Exchange-Traded Equity Funds 1.6%** | **Exchange-Traded Equity Funds 1.6%** | **Exchange-Traded Equity Funds 1.6%** |
|  | **Shares** | **Value ($)** |
| **Sector 1.6%** | **Sector 1.6%** | **Sector 1.6%** |
| State Street SPDR S&P Biotech ETF | &nbsp;&nbsp; 278004 | &nbsp;&nbsp; 35509451 |
| &nbsp;&nbsp; Total Exchange-Traded Equity Funds <br>(Cost $28,404,882) | &nbsp;&nbsp; Total Exchange-Traded Equity Funds <br>(Cost $28,404,882) | &nbsp;&nbsp; **35509451** |
| **Money Market Funds 0.9%** | **Money Market Funds 0.9%** | **Money Market Funds 0.9%** |
| Columbia Short-Term Cash Fund, 3.790%<sup>(b),(f)</sup> <br>| &nbsp;&nbsp; 19982329 | &nbsp;&nbsp; 19974336 |
| &nbsp;&nbsp; Total Money Market Funds <br>(Cost $19,971,514) | &nbsp;&nbsp; Total Money Market Funds <br>(Cost $19,971,514) | &nbsp;&nbsp; **19974336** |
| **Total Investments in Securities** <br>**(Cost: $1,973,616,296)** | **Total Investments in Securities** <br>**(Cost: $1,973,616,296)** | &nbsp;&nbsp; **2249674297** |
| **Other Assets & Liabilities, Net** |  | &nbsp;&nbsp; **(5875383)**<br>|
| **Net Assets** | **Net Assets** | &nbsp;&nbsp; **2243798914** |

---

**Notes to Portfolio of Investments** 

&nbsp;&nbsp;&nbsp;&nbsp;(a) Non-income producing investment.

&nbsp;&nbsp;&nbsp;&nbsp;(b) Under the Investment Company Act of 1940, an affiliated company is one in which the Fund owns 5% or more of the company's outstanding voting securities, or a company which is under common ownership or control with the Fund. The value of the
 holdings and transactions in these affiliated companies during the period ended March 31, 2026 are as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Affiliated issuers** | **Beginning** <br>**of period($)**<br>| **Purchases($)** | **Sales($)** | **Net change in** <br>**unrealized** <br>**appreciation** <br>**(depreciation)($)**<br>| **End of** <br>**period($)**<br>| **Realized gain** <br>**(loss)($)**<br>| **Dividends($)** | **End of** <br>**period shares**<br>|
| Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% |
|  | 15279105 | &nbsp;&nbsp; 183720481 | &nbsp;&nbsp; (179024209)<br>| &nbsp;&nbsp; (1041)<br>| &nbsp;&nbsp; 19974336 | &nbsp;&nbsp; (2205)<br>| &nbsp;&nbsp; 261355 | &nbsp;&nbsp; 19982329 |
| GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A | GCM Grosvenor, Inc., Class A |
|  | 50133450 | &nbsp;&nbsp; — | &nbsp;&nbsp; — | &nbsp;&nbsp; (6731700)<br>| &nbsp;&nbsp; 43401750 | &nbsp;&nbsp; — | &nbsp;&nbsp; 531450 | &nbsp;&nbsp; 4428750 |
| Total | 65412555 |  |  | &nbsp;&nbsp; (6732741)<br>| &nbsp;&nbsp; 63376086 | &nbsp;&nbsp; (2205)<br>| &nbsp;&nbsp; 792805 |  |

---

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

Columbia Acorn<sup>®</sup> Fund \| 2026

------

Portfolio of Investments (continued)

Columbia Acorn<sup>®</sup> Fund, March 31, 2026 (Unaudited)

**Notes to Portfolio of Investments (continued)**

&nbsp;&nbsp;&nbsp;&nbsp;(c) Represents fair value as determined in good faith under procedures approved by the
 Board of Trustees. At March 31, 2026, the total value of these securities amounted to $94,710, which represents less than 0.01% of total net assets.

&nbsp;&nbsp;&nbsp;&nbsp;(d) Denotes a restricted security, which is subject to legal or contractual restrictions
 on resale under federal securities laws. Disposal of a restricted investment may involve time-consuming negotiations and expenses, and prompt sale at an acceptable
 price may be difficult to achieve. Private placement securities are generally considered to be restricted, although certain of those securities may be traded between
 qualified institutional investors under the provisions of Section 4(a)(2) and Rule 144A. The Fund will not incur any registration costs upon such a trade. At March
 31, 2026, the total market value of these securities amounted to $94,710, which represents less than 0.01% of total net assets. Additional information on these securities
 is as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | |
|:---|:---|:---|:---|:---|
| **Security** | **Acquisition** <br>**Dates**<br>| **Shares** | **Cost ($)** | **Value ($)** |
| DNA Script | 10/01/2021 | &nbsp;&nbsp; 1139 | &nbsp;&nbsp; 993179 | &nbsp;&nbsp; 94710 |

---

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;(e) Valuation based on significant unobservable inputs.

&nbsp;&nbsp;&nbsp;&nbsp;(f) The rate shown is the seven-day current annualized yield at March 31, 2026.

Investments are valued using policies described in the Notes to Financial Statements included within the most recent financial statements and additional information.

Columbia Acorn<sup>®</sup> Fund \| 2026

------

Portfolio of Investments

Columbia Acorn International<sup>®</sup>, March 31, 2026 (Unaudited)

(Percentages represent value of investments compared to net assets)

**Investments in securities**

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Common Stocks 98.5%** | **Common Stocks 98.5%** | **Common Stocks 98.5%** |
| **Issuer** | **Shares** | **Value ($)** |
| **Australia 2.0%** | **Australia 2.0%** | **Australia 2.0%** |
| ALS Ltd. | &nbsp;&nbsp; 763610 | &nbsp;&nbsp; 11209483 |
| CAR Group Ltd. | &nbsp;&nbsp; 245518 | &nbsp;&nbsp; 3919294 |
| **Total** | **Total** | &nbsp;&nbsp; **15128777** |
| **Canada 1.5%** | **Canada 1.5%** | **Canada 1.5%** |
| CCL Industries, Inc., Class B | &nbsp;&nbsp; 64213 | &nbsp;&nbsp; 4023294 |
| Finning International, Inc. | &nbsp;&nbsp; 119677 | &nbsp;&nbsp; 7405504 |
| **Total** | **Total** | &nbsp;&nbsp; **11428798** |
| **China 1.4%** | **China 1.4%** | **China 1.4%** |
| Silergy Corp. | &nbsp;&nbsp; 1138000 | &nbsp;&nbsp; 10371353 |
| **Denmark 1.0%** | **Denmark 1.0%** | **Denmark 1.0%** |
| FLSmidth & Co. A/S | &nbsp;&nbsp; 95384 | &nbsp;&nbsp; 7225956 |
| **Finland 2.8%** | **Finland 2.8%** | **Finland 2.8%** |
| Konecranes OYJ | &nbsp;&nbsp; 307904 | &nbsp;&nbsp; 10105628 |
| Valmet OYJ | &nbsp;&nbsp; 382556 | &nbsp;&nbsp; 10946292 |
| **Total** | **Total** | &nbsp;&nbsp; **21051920** |
| **France 4.3%** | **France 4.3%** | **France 4.3%** |
| Gaztransport Et Technigaz SA | &nbsp;&nbsp; 76782 | &nbsp;&nbsp; 18058645 |
| Lisi SA | &nbsp;&nbsp; 84378 | &nbsp;&nbsp; 5159101 |
| Robertet SA | &nbsp;&nbsp; 4147 | &nbsp;&nbsp; 3874920 |
| Virbac SA | &nbsp;&nbsp; 13867 | &nbsp;&nbsp; 5727256 |
| **Total** | **Total** | &nbsp;&nbsp; **32819922** |
| **Germany 4.5%** | **Germany 4.5%** | **Germany 4.5%** |
| CTS Eventim AG & Co. KGaA | &nbsp;&nbsp; 96909 | &nbsp;&nbsp; 5676029 |
| flatexDEGIRO AG | &nbsp;&nbsp; 262133 | &nbsp;&nbsp; 9039748 |
| Nemetschek SE | &nbsp;&nbsp; 111618 | &nbsp;&nbsp; 8355724 |
| Renk Group AG | &nbsp;&nbsp; 187745 | &nbsp;&nbsp; 11221836 |
| **Total** | **Total** | &nbsp;&nbsp; **34293337** |
| **Greece 1.9%** | **Greece 1.9%** | **Greece 1.9%** |
| National Bank of Greece SA | &nbsp;&nbsp; 914195 | &nbsp;&nbsp; 14122379 |
| **Ireland 2.6%** | **Ireland 2.6%** | **Ireland 2.6%** |
| Bank of Ireland Group PLC | &nbsp;&nbsp; 1089587 | &nbsp;&nbsp; 19768641 |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Italy 3.9%** | **Italy 3.9%** | **Italy 3.9%** |
| Carel Industries SpA | &nbsp;&nbsp; 241694 | &nbsp;&nbsp; 6159932 |
| Prysmian SpA | &nbsp;&nbsp; 194146 | &nbsp;&nbsp; 22926912 |
| **Total** | **Total** | &nbsp;&nbsp; **29086844** |
| **Japan 41.0%** | **Japan 41.0%** | **Japan 41.0%** |
| Anritsu Corp. | &nbsp;&nbsp; 207600 | &nbsp;&nbsp; 3769674 |
| Asics Corp. | &nbsp;&nbsp; 566200 | &nbsp;&nbsp; 15222352 |
| BayCurrent, Inc. | &nbsp;&nbsp; 194400 | &nbsp;&nbsp; 5627167 |
| Capcom Co., Ltd. | &nbsp;&nbsp; 714600 | &nbsp;&nbsp; 15100877 |
| Disco Corp. | &nbsp;&nbsp; 31400 | &nbsp;&nbsp; 12798116 |
| DMG Mori Co., Ltd. | &nbsp;&nbsp; 312400 | &nbsp;&nbsp; 4826020 |
| Gunma Bank Ltd. (The) | &nbsp;&nbsp; 748400 | &nbsp;&nbsp; 9973934 |
| Kokusai Electric Corp. | &nbsp;&nbsp; 360100 | &nbsp;&nbsp; 12176672 |
| Konami Holdings Corp. | &nbsp;&nbsp; 47500 | &nbsp;&nbsp; 5853338 |
| Kraftia Corp. | &nbsp;&nbsp; 308900 | &nbsp;&nbsp; 18683980 |
| Kyoritsu Maintenance Co., Ltd. | &nbsp;&nbsp; 607100 | &nbsp;&nbsp; 9422009 |
| MatsukiyoCocokara & Co. | &nbsp;&nbsp; 538300 | &nbsp;&nbsp; 8567756 |
| Modec, Inc. | &nbsp;&nbsp; 87600 | &nbsp;&nbsp; 8292576 |
| Nippon Sanso Holdings Corp. | &nbsp;&nbsp; 432900 | &nbsp;&nbsp; 15349942 |
| Nissin Foods Holdings Co., Ltd. | &nbsp;&nbsp; 629100 | &nbsp;&nbsp; 11940135 |
| Niterra Co., Ltd. | &nbsp;&nbsp; 444100 | &nbsp;&nbsp; 20940012 |
| Nomura Real Estate Holdings, Inc. | &nbsp;&nbsp; 2734500 | &nbsp;&nbsp; 17729907 |
| Omron Corp. | &nbsp;&nbsp; 409300 | &nbsp;&nbsp; 11775190 |
| Open House Co., Ltd. | &nbsp;&nbsp; 175800 | &nbsp;&nbsp; 11267328 |
| Recruit Holdings Co., Ltd. | &nbsp;&nbsp; 105600 | &nbsp;&nbsp; 4600989 |
| Renesas Electronics Corp. | &nbsp;&nbsp; 697200 | &nbsp;&nbsp; 9968029 |
| Sanwa Holdings Corp. | &nbsp;&nbsp; 495050 | &nbsp;&nbsp; 11258555 |
| Sekisui Chemical Co., Ltd. | &nbsp;&nbsp; 720000 | &nbsp;&nbsp; 12087839 |
| Shimadzu Corp. | &nbsp;&nbsp; 299800 | &nbsp;&nbsp; 7124106 |
| Simplex Holdings, Inc. | &nbsp;&nbsp; 2400400 | &nbsp;&nbsp; 12340130 |
| Suntory Beverage & Food Ltd. | &nbsp;&nbsp; 463600 | &nbsp;&nbsp; 13092112 |
| Taisei Corp. | &nbsp;&nbsp; 186800 | &nbsp;&nbsp; 19352806 |
| **Total** | **Total** | &nbsp;&nbsp; **309141551** |
| **Jersey 1.8%** | **Jersey 1.8%** | **Jersey 1.8%** |
| Yellow Cake PLC<sup>(a)</sup> <br>| &nbsp;&nbsp; 1722108 | &nbsp;&nbsp; 13791748 |

---

Columbia Acorn International<sup>®</sup> \| 2026

------

Portfolio of Investments (continued)

Columbia Acorn International<sup>®</sup>, March 31, 2026 (Unaudited)

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Netherlands 2.3%** | **Netherlands 2.3%** | **Netherlands 2.3%** |
| BE Semiconductor Industries NV | &nbsp;&nbsp; 79720 | &nbsp;&nbsp; 17075535 |
| **New Zealand 2.3%** | **New Zealand 2.3%** | **New Zealand 2.3%** |
| Fisher & Paykel Healthcare Corp., Ltd. | &nbsp;&nbsp; 819257 | &nbsp;&nbsp; 17764346 |
| **Sweden 2.9%** | **Sweden 2.9%** | **Sweden 2.9%** |
| AddTech AB, B Shares | &nbsp;&nbsp; 315479 | &nbsp;&nbsp; 10825644 |
| Hexagon AB, Class B | &nbsp;&nbsp; 1133858 | &nbsp;&nbsp; 11034839 |
| **Total** | **Total** | &nbsp;&nbsp; **21860483** |
| **Switzerland 5.7%** | **Switzerland 5.7%** | **Switzerland 5.7%** |
| Belimo Holding AG, Registered Shares | &nbsp;&nbsp; 10342 | &nbsp;&nbsp; 8399243 |
| Inficon Holding AG, Registered Shares | &nbsp;&nbsp; 60601 | &nbsp;&nbsp; 7669748 |
| Kardex Holding AG | &nbsp;&nbsp; 24629 | &nbsp;&nbsp; 7410232 |
| SGS SA, Registered Shares | &nbsp;&nbsp; 72936 | &nbsp;&nbsp; 7686652 |
| VAT Group AG | &nbsp;&nbsp; 19160 | &nbsp;&nbsp; 11952808 |
| **Total** | **Total** | &nbsp;&nbsp; **43118683** |
| **United Kingdom 11.9%** | **United Kingdom 11.9%** | **United Kingdom 11.9%** |
| Babcock International Group PLC | &nbsp;&nbsp; 1053570 | &nbsp;&nbsp; 16356265 |
| Baltic Classifieds Group PLC | &nbsp;&nbsp; 2200698 | &nbsp;&nbsp; 5330504 |
| ConvaTec Group PLC | &nbsp;&nbsp; 3120353 | &nbsp;&nbsp; 9000465 |
| Endeavour Mining PLC | &nbsp;&nbsp; 246358 | &nbsp;&nbsp; 14843017 |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| Halma PLC | &nbsp;&nbsp; 409137 | &nbsp;&nbsp; 20878575 |
| ICG PLC | &nbsp;&nbsp; 465471 | &nbsp;&nbsp; 9546474 |
| Rightmove PLC | &nbsp;&nbsp; 754204 | &nbsp;&nbsp; 4315039 |
| Safestore Holdings PLC | &nbsp;&nbsp; 1099285 | &nbsp;&nbsp; 9232062 |
| **Total** | **Total** | &nbsp;&nbsp; **89502401** |
| **United States 4.7%** | **United States 4.7%** | **United States 4.7%** |
| CRH PLC | &nbsp;&nbsp; 138047 | &nbsp;&nbsp; 14442549 |
| Interparfums, Inc. | &nbsp;&nbsp; 93939 | &nbsp;&nbsp; 8533419 |
| Sunbelt Rentals Holdings, Inc. | &nbsp;&nbsp; 201715 | &nbsp;&nbsp; 12867673 |
| **Total** | **Total** | &nbsp;&nbsp; **35843641** |
| &nbsp;&nbsp; Total Common Stocks <br>(Cost $590,592,640) | &nbsp;&nbsp; Total Common Stocks <br>(Cost $590,592,640) | &nbsp;&nbsp; **743396315** |
| **Money Market Funds 0.8%** | **Money Market Funds 0.8%** | **Money Market Funds 0.8%** |
|  | **Shares** | **Value ($)** |
| Columbia Short-Term Cash Fund, 3.790%<sup>(b),(c)</sup> <br>| &nbsp;&nbsp; 5931736 | &nbsp;&nbsp; 5929363 |
| &nbsp;&nbsp; Total Money Market Funds <br>(Cost $5,929,711) | &nbsp;&nbsp; Total Money Market Funds <br>(Cost $5,929,711) | &nbsp;&nbsp; **5929363** |
| **Total Investments in Securities** <br>**(Cost $596,522,351)** | **Total Investments in Securities** <br>**(Cost $596,522,351)** | &nbsp;&nbsp; **749325678** |
| **Other Assets & Liabilities, Net** |  | &nbsp;&nbsp; **5326459** |
| **Net Assets** | **Net Assets** | &nbsp;&nbsp; **$754652137** |

---

**Notes to Portfolio of Investments** 

&nbsp;&nbsp;&nbsp;&nbsp;(a) Non-income producing investment.

&nbsp;&nbsp;&nbsp;&nbsp;(b) The rate shown is the seven-day current annualized yield at March 31, 2026.

&nbsp;&nbsp;&nbsp;&nbsp;(c) Under the Investment Company Act of 1940, an affiliated company is one in which the Fund owns 5% or more of the company's outstanding voting securities, or a company which is under common ownership or control with the Fund. The value of the
 holdings and transactions in these affiliated companies during the period ended March 31, 2026 are as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Affiliated issuers** | **Beginning** <br>**of period($)**<br>| **Purchases($)** | **Sales($)** | **Net change in** <br>**unrealized** <br>**appreciation** <br>**(depreciation)($)**<br>| **End of** <br>**period($)**<br>| **Realized gain** <br>**(loss)($)**<br>| **Dividends($)** | **End of** <br>**period shares**<br>|
| Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% |
|  | 4175673 | &nbsp;&nbsp; 71937529 | &nbsp;&nbsp; (70183315)<br>| &nbsp;&nbsp; (524)<br>| &nbsp;&nbsp; 5929363 | &nbsp;&nbsp; (680)<br>| &nbsp;&nbsp; 74853 | &nbsp;&nbsp; 5931736 |

---

Investments are valued using policies described in the Notes to Financial Statements included within the most recent financial statements and additional information.

Columbia Acorn International<sup>®</sup> \| 2026

------

Portfolio of Investments

Columbia Acorn International Select<sup>SM</sup>, March 31, 2026 (Unaudited)

(Percentages represent value of investments compared to net assets)

**Investments in securities**

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Common Stocks 96.9%** | **Common Stocks 96.9%** | **Common Stocks 96.9%** |
| **Issuer** | **Shares** | **Value ($)** |
| **Australia 1.3%** | **Australia 1.3%** | **Australia 1.3%** |
| ALS Ltd. | &nbsp;&nbsp; 157176 | &nbsp;&nbsp; 2307280 |
| **Belgium 1.2%** | **Belgium 1.2%** | **Belgium 1.2%** |
| Anheuser-Busch InBev SA/NV | &nbsp;&nbsp; 32826 | &nbsp;&nbsp; 2271103 |
| **Canada 1.4%** | **Canada 1.4%** | **Canada 1.4%** |
| Alimentation Couche-Tard, Inc. | &nbsp;&nbsp; 32122 | &nbsp;&nbsp; 1820731 |
| Shopify, Inc., Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 5650 | &nbsp;&nbsp; 670397 |
| **Total** | **Total** | &nbsp;&nbsp; **2491128** |
| **Denmark 2.7%** | **Denmark 2.7%** | **Denmark 2.7%** |
| FLSmidth & Co. A/S | &nbsp;&nbsp; 20204 | &nbsp;&nbsp; 1530584 |
| Novo Nordisk A/S, Class B | &nbsp;&nbsp; 92938 | &nbsp;&nbsp; 3401019 |
| **Total** | **Total** | &nbsp;&nbsp; **4931603** |
| **Finland 0.7%** | **Finland 0.7%** | **Finland 0.7%** |
| Konecranes OYJ | &nbsp;&nbsp; 41950 | &nbsp;&nbsp; 1376829 |
| **France 14.8%** | **France 14.8%** | **France 14.8%** |
| Airbus Group SE | &nbsp;&nbsp; 29167 | &nbsp;&nbsp; 5514691 |
| Cie de Saint-Gobain SA | &nbsp;&nbsp; 36439 | &nbsp;&nbsp; 3016947 |
| EssilorLuxottica SA | &nbsp;&nbsp; 15082 | &nbsp;&nbsp; 3514485 |
| LVMH Moet Hennessy Louis Vuitton SE | &nbsp;&nbsp; 8139 | &nbsp;&nbsp; 4449181 |
| Publicis Groupe SA | &nbsp;&nbsp; 26316 | &nbsp;&nbsp; 2178162 |
| Schneider Electric SE | &nbsp;&nbsp; 20193 | &nbsp;&nbsp; 5500202 |
| TotalEnergies SE | &nbsp;&nbsp; 32861 | &nbsp;&nbsp; 3015790 |
| **Total** | **Total** | &nbsp;&nbsp; **27189458** |
| **Germany 6.2%** | **Germany 6.2%** | **Germany 6.2%** |
| Adidas AG | &nbsp;&nbsp; 15472 | &nbsp;&nbsp; 2504247 |
| Deutsche Telekom AG, Registered Shares | &nbsp;&nbsp; 123264 | &nbsp;&nbsp; 4600615 |
| Nemetschek SE | &nbsp;&nbsp; 25067 | &nbsp;&nbsp; 1876516 |
| Renk Group AG | &nbsp;&nbsp; 41899 | &nbsp;&nbsp; 2504374 |
| **Total** | **Total** | &nbsp;&nbsp; **11485752** |
| **Ireland 2.3%** | **Ireland 2.3%** | **Ireland 2.3%** |
| Bank of Ireland Group PLC | &nbsp;&nbsp; 236857 | &nbsp;&nbsp; 4297354 |
| **Italy 2.4%** | **Italy 2.4%** | **Italy 2.4%** |
| Prysmian SpA | &nbsp;&nbsp; 36742 | &nbsp;&nbsp; 4338903 |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Japan 25.1%** | **Japan 25.1%** | **Japan 25.1%** |
| Asics Corp. | &nbsp;&nbsp; 83300 | &nbsp;&nbsp; 2239530 |
| BayCurrent, Inc. | &nbsp;&nbsp; 26300 | &nbsp;&nbsp; 761289 |
| Capcom Co., Ltd. | &nbsp;&nbsp; 212900 | &nbsp;&nbsp; 4498988 |
| Disco Corp. | &nbsp;&nbsp; 9300 | &nbsp;&nbsp; 3790525 |
| Gunma Bank Ltd. (The) | &nbsp;&nbsp; 168500 | &nbsp;&nbsp; 2245601 |
| Kokusai Electric Corp. | &nbsp;&nbsp; 76800 | &nbsp;&nbsp; 2596969 |
| MatsukiyoCocokara & Co. | &nbsp;&nbsp; 129100 | &nbsp;&nbsp; 2054797 |
| Niterra Co., Ltd. | &nbsp;&nbsp; 76900 | &nbsp;&nbsp; 3625956 |
| Nomura Real Estate Holdings, Inc. | &nbsp;&nbsp; 322000 | &nbsp;&nbsp; 2087778 |
| Omron Corp. | &nbsp;&nbsp; 98800 | &nbsp;&nbsp; 2842386 |
| Recruit Holdings Co., Ltd. | &nbsp;&nbsp; 59500 | &nbsp;&nbsp; 2592413 |
| Sekisui Chemical Co., Ltd. | &nbsp;&nbsp; 113400 | &nbsp;&nbsp; 1903835 |
| Shimadzu Corp. | &nbsp;&nbsp; 106600 | &nbsp;&nbsp; 2533121 |
| Suntory Beverage & Food Ltd. | &nbsp;&nbsp; 98100 | &nbsp;&nbsp; 2770354 |
| Taisei Corp. | &nbsp;&nbsp; 49800 | &nbsp;&nbsp; 5159367 |
| Tokyo Electron Ltd. | &nbsp;&nbsp; 18400 | &nbsp;&nbsp; 4571497 |
| **Total** | **Total** | &nbsp;&nbsp; **46274406** |
| **Netherlands 4.8%** | **Netherlands 4.8%** | **Netherlands 4.8%** |
| ASML Holding NV | &nbsp;&nbsp; 6671 | &nbsp;&nbsp; 8870807 |
| **New Zealand 1.1%** | **New Zealand 1.1%** | **New Zealand 1.1%** |
| Fisher & Paykel Healthcare Corp., Ltd. | &nbsp;&nbsp; 89689 | &nbsp;&nbsp; 1944770 |
| **Spain 2.6%** | **Spain 2.6%** | **Spain 2.6%** |
| Industria de Diseno Textil SA | &nbsp;&nbsp; 83602 | &nbsp;&nbsp; 4866371 |
| **Switzerland 8.0%** | **Switzerland 8.0%** | **Switzerland 8.0%** |
| Belimo Holding AG, Registered Shares | &nbsp;&nbsp; 1722 | &nbsp;&nbsp; 1398520 |
| &nbsp;&nbsp; Cie Financiere Richemont SA, Class A, Registered <br> Shares<br>| &nbsp;&nbsp; 7593 | &nbsp;&nbsp; 1340667 |
| Lonza Group AG, Registered Shares | &nbsp;&nbsp; 6512 | &nbsp;&nbsp; 4177569 |
| Nestlé SA, Registered Shares | &nbsp;&nbsp; 36109 | &nbsp;&nbsp; 3541970 |
| SGS SA, Registered Shares | &nbsp;&nbsp; 17926 | &nbsp;&nbsp; 1889203 |
| VAT Group AG | &nbsp;&nbsp; 3930 | &nbsp;&nbsp; 2451698 |
| **Total** | **Total** | &nbsp;&nbsp; **14799627** |

---

Columbia Acorn International Select<sup>SM</sup> \| 2026

------

Portfolio of Investments (continued)

Columbia Acorn International Select<sup>SM</sup>, March 31, 2026 (Unaudited)

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **United Kingdom 18.7%** | **United Kingdom 18.7%** | **United Kingdom 18.7%** |
| 3i Group PLC | &nbsp;&nbsp; 92270 | &nbsp;&nbsp; 3007172 |
| Babcock International Group PLC | &nbsp;&nbsp; 195921 | &nbsp;&nbsp; 3041597 |
| BT Group PLC | &nbsp;&nbsp; 1857140 | &nbsp;&nbsp; 5204759 |
| ConvaTec Group PLC | &nbsp;&nbsp; 665130 | &nbsp;&nbsp; 1918526 |
| Endeavour Mining PLC | &nbsp;&nbsp; 45050 | &nbsp;&nbsp; 2714253 |
| GSK PLC | &nbsp;&nbsp; 70565 | &nbsp;&nbsp; 1943747 |
| Halma PLC | &nbsp;&nbsp; 44904 | &nbsp;&nbsp; 2291486 |
| ICG PLC | &nbsp;&nbsp; 104163 | &nbsp;&nbsp; 2136308 |
| London Stock Exchange Group PLC | &nbsp;&nbsp; 17675 | &nbsp;&nbsp; 2087201 |
| Reckitt Benckiser Group PLC | &nbsp;&nbsp; 42587 | &nbsp;&nbsp; 2863564 |
| Rightmove PLC | &nbsp;&nbsp; 190972 | &nbsp;&nbsp; 1092611 |
| Rio Tinto PLC | &nbsp;&nbsp; 23486 | &nbsp;&nbsp; 2178884 |
| Standard Chartered PLC | &nbsp;&nbsp; 194338 | &nbsp;&nbsp; 4049975 |
| **Total** | **Total** | &nbsp;&nbsp; **34530083** |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **United States 3.6%** | **United States 3.6%** | **United States 3.6%** |
| CRH PLC | &nbsp;&nbsp; 31618 | &nbsp;&nbsp; 3307891 |
| Sunbelt Rentals Holdings, Inc. | &nbsp;&nbsp; 52488 | &nbsp;&nbsp; 3348281 |
| **Total** | **Total** | &nbsp;&nbsp; **6656172** |
| &nbsp;&nbsp; Total Common Stocks <br>(Cost $151,714,694) | &nbsp;&nbsp; Total Common Stocks <br>(Cost $151,714,694) | &nbsp;&nbsp; **178631646** |
| **Money Market Funds 2.3%** | **Money Market Funds 2.3%** | **Money Market Funds 2.3%** |
|  | **Shares** | **Value ($)** |
| Columbia Short-Term Cash Fund, 3.790%<sup>(b),(c)</sup> <br>| &nbsp;&nbsp; 4185522 | &nbsp;&nbsp; 4183848 |
| &nbsp;&nbsp; Total Money Market Funds <br>(Cost $4,184,157) | &nbsp;&nbsp; Total Money Market Funds <br>(Cost $4,184,157) | &nbsp;&nbsp; **4183848** |
| **Total Investments in Securities** <br>**(Cost $155,898,851)** | **Total Investments in Securities** <br>**(Cost $155,898,851)** | &nbsp;&nbsp; **182815494** |
| **Other Assets & Liabilities, Net** |  | &nbsp;&nbsp; **1459848** |
| **Net Assets** | **Net Assets** | &nbsp;&nbsp; **$184275342** |

---

**Notes to Portfolio of Investments** 

&nbsp;&nbsp;&nbsp;&nbsp;(a) Non-income producing investment.

&nbsp;&nbsp;&nbsp;&nbsp;(b) The rate shown is the seven-day current annualized yield at March 31, 2026.

&nbsp;&nbsp;&nbsp;&nbsp;(c) Under the Investment Company Act of 1940, an affiliated company is one in which the Fund owns 5% or more of the company's outstanding voting securities, or a company which is under common ownership or control with the Fund. The value of the
 holdings and transactions in these affiliated companies during the period ended March 31, 2026 are as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Affiliated issuers** | **Beginning** <br>**of period($)**<br>| **Purchases($)** | **Sales($)** | **Net change in** <br>**unrealized** <br>**appreciation** <br>**(depreciation)($)**<br>| **End of** <br>**period($)**<br>| **Realized gain** <br>**(loss)($)**<br>| **Dividends($)** | **End of** <br>**period shares**<br>|
| Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% |
|  | 2432784 | &nbsp;&nbsp; 17846807 | &nbsp;&nbsp; (16095404)<br>| &nbsp;&nbsp; (339)<br>| &nbsp;&nbsp; 4183848 | &nbsp;&nbsp; 556 | &nbsp;&nbsp; 25596 | &nbsp;&nbsp; 4185522 |

---

Investments are valued using policies described in the Notes to Financial Statements included within the most recent financial statements and additional information.

Columbia Acorn International Select<sup>SM</sup> \| 2026

------

Portfolio of Investments

Columbia Thermostat Fund<sup>SM</sup>, March 31, 2026 (Unaudited)

(Percentages represent value of investments compared to net assets)

**Investments in securities**

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Equity Funds 27.9%** | **Equity Funds 27.9%** | **Equity Funds 27.9%** |
|  | **Shares** | **Value ($)** |
| **U.S. Large Cap 27.9%** | **U.S. Large Cap 27.9%** | **U.S. Large Cap 27.9%** |
| &nbsp;&nbsp; Columbia Contrarian Core Fund, Institutional 3 <br> Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 2907531 | &nbsp;&nbsp; 109672067 |
| &nbsp;&nbsp; Columbia Cornerstone Growth Fund, Institutional <br> 3 Class<sup>(a),(b)</sup> <br>| &nbsp;&nbsp; 587851 | &nbsp;&nbsp; 44094728 |
| &nbsp;&nbsp; Columbia Large Cap Enhanced Core Fund, <br> Institutional 3 Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 4075657 | &nbsp;&nbsp; 109961225 |
| &nbsp;&nbsp; Columbia Large Cap Index Fund, Institutional 3 <br> Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 1766694 | &nbsp;&nbsp; 87822376 |
| **Total** | **Total** | &nbsp;&nbsp; **351550396** |
| &nbsp;&nbsp; Total Equity Funds <br>(Cost $316,007,299) | &nbsp;&nbsp; Total Equity Funds <br>(Cost $316,007,299) | &nbsp;&nbsp; **351550396** |
| **Exchange-Traded Equity Funds 7.0%** | **Exchange-Traded Equity Funds 7.0%** | **Exchange-Traded Equity Funds 7.0%** |
| **U.S. Large Cap 7.0%** | **U.S. Large Cap 7.0%** | **U.S. Large Cap 7.0%** |
| Columbia Research Enhanced Core ETF<sup>(a)</sup> <br>| &nbsp;&nbsp; 2245815 | &nbsp;&nbsp; 87541869 |
| &nbsp;&nbsp; Total Exchange-Traded Equity Funds <br>(Cost $72,933,543) | &nbsp;&nbsp; Total Exchange-Traded Equity Funds <br>(Cost $72,933,543) | &nbsp;&nbsp; **87541869** |
| **Exchange-Traded Fixed Income Funds 6.4%** | **Exchange-Traded Fixed Income Funds 6.4%** | **Exchange-Traded Fixed Income Funds 6.4%** |
| **Multisector 6.4%** | **Multisector 6.4%** | **Multisector 6.4%** |
| &nbsp;&nbsp; Columbia Diversified Fixed Income Allocation <br> ETF<sup>(a)</sup> <br>| &nbsp;&nbsp; 4487194 | &nbsp;&nbsp; 81128467 |
| &nbsp;&nbsp; Total Exchange-Traded Fixed Income Funds <br>(Cost $81,418,234) | &nbsp;&nbsp; Total Exchange-Traded Fixed Income Funds <br>(Cost $81,418,234) | &nbsp;&nbsp; **81128467** |

---

---

| | | |
|:---|:---|:---|
| **Fixed Income Funds 58.2%** | **Fixed Income Funds 58.2%** | **Fixed Income Funds 58.2%** |
|  | **Shares** | **Value ($)** |
| **High Yield 6.5%** | **High Yield 6.5%** | **High Yield 6.5%** |
| &nbsp;&nbsp; Columbia High Yield Bond Fund, Institutional 3 <br> Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 7397286 | &nbsp;&nbsp; 81296177 |
| **Investment Grade 51.7%** | **Investment Grade 51.7%** | **Investment Grade 51.7%** |
| &nbsp;&nbsp; Columbia Quality Income Fund, Institutional 3 <br> Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 6760368 | &nbsp;&nbsp; 122633072 |
| &nbsp;&nbsp; Columbia Select Corporate Income Fund, <br> Institutional 3 Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 17866160 | &nbsp;&nbsp; 163118039 |
| &nbsp;&nbsp; Columbia Total Return Bond Fund, Institutional 3 <br> Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 6667403 | &nbsp;&nbsp; 203822495 |
| &nbsp;&nbsp; Columbia U.S. Treasury Index Fund, Institutional <br> 3 Class<sup>(a)</sup> <br>| &nbsp;&nbsp; 16372094 | &nbsp;&nbsp; 162902339 |
| **Total** | **Total** | &nbsp;&nbsp; **652475945** |
| &nbsp;&nbsp; Total Fixed Income Funds <br>(Cost $716,854,863) | &nbsp;&nbsp; Total Fixed Income Funds <br>(Cost $716,854,863) | &nbsp;&nbsp; **733772122** |
| **Money Market Funds 0.5%** | **Money Market Funds 0.5%** | **Money Market Funds 0.5%** |
| Columbia Short-Term Cash Fund, 3.790%<sup>(a),(c)</sup> <br>| &nbsp;&nbsp; 6775013 | &nbsp;&nbsp; 6772303 |
| &nbsp;&nbsp; Total Money Market Funds <br>(Cost $6,772,408) | &nbsp;&nbsp; Total Money Market Funds <br>(Cost $6,772,408) | &nbsp;&nbsp; **6772303** |
| **Total Investments in Securities** <br>**(Cost: $1,193,986,347)** | **Total Investments in Securities** <br>**(Cost: $1,193,986,347)** | &nbsp;&nbsp; **1260765157** |
| **Other Assets & Liabilities, Net** |  | &nbsp;&nbsp; **500789** |
| **Net Assets** | **Net Assets** | &nbsp;&nbsp; **1261265946** |

---

**Notes to Portfolio of Investments** 

&nbsp;&nbsp;&nbsp;&nbsp;(a) Under the Investment Company Act of 1940, an affiliated company is one in which the Fund owns 5% or more of the company's outstanding voting securities, or a company which is under common ownership or control with the Fund. The value of the
 holdings and transactions in these affiliated companies during the period ended March 31, 2026 are as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Affiliated issuers** | **Beginning** <br>**of period($)**<br>| **Purchases($)** | **Sales($)** | **Net change in** <br>**unrealized** <br>**appreciation** <br>**(depreciation)($)**<br>| **End of** <br>**period($)**<br>| **Realized gain** <br>**(loss)($)**<br>| **Dividends($)** | **End of** <br>**period shares**<br>|
| Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class | Columbia Contrarian Core Fund, Institutional 3 Class |
|  | 97651520 | &nbsp;&nbsp; 34960524 | &nbsp;&nbsp; (18080618)<br>| &nbsp;&nbsp; (4859359)<br>| &nbsp;&nbsp; 109672067 | &nbsp;&nbsp; (62084)<br>| &nbsp;&nbsp; — | &nbsp;&nbsp; 2907531 |
| Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class | Columbia Cornerstone Growth Fund, Institutional 3 Class |
|  | 39255313 | &nbsp;&nbsp; 15261770 | &nbsp;&nbsp; (7162411)<br>| &nbsp;&nbsp; (3259944)<br>| &nbsp;&nbsp; 44094728 | &nbsp;&nbsp; (172970)<br>| &nbsp;&nbsp; — | &nbsp;&nbsp; 587851 |
| Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF | Columbia Diversified Fixed Income Allocation ETF |
|  | 89709310 | &nbsp;&nbsp; 6997689 | &nbsp;&nbsp; (14404920)<br>| &nbsp;&nbsp; (1173612)<br>| &nbsp;&nbsp; 81128467 | &nbsp;&nbsp; (283664)<br>| &nbsp;&nbsp; 799536 | &nbsp;&nbsp; 4487194 |
| Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class | Columbia High Yield Bond Fund, Institutional 3 Class |
|  | 90648928 | &nbsp;&nbsp; 7239284 | &nbsp;&nbsp; (15215546)<br>| &nbsp;&nbsp; (1376489)<br>| &nbsp;&nbsp; 81296177 | &nbsp;&nbsp; (269067)<br>| &nbsp;&nbsp; 1436032 | &nbsp;&nbsp; 7397286 |

---

Columbia Thermostat Fund<sup>SM</sup> \| 2026

------

Portfolio of Investments (continued)

Columbia Thermostat Fund<sup>SM</sup>, March 31, 2026 (Unaudited)

**Notes to Portfolio of Investments (continued)**

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Affiliated issuers** | **Beginning**<br> **of period($)**<br>| **Purchases($)** | **Sales($)** | **Net change in** <br> **unrealized** <br> **appreciation** <br> **(depreciation)($)**<br>| **End of** <br> **period($)**<br>| **Realized gain**<br> **(loss)($)**<br>| **Dividends($)** | **End of**<br> **period shares**<br>|
| Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class | Columbia Large Cap Enhanced Core Fund, Institutional 3 Class |
|  | 97597120 | &nbsp;&nbsp; 35228683 | &nbsp;&nbsp; (18412492)<br>| &nbsp;&nbsp; (4452086)<br>| &nbsp;&nbsp; 109961225 | &nbsp;&nbsp; (114020)<br>| &nbsp;&nbsp; — | &nbsp;&nbsp; 4075657 |
| Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class | Columbia Large Cap Index Fund, Institutional 3 Class |
|  | 77979626 | &nbsp;&nbsp; 27819515 | &nbsp;&nbsp; (14996151)<br>| &nbsp;&nbsp; (2980614)<br>| &nbsp;&nbsp; 87822376 | &nbsp;&nbsp; (62135)<br>| &nbsp;&nbsp; — | &nbsp;&nbsp; 1766694 |
| Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class | Columbia Quality Income Fund, Institutional 3 Class |
|  | 134926489 | &nbsp;&nbsp; 11986143 | &nbsp;&nbsp; (23356286)<br>| &nbsp;&nbsp; (923274)<br>| &nbsp;&nbsp; 122633072 | &nbsp;&nbsp; 64440 | &nbsp;&nbsp; 1452469 | &nbsp;&nbsp; 6760368 |
| Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF | Columbia Research Enhanced Core ETF |
|  | 77584688 | &nbsp;&nbsp; 27385576 | &nbsp;&nbsp; (14150554)<br>| &nbsp;&nbsp; (3277841)<br>| &nbsp;&nbsp; 87541869 | &nbsp;&nbsp; 390611 | &nbsp;&nbsp; — | &nbsp;&nbsp; 2245815 |
| Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class | Columbia Select Corporate Income Fund, Institutional 3 Class |
|  | 180248809 | &nbsp;&nbsp; 15265284 | &nbsp;&nbsp; (31579683)<br>| &nbsp;&nbsp; (816371)<br>| &nbsp;&nbsp; 163118039 | &nbsp;&nbsp; (1748372)<br>| &nbsp;&nbsp; 2155735 | &nbsp;&nbsp; 17866160 |
| Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% |
|  | 6220261 | &nbsp;&nbsp; 20997882 | &nbsp;&nbsp; (20445420)<br>| &nbsp;&nbsp; (420)<br>| &nbsp;&nbsp; 6772303 | &nbsp;&nbsp; 50 | &nbsp;&nbsp; 47161 | &nbsp;&nbsp; 6775013 |
| Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class | Columbia Total Return Bond Fund, Institutional 3 Class |
|  | 225428660 | &nbsp;&nbsp; 19764450 | &nbsp;&nbsp; (38847707)<br>| &nbsp;&nbsp; (2522908)<br>| &nbsp;&nbsp; 203822495 | &nbsp;&nbsp; (104258)<br>| &nbsp;&nbsp; 2741823 | &nbsp;&nbsp; 6667403 |
| Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class | Columbia U.S. Treasury Index Fund, Institutional 3 Class |
|  | 179934702 | &nbsp;&nbsp; 15661956 | &nbsp;&nbsp; (31152943)<br>| &nbsp;&nbsp; (1541376)<br>| &nbsp;&nbsp; 162902339 | &nbsp;&nbsp; (5300)<br>| &nbsp;&nbsp; 1685211 | &nbsp;&nbsp; 16372094 |
| Total | 1297185426 |  |  | &nbsp;&nbsp; (27184294)<br>| &nbsp;&nbsp; 1260765157 | &nbsp;&nbsp; (2366769)<br>| &nbsp;&nbsp; 10317967 |  |

---

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;(b) Non-income producing investment.

&nbsp;&nbsp;&nbsp;&nbsp;(c) The rate shown is the seven-day current annualized yield at March 31, 2026.

Investments are valued using policies described in the Notes to Financial Statements included within the most recent financial statements and additional information.

Columbia Thermostat Fund<sup>SM</sup> \| 2026

------

Portfolio of Investments

Columbia Acorn European Fund<sup>SM</sup>, March 31, 2026 (Unaudited)

(Percentages represent value of investments compared to net assets)

**Investments in securities**

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Common Stocks 98.1%** | **Common Stocks 98.1%** | **Common Stocks 98.1%** |
| **Issuer** | **Shares** | **Value ($)** |
| **Austria 3.0%** | **Austria 3.0%** | **Austria 3.0%** |
| BAWAG Group AG | &nbsp;&nbsp; 1258 | &nbsp;&nbsp; 191251 |
| DO & CO AG | &nbsp;&nbsp; 2492 | &nbsp;&nbsp; 466621 |
| Wienerberger AG | &nbsp;&nbsp; 12494 | &nbsp;&nbsp; 332266 |
| **Total** | **Total** | &nbsp;&nbsp; **990138** |
| **Belgium 1.4%** | **Belgium 1.4%** | **Belgium 1.4%** |
| Cenergy Holdings SA | &nbsp;&nbsp; 8678 | &nbsp;&nbsp; 190872 |
| Warehouses De Pauw CVA | &nbsp;&nbsp; 10879 | &nbsp;&nbsp; 283394 |
| **Total** | **Total** | &nbsp;&nbsp; **474266** |
| **Denmark 2.9%** | **Denmark 2.9%** | **Denmark 2.9%** |
| FLSmidth & Co. A/S | &nbsp;&nbsp; 3250 | &nbsp;&nbsp; 246209 |
| NKT A/S<sup>(a)</sup> <br>| &nbsp;&nbsp; 3759 | &nbsp;&nbsp; 489350 |
| Per Aarsleff Holding A/S | &nbsp;&nbsp; 2044 | &nbsp;&nbsp; 236681 |
| **Total** | **Total** | &nbsp;&nbsp; **972240** |
| **Finland 2.0%** | **Finland 2.0%** | **Finland 2.0%** |
| Cargotec OYJ, Class B | &nbsp;&nbsp; 3490 | &nbsp;&nbsp; 165037 |
| Konecranes OYJ | &nbsp;&nbsp; 9642 | &nbsp;&nbsp; 316457 |
| Valmet OYJ | &nbsp;&nbsp; 6095 | &nbsp;&nbsp; 174400 |
| **Total** | **Total** | &nbsp;&nbsp; **655894** |
| **France 16.2%** | **France 16.2%** | **France 16.2%** |
| Elis SA | &nbsp;&nbsp; 35813 | &nbsp;&nbsp; 1014168 |
| Exosens SAS | &nbsp;&nbsp; 3275 | &nbsp;&nbsp; 235828 |
| Gaztransport Et Technigaz SA | &nbsp;&nbsp; 1653 | &nbsp;&nbsp; 388775 |
| Lisi SA | &nbsp;&nbsp; 6279 | &nbsp;&nbsp; 383915 |
| Nexans SA | &nbsp;&nbsp; 5351 | &nbsp;&nbsp; 727944 |
| SCOR SE | &nbsp;&nbsp; 16193 | &nbsp;&nbsp; 578744 |
| SPIE SA | &nbsp;&nbsp; 14283 | &nbsp;&nbsp; 715230 |
| Technip Energies NV | &nbsp;&nbsp; 16727 | &nbsp;&nbsp; 706842 |
| Vallourec SACA | &nbsp;&nbsp; 7926 | &nbsp;&nbsp; 200636 |
| Virbac SA | &nbsp;&nbsp; 1092 | &nbsp;&nbsp; 451011 |
| **Total** | **Total** | &nbsp;&nbsp; **5403093** |
| **Germany 9.2%** | **Germany 9.2%** | **Germany 9.2%** |
| AlzChem Group AG | &nbsp;&nbsp; 1303 | &nbsp;&nbsp; 260789 |
| Bilfinger SE | &nbsp;&nbsp; 3898 | &nbsp;&nbsp; 451350 |
| CTS Eventim AG & Co. KGaA | &nbsp;&nbsp; 1991 | &nbsp;&nbsp; 116614 |
| DEUTZ AG | &nbsp;&nbsp; 18207 | &nbsp;&nbsp; 183060 |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| flatexDEGIRO AG | &nbsp;&nbsp; 17560 | &nbsp;&nbsp; 605563 |
| JOST Werke AG | &nbsp;&nbsp; 2575 | &nbsp;&nbsp; 149942 |
| Nordex SE<sup>(a)</sup> <br>| &nbsp;&nbsp; 4990 | &nbsp;&nbsp; 270420 |
| PVA TePla AG<sup>(a)</sup> <br>| &nbsp;&nbsp; 4693 | &nbsp;&nbsp; 162411 |
| Renk Group AG | &nbsp;&nbsp; 4562 | &nbsp;&nbsp; 272679 |
| SUSS MicroTec SE | &nbsp;&nbsp; 6973 | &nbsp;&nbsp; 414683 |
| Vincorion SE<sup>(a)</sup> <br>| &nbsp;&nbsp; 9150 | &nbsp;&nbsp; 171649 |
| **Total** | **Total** | &nbsp;&nbsp; **3059160** |
| **Greece 2.5%** | **Greece 2.5%** | **Greece 2.5%** |
| Alpha Bank SA | &nbsp;&nbsp; 108416 | &nbsp;&nbsp; 402150 |
| Piraeus Bank SA<sup>(a)</sup> <br>| &nbsp;&nbsp; 52189 | &nbsp;&nbsp; 428785 |
| **Total** | **Total** | &nbsp;&nbsp; **830935** |
| **Ireland 7.5%** | **Ireland 7.5%** | **Ireland 7.5%** |
| Cairn Homes PLC | &nbsp;&nbsp; 207880 | &nbsp;&nbsp; 511190 |
| Glanbia PLC | &nbsp;&nbsp; 36364 | &nbsp;&nbsp; 712431 |
| Irish Continental Group PLC | &nbsp;&nbsp; 56644 | &nbsp;&nbsp; 415092 |
| Permanent TSB Group Holdings PLC<sup>(a)</sup> <br>| &nbsp;&nbsp; 53289 | &nbsp;&nbsp; 178700 |
| Uniphar PLC | &nbsp;&nbsp; 156292 | &nbsp;&nbsp; 697461 |
| **Total** | **Total** | &nbsp;&nbsp; **2514874** |
| **Italy 5.7%** | **Italy 5.7%** | **Italy 5.7%** |
| Buzzi SpA | &nbsp;&nbsp; 4748 | &nbsp;&nbsp; 240045 |
| Carel Industries SpA | &nbsp;&nbsp; 6596 | &nbsp;&nbsp; 168109 |
| De' Longhi SpA | &nbsp;&nbsp; 10153 | &nbsp;&nbsp; 356829 |
| Hera | &nbsp;&nbsp; 149931 | &nbsp;&nbsp; 691584 |
| Intercos SpA | &nbsp;&nbsp; 13946 | &nbsp;&nbsp; 199881 |
| Reply SpA | &nbsp;&nbsp; 1866 | &nbsp;&nbsp; 175608 |
| Sol Spa | &nbsp;&nbsp; 1220 | &nbsp;&nbsp; 83057 |
| **Total** | **Total** | &nbsp;&nbsp; **1915113** |
| **Luxembourg 0.6%** | **Luxembourg 0.6%** | **Luxembourg 0.6%** |
| Tonies SE, Class A<sup>(a)</sup> <br>| &nbsp;&nbsp; 16151 | &nbsp;&nbsp; 192671 |
| **Netherlands 2.8%** | **Netherlands 2.8%** | **Netherlands 2.8%** |
| CTP NV<sup>(b)</sup> <br>| &nbsp;&nbsp; 30147 | &nbsp;&nbsp; 504555 |
| Van Lanschot Kempen NV | &nbsp;&nbsp; 6287 | &nbsp;&nbsp; 417524 |
| **Total** | **Total** | &nbsp;&nbsp; **922079** |

---

Columbia Acorn European Fund<sup>SM</sup> \| 2026

------

Portfolio of Investments (continued)

Columbia Acorn European Fund<sup>SM</sup>, March 31, 2026 (Unaudited)

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| **Norway 3.1%** | **Norway 3.1%** | **Norway 3.1%** |
| SalMar ASA | &nbsp;&nbsp; 5914 | &nbsp;&nbsp; 345084 |
| Storebrand ASA | &nbsp;&nbsp; 38943 | &nbsp;&nbsp; 702912 |
| **Total** | **Total** | &nbsp;&nbsp; **1047996** |
| **Portugal 1.0%** | **Portugal 1.0%** | **Portugal 1.0%** |
| Galp Energia SGPS SA | &nbsp;&nbsp; 14587 | &nbsp;&nbsp; 349761 |
| **Spain 2.2%** | **Spain 2.2%** | **Spain 2.2%** |
| Fluidra SA | &nbsp;&nbsp; 12694 | &nbsp;&nbsp; 294452 |
| Merlin Properties Socimi SA | &nbsp;&nbsp; 14446 | &nbsp;&nbsp; 235359 |
| Viscofan SA | &nbsp;&nbsp; 2852 | &nbsp;&nbsp; 198596 |
| **Total** | **Total** | &nbsp;&nbsp; **728407** |
| **Sweden 2.6%** | **Sweden 2.6%** | **Sweden 2.6%** |
| Hexpol AB | &nbsp;&nbsp; 22265 | &nbsp;&nbsp; 169841 |
| Nordnet AB | &nbsp;&nbsp; 10816 | &nbsp;&nbsp; 352215 |
| SSAB AB, Class B | &nbsp;&nbsp; 42332 | &nbsp;&nbsp; 333708 |
| **Total** | **Total** | &nbsp;&nbsp; **855764** |
| **Switzerland 5.2%** | **Switzerland 5.2%** | **Switzerland 5.2%** |
| Burckhardt Compression Holding AG | &nbsp;&nbsp; 398 | &nbsp;&nbsp; 238211 |
| Cicor Technologies Ltd.<sup>(a)</sup> <br>| &nbsp;&nbsp; 690 | &nbsp;&nbsp; 102548 |
| Comet Holding AG, Registered Shares | &nbsp;&nbsp; 1047 | &nbsp;&nbsp; 333898 |
| Galenica AG | &nbsp;&nbsp; 2895 | &nbsp;&nbsp; 329590 |
| Huber + Suhner AG, Registered Shares | &nbsp;&nbsp; 1111 | &nbsp;&nbsp; 248161 |
| Kardex Holding AG | &nbsp;&nbsp; 884 | &nbsp;&nbsp; 265973 |
| R&S Group Holding AG<sup>(a)</sup> <br>| &nbsp;&nbsp; 8852 | &nbsp;&nbsp; 229167 |
| **Total** | **Total** | &nbsp;&nbsp; **1747548** |
| **United Kingdom 29.7%** | **United Kingdom 29.7%** | **United Kingdom 29.7%** |
| Ashmore Group PLC | &nbsp;&nbsp; 68281 | &nbsp;&nbsp; 191056 |
| Avon Technologies PLC | &nbsp;&nbsp; 8193 | &nbsp;&nbsp; 179798 |
| Babcock International Group PLC | &nbsp;&nbsp; 24931 | &nbsp;&nbsp; 387044 |
| Breedon Group PLC | &nbsp;&nbsp; 63212 | &nbsp;&nbsp; 249329 |
| Bridgepoint Group Ltd., Registered Shares<sup>(b)</sup> <br>| &nbsp;&nbsp; 61540 | &nbsp;&nbsp; 191741 |
| Capital & Counties Properties PLC | &nbsp;&nbsp; 244237 | &nbsp;&nbsp; 412172 |
| Chemring Group PLC | &nbsp;&nbsp; 66020 | &nbsp;&nbsp; 450132 |
| Clarkson PLC | &nbsp;&nbsp; 4702 | &nbsp;&nbsp; 289201 |
| Cohort PLC | &nbsp;&nbsp; 15558 | &nbsp;&nbsp; 252023 |
| ConvaTec Group PLC | &nbsp;&nbsp; 96569 | &nbsp;&nbsp; 278547 |
| Croda International PLC | &nbsp;&nbsp; 7481 | &nbsp;&nbsp; 280932 |

---

---

| | | |
|:---|:---|:---|
| **Common Stocks (continued)** | **Common Stocks (continued)** | **Common Stocks (continued)** |
| **Issuer** | **Shares** | **Value ($)** |
| CVS Group PLC | &nbsp;&nbsp; 45389 | &nbsp;&nbsp; 678444 |
| discoverIE Group PLC | &nbsp;&nbsp; 27711 | &nbsp;&nbsp; 197329 |
| Endeavour Mining PLC | &nbsp;&nbsp; 3150 | &nbsp;&nbsp; 189787 |
| Genus PLC | &nbsp;&nbsp; 6183 | &nbsp;&nbsp; 196072 |
| ICG PLC | &nbsp;&nbsp; 8273 | &nbsp;&nbsp; 169673 |
| IG Group Holdings PLC | &nbsp;&nbsp; 27224 | &nbsp;&nbsp; 518412 |
| IMI PLC | &nbsp;&nbsp; 9205 | &nbsp;&nbsp; 313145 |
| Keller Group PLC | &nbsp;&nbsp; 8648 | &nbsp;&nbsp; 220002 |
| LondonMetric Property PLC | &nbsp;&nbsp; 244406 | &nbsp;&nbsp; 590386 |
| Moonpig Group PLC | &nbsp;&nbsp; 190740 | &nbsp;&nbsp; 528911 |
| Quilter PLC<sup>(b)</sup> <br>| &nbsp;&nbsp; 342811 | &nbsp;&nbsp; 792238 |
| Rotork PLC | &nbsp;&nbsp; 81821 | &nbsp;&nbsp; 340223 |
| Safestore Holdings PLC | &nbsp;&nbsp; 64016 | &nbsp;&nbsp; 537622 |
| Shawbrook Group PLC<sup>(a),(b)</sup> <br>| &nbsp;&nbsp; 91314 | &nbsp;&nbsp; 386762 |
| SigmaRoc PLC<sup>(a)</sup> <br>| &nbsp;&nbsp; 262016 | &nbsp;&nbsp; 408706 |
| Subsea 7 SA | &nbsp;&nbsp; 13087 | &nbsp;&nbsp; 405419 |
| WPP PLC | &nbsp;&nbsp; 89170 | &nbsp;&nbsp; 278876 |
| **Total** | **Total** | &nbsp;&nbsp; **9913982** |
| **United States 0.5%** | **United States 0.5%** | **United States 0.5%** |
| Boku, Inc.<sup>(a)</sup> <br>| &nbsp;&nbsp; 70611 | &nbsp;&nbsp; 153743 |
| &nbsp;&nbsp; Total Common Stocks <br>(Cost $30,061,070) | &nbsp;&nbsp; Total Common Stocks <br>(Cost $30,061,070) | &nbsp;&nbsp; **32727664** |

---

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Preferred Stocks 0.5%** | **Preferred Stocks 0.5%** | **Preferred Stocks 0.5%** |
| **Issuer** | **Shares** | **Value ($)** |
| **Germany 0.5%** | **Germany 0.5%** | **Germany 0.5%** |
| KSB SE & Co. KGAA | &nbsp;&nbsp; 164 | &nbsp;&nbsp; 174772 |
| &nbsp;&nbsp; Total Preferred Stocks <br>(Cost $212,981) | &nbsp;&nbsp; Total Preferred Stocks <br>(Cost $212,981) | &nbsp;&nbsp; **174772** |

---

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | |
|:---|:---|:---|
| **Money Market Funds 0.8%** | **Money Market Funds 0.8%** | **Money Market Funds 0.8%** |
|  | **Shares** | **Value ($)** |
| Columbia Short-Term Cash Fund, 3.790%<sup>(c),(d)</sup> <br>| &nbsp;&nbsp; 275393 | &nbsp;&nbsp; 275283 |
| &nbsp;&nbsp; Total Money Market Funds <br>(Cost $275,271) | &nbsp;&nbsp; Total Money Market Funds <br>(Cost $275,271) | &nbsp;&nbsp; **275283** |
| **Total Investments in Securities** <br>**(Cost $30,549,322)** | **Total Investments in Securities** <br>**(Cost $30,549,322)** | &nbsp;&nbsp; **33177719** |
| **Other Assets & Liabilities, Net** |  | &nbsp;&nbsp; **194593** |
| **Net Assets** | **Net Assets** | &nbsp;&nbsp; **$33372312** |

---

Columbia Acorn European Fund<sup>SM</sup> \| 2026

------

Portfolio of Investments (continued)

Columbia Acorn European Fund<sup>SM</sup>, March 31, 2026 (Unaudited)

**Notes to Portfolio of Investments** 

&nbsp;&nbsp;&nbsp;&nbsp;(a) Non-income producing investment.

&nbsp;&nbsp;&nbsp;&nbsp;(b) Represents privately placed and other securities and instruments exempt from Securities
 and Exchange Commission registration (collectively, private placements), such as Section 4(a)(2) and Rule 144A eligible securities, which are often sold only
 to qualified institutional buyers. At March 31, 2026, the total value of these securities amounted to $1,875,296, which represents 5.62% of total net assets.

&nbsp;&nbsp;&nbsp;&nbsp;(c) The rate shown is the seven-day current annualized yield at March 31, 2026.

&nbsp;&nbsp;&nbsp;&nbsp;(d) Under the Investment Company Act of 1940, an affiliated company is one in which the Fund owns 5% or more of the company's outstanding voting securities, or a company which is under common ownership or control with the Fund. The value of the
 holdings and transactions in these affiliated companies during the period ended March 31, 2026 are as follows:

&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;

---

| | | | | | | | | |
|:---|:---|:---|:---|:---|:---|:---|:---|:---|
| **Affiliated issuers** | **Beginning** <br>**of period($)**<br>| **Purchases($)** | **Sales($)** | **Net change in** <br>**unrealized** <br>**appreciation** <br>**(depreciation)($)**<br>| **End of** <br>**period($)**<br>| **Realized gain** <br>**(loss)($)**<br>| **Dividends($)** | **End of** <br>**period shares**<br>|
| Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% | Columbia Short-Term Cash Fund, 3.790% |
|  | 326154 | &nbsp;&nbsp; 3939973 | &nbsp;&nbsp; (3990855)<br>| &nbsp;&nbsp; 11 | &nbsp;&nbsp; 275283 | &nbsp;&nbsp; 6 | &nbsp;&nbsp; 2854 | &nbsp;&nbsp; 275393 |

---

Investments are valued using policies described in the Notes to Financial Statements included within the most recent financial statements and additional information.

Columbia Acorn European Fund<sup>SM</sup> \| 2026

------

[THIS PAGE INTENTIONALLY LEFT BLANK]

------

You may at any time request, free of charge, to receive a paper copy of this report by calling 800.345.6611.

1QT110_(05/26)

------

## Form NPORT-P: Monthly Portfolio Investments Report

### NPORT-P: Part A: General Information

**Item A.1. Information about the Registrant.**

- **a. Name of Registrant:** COLUMBIA ACORN TRUST

- **b. Investment Company Act file number:** 811-01829

- **c. CIK number of Registrant:** 0000002110

- **d. LEI of Registrant:** 549300DX0CWIN8VE1908

- **e. Address and telephone number of Registrant.**

  - **Street Address 1:** 290 Congress Street

  - **City:** Boston

  - **State:** MA

  - **Foreign country:** US

  - **Zip / Postal Code:** 02210

  - **Telephone number:** 1800-345-6611

**Item A.2. Information about the Series.**

- **a. Name of Series:** Columbia Acorn Fund

- **b. EDGAR series identifier (if any):** S000009184

- **c. LEI of Series:** 549300X3KJ5X2R54O387

**Item A.3. Reporting period.**

- **a. Date of fiscal year-end:** 2026-12-31

- **b. Date as of which information is reported:** 2026-03-31

**Item A.4. Final filing**

Does the Fund anticipate that this will be its final filing on Form N-PORT? **No**

### Fund Information

**Total Assets:** $2249862358.60

**Total Liabilities:** $4399447.15

**Net Assets:** $2245462911.45

**Cash Not Reported:** $20.71

**Monthly Return Information**

| Class               | Month 1 Return (%)   | Month 2 Return (%)   | Month 3 Return (%)   |
|:---|:---|:---|:---|
| Class ID C000024954 | 4.81%                | -0.32%               | -8.78%               |
| Class ID C000024956 | 4.75%                | -0.47%               | -8.79%               |
| Class ID C000024957 | 4.81%                | -0.29%               | -8.76%               |
| Class ID C000122736 | 4.83%                | -0.33%               | -8.79%               |
| Class ID C000122737 | 4.81%                | -0.32%               | -8.76%               |
| Class ID C000251059 | 4.73%                | -0.29%               | -8.76%               |

**Monthly Gains & Losses**

| Period   | Net Realized Gain/Loss   | Net Unrealized Appreciation/Depreciation   |
|:---|:---|:---|
| Month 1  | $43703763.87             | $74586173.39                               |
| Month 2  | $19470405.00             | $-27752986.20                              |
| Month 3  | $-23618025.48            | $-193217801.26                             |

### Schedule of Portfolio Investments

| Name                                         | Title                                        | Identifiers                                   | Payoff Profile   | Asset Category   | Issuer Category   | Country   |   Balance | Units   | Value (USD)   | % of Net Assets   | Maturity Date   | Coupon Type   | Annualized Rate (%)   | Restricted?   |   Fair Value Level | Lending Status   |
|:---|:---|:---|:---|:---|:---|:---|---:|:---|:---|:---|:---|:---|:---|:---|---:|:---|
| Absci Corp                                   | Absci Corp                                   | CUSIP: 00091E109<br>LEI: N/A                  | Long             | EC               | CORP              | US        |   2566842 | NS      | $7700526.00   | 0.34%             |  |  |  | No            |                  1 | On Loan: No      |
| Adaptive Biotechnologies Corp                | Adaptive Biotechnologies Corp                | CUSIP: 00650F109<br>LEI: 549300ZTF7OT1FW66Q96 | Long             | EC               | CORP              | US        |   1148404 | NS      | $15939847.52  | 0.71%             |  |  |  | No            |                  1 | On Loan: No      |
| Advanced Energy Industries Inc               | Advanced Energy Industries Inc               | CUSIP: 007973100<br>LEI: 549300756XI3QLFT2U27 | Long             | EC               | CORP              | US        |     28789 | NS      | $9290498.19   | 0.41%             |  |  |  | No            |                  1 | On Loan: No      |
| AeroVironment Inc                            | AeroVironment Inc                            | CUSIP: 008073108<br>LEI: 529900PGHVI29YLZ6221 | Long             | EC               | CORP              | US        |    153583 | NS      | $28113368.15  | 1.25%             |  |  |  | No            |                  1 | On Loan: No      |
| Alignment Healthcare Inc                     | Alignment Healthcare Inc                     | CUSIP: 01625V104<br>LEI: N/A                  | Long             | EC               | CORP              | US        |   1618094 | NS      | $28510816.28  | 1.27%             |  |  |  | No            |                  1 | On Loan: No      |
| Annexon Inc                                  | Annexon Inc                                  | CUSIP: 03589W102<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    975637 | NS      | $5405028.98   | 0.24%             |  |  |  | No            |                  1 | On Loan: No      |
| Antero Resources Corp                        | Antero Resources Corp                        | CUSIP: 03674X106<br>LEI: APVBKPOULHUX7YZLKU17 | Long             | EC               | CORP              | US        |    355938 | NS      | $15106008.72  | 0.67%             |  |  |  | No            |                  1 | On Loan: No      |
| Applied Digital Corp                         | Applied Digital Corp                         | CUSIP: 038169207<br>LEI: 2549004NE2B6RYXINZ02 | Long             | EC               | CORP              | US        |    439896 | NS      | $10443131.04  | 0.47%             |  |  |  | No            |                  1 | On Loan: No      |
| Applied Industrial Technologies Inc          | Applied Industrial Technologies Inc          | CUSIP: 03820C105<br>LEI: 549300NU173IJRC6PO38 | Long             | EC               | CORP              | US        |     84867 | NS      | $22516912.44  | 1.00%             |  |  |  | No            |                  1 | On Loan: No      |
| Astronics Corp                               | Astronics Corp                               | CUSIP: 046433108<br>LEI: 549300NNXCF8H4MSMK17 | Long             | EC               | CORP              | US        |    340163 | NS      | $22699076.99  | 1.01%             |  |  |  | No            |                  1 | On Loan: No      |
| Axon Enterprise Inc                          | Axon Enterprise Inc                          | CUSIP: 05464C101<br>LEI: 549300QP2IEEGFE16681 | Long             | EC               | CORP              | US        |     61466 | NS      | $26103995.54  | 1.16%             |  |  |  | No            |                  1 | On Loan: No      |
| BJ's Wholesale Club Holdings Inc             | BJ's Wholesale Club Holdings Inc             | CUSIP: 05550J101<br>LEI: 549300UCIWH1C93V0J03 | Long             | EC               | CORP              | US        |    350621 | NS      | $34508118.82  | 1.54%             |  |  |  | No            |                  1 | On Loan: No      |
| Billiontoone Inc                             | Billiontoone Inc                             | CUSIP: 090168105<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    166205 | NS      | $13120222.70  | 0.58%             |  |  |  | No            |                  1 | On Loan: No      |
| BioLife Solutions Inc                        | BioLife Solutions Inc                        | CUSIP: 09062W204<br>LEI: 549300G2IVCMC2TJDW72 | Long             | EC               | CORP              | US        |    696462 | NS      | $13288494.96  | 0.59%             |  |  |  | No            |                  1 | On Loan: No      |
| Bloom Energy Corp                            | Bloom Energy Corp                            | CUSIP: 093712107<br>LEI: 549300XB5J9BFW5JXV97 | Long             | EC               | CORP              | US        |    288864 | NS      | $39138183.36  | 1.74%             |  |  |  | No            |                  1 | On Loan: No      |
| Blue Owl Capital Inc                         | Blue Owl Capital Inc                         | CUSIP: 09581B103<br>LEI: N/A                  | Long             | EC               | CORP              | US        |   1391042 | NS      | $12700213.46  | 0.57%             |  |  |  | No            |                  1 | On Loan: No      |
| Bridgebio Pharma Inc                         | Bridgebio Pharma Inc                         | CUSIP: 10806X102<br>LEI: 54930028YG89VTY2J114 | Long             | EC               | CORP              | US        |    316495 | NS      | $23502918.70  | 1.05%             |  |  |  | No            |                  1 | On Loan: No      |
| Carlyle Group Inc/The                        | Carlyle Group Inc/The                        | CUSIP: 14316J108<br>LEI: 529900NGGVF94UK6PG95 | Long             | EC               | CORP              | US        |    950835 | NS      | $46010905.65  | 2.05%             |  |  |  | No            |                  1 | On Loan: No      |
| Cavco Industries Inc                         | Cavco Industries Inc                         | CUSIP: 149568107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |     83379 | NS      | $40379615.91  | 1.80%             |  |  |  | No            |                  1 | On Loan: No      |
| Celestica Inc                                | Celestica Inc                                | CUSIP: 15101Q207<br>LEI: 549300J3CG5T6RZWLN53 | Long             | EC               | CORP              | CA        |    172300 | NS      | $48533464.00  | 2.16%             |  |  |  | No            |                  1 | On Loan: No      |
| Charles River Laboratories International Inc | Charles River Laboratories International Inc | CUSIP: 159864107<br>LEI: 549300BSQ0R4UZ5KX287 | Long             | EC               | CORP              | US        |     70538 | NS      | $12167805.00  | 0.54%             |  |  |  | No            |                  1 | On Loan: No      |
| Churchill Downs Inc                          | Churchill Downs Inc                          | CUSIP: 171484108<br>LEI: 529900FODTBQC8MIS994 | Long             | EC               | CORP              | US        |    539022 | NS      | $48420346.26  | 2.16%             |  |  |  | No            |                  1 | On Loan: No      |
| Coherent Corp                                | Coherent Corp                                | CUSIP: 19247G107<br>LEI: 549300O5C25A0MMHHU33 | Long             | EC               | CORP              | US        |    172879 | NS      | $41181506.59  | 1.83%             |  |  |  | No            |                  1 | On Loan: No      |
| COLUMBIA SHORT TERM CASH FUND                | COLUMBIA SHORT TERM CASH FUND                | CUSIP: 19766H239<br>LEI: N/A                  | Long             | STIV             | RF                | US        |  19982329 | NS      | $19974336.14  | 0.89%             |  |  |  | No            |                  1 | On Loan: No      |
| Crinetics Pharmaceuticals Inc                | Crinetics Pharmaceuticals Inc                | CUSIP: 22663K107<br>LEI: 549300Q1CNFXK5P31D28 | Long             | EC               | CORP              | US        |    252431 | NS      | $9168293.92   | 0.41%             |  |  |  | No            |                  1 | On Loan: No      |
| Curtiss-Wright Corp                          | Curtiss-Wright Corp                          | CUSIP: 231561101<br>LEI: TEL51ETZWP7D0ZM4X325 | Long             | EC               | CORP              | US        |     65825 | NS      | $44834724.00  | 2.00%             |  |  |  | No            |                  1 | On Loan: No      |
| Definium Therapeutics Inc                    | Definium Therapeutics Inc                    | CUSIP: 24477V105<br>LEI: 549300QY8KN1O6RPFJ13 | Long             | EC               | CORP              | CA        |    512390 | NS      | $9684171.00   | 0.43%             |  |  |  | No            |                  1 | On Loan: No      |
| Devon Energy Corp                            | Devon Energy Corp                            | CUSIP: 25179M103<br>LEI: 54930042348RKR3ZPN35 | Long             | EC               | CORP              | US        |    421587 | NS      | $21214257.84  | 0.94%             |  |  |  | No            |                  1 | On Loan: No      |
| Dorman Products Inc                          | Dorman Products Inc                          | CUSIP: 258278100<br>LEI: 2549000O0BBUX740XR59 | Long             | EC               | CORP              | US        |    344653 | NS      | $35967987.08  | 1.60%             |  |  |  | No            |                  1 | On Loan: No      |
| Doximity Inc                                 | Doximity Inc                                 | CUSIP: 26622P107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    432308 | NS      | $10072776.40  | 0.45%             |  |  |  | No            |                  1 | On Loan: No      |
| Dynatrace Inc                                | Dynatrace Inc                                | CUSIP: 268150109<br>LEI: 6488T0Z33CE9L61EX904 | Long             | EC               | CORP              | US        |    695469 | NS      | $25718443.62  | 1.15%             |  |  |  | No            |                  1 | On Loan: No      |
| elf Beauty Inc                               | elf Beauty Inc                               | CUSIP: 26856L103<br>LEI: 549300U2K9TCON25C029 | Long             | EC               | CORP              | US        |    444462 | NS      | $26938841.82  | 1.20%             |  |  |  | No            |                  1 | On Loan: No      |
| EQT Corp                                     | EQT Corp                                     | CUSIP: 26884L109<br>LEI: 4NT01YGM4X7ZX86ISY52 | Long             | EC               | CORP              | US        |    311439 | NS      | $19819977.96  | 0.88%             |  |  |  | No            |                  1 | On Loan: No      |
| Encompass Health Corp                        | Encompass Health Corp                        | CUSIP: 29261A100<br>LEI: 549300FVRPM9DRQZRG64 | Long             | EC               | CORP              | US        |    230244 | NS      | $22271502.12  | 0.99%             |  |  |  | No            |                  1 | On Loan: No      |
| Esab Corp                                    | Esab Corp                                    | CUSIP: 29605J106<br>LEI: 549300ZNV0SWBXEIWT04 | Long             | EC               | CORP              | US        |    372810 | NS      | $36035814.60  | 1.60%             |  |  |  | No            |                  1 | On Loan: No      |
| Fortrea Holdings Inc                         | Fortrea Holdings Inc                         | CUSIP: 34965K107<br>LEI: 5493000FIVFABOVCDU97 | Long             | EC               | CORP              | US        |   1173131 | NS      | $11050894.02  | 0.49%             |  |  |  | No            |                  1 | On Loan: No      |
| GCM Grosvenor Inc                            | GCM Grosvenor Inc                            | CUSIP: 36831E108<br>LEI: N/A                  | Long             | EC               | CORP              | US        |   4428750 | NS      | $43401750.00  | 1.93%             |  |  |  | No            |                  1 | On Loan: No      |
| Glaukos Corp                                 | Glaukos Corp                                 | CUSIP: 377322102<br>LEI: 5299004JA1LHE21M6T14 | Long             | EC               | CORP              | US        |    311474 | NS      | $33533290.84  | 1.49%             |  |  |  | No            |                  1 | On Loan: No      |
| Guardant Health Inc                          | Guardant Health Inc                          | CUSIP: 40131M109<br>LEI: 254900M8C3E5VC8BR186 | Long             | EC               | CORP              | US        |    303784 | NS      | $28060528.08  | 1.25%             |  |  |  | No            |                  1 | On Loan: No      |
| Hamilton Lane Inc                            | Hamilton Lane Inc                            | CUSIP: 407497106<br>LEI: 5493006OKN3V1NORM833 | Long             | EC               | CORP              | US        |    169441 | NS      | $16842435.40  | 0.75%             |  |  |  | No            |                  1 | On Loan: No      |
| Healthcare Services Group Inc                | Healthcare Services Group Inc                | CUSIP: 421906108<br>LEI: 529900GVZ1D2FBYMS719 | Long             | EC               | CORP              | US        |    784900 | NS      | $14559895.00  | 0.65%             |  |  |  | No            |                  1 | On Loan: No      |
| HealthEquity Inc                             | HealthEquity Inc                             | CUSIP: 42226A107<br>LEI: 529900LRJU1RAFQHM467 | Long             | EC               | CORP              | US        |    422315 | NS      | $35292864.55  | 1.57%             |  |  |  | No            |                  1 | On Loan: No      |
| Hut 8 Corp                                   | Hut 8 Corp                                   | CUSIP: 44812J104<br>LEI: 98450066FEF9C1D3FC20 | Long             | EC               | CORP              | US        |    240717 | NS      | $11292034.47  | 0.50%             |  |  |  | No            |                  1 | On Loan: No      |
| IRhythm Holdings Inc                         | IRhythm Holdings Inc                         | CUSIP: 450056106<br>LEI: 549300FJI72RJGDQS384 | Long             | EC               | CORP              | US        |    169788 | NS      | $20038379.76  | 0.89%             |  |  |  | No            |                  1 | On Loan: No      |
| Insmed Inc                                   | Insmed Inc                                   | CUSIP: 457669307<br>LEI: 529900Q55QAG41CCAC11 | Long             | EC               | CORP              | US        |    336411 | NS      | $55009926.72  | 2.45%             |  |  |  | No            |                  1 | On Loan: No      |
| InterDigital Inc                             | InterDigital Inc                             | CUSIP: 45867G101<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    140996 | NS      | $42580792.00  | 1.90%             |  |  |  | No            |                  1 | On Loan: No      |
| Ionis Pharmaceuticals Inc                    | Ionis Pharmaceuticals Inc                    | CUSIP: 462222100<br>LEI: 549300SI4ZGLG0BLUZ92 | Long             | EC               | CORP              | US        |    148670 | NS      | $11163630.30  | 0.50%             |  |  |  | No            |                  1 | On Loan: No      |
| Janus International Group Inc                | Janus International Group Inc                | CUSIP: 47103N106<br>LEI: N/A                  | Long             | EC               | CORP              | US        |   3345300 | NS      | $17228295.00  | 0.77%             |  |  |  | No            |                  1 | On Loan: No      |
| Karman Holdings Inc                          | Karman Holdings Inc                          | CUSIP: 485924104<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    220172 | NS      | $17624768.60  | 0.78%             |  |  |  | No            |                  1 | On Loan: No      |
| Knight-Swift Transportation Holdings Inc     | Knight-Swift Transportation Holdings Inc     | CUSIP: 499049104<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    437705 | NS      | $25203053.90  | 1.12%             |  |  |  | No            |                  1 | On Loan: No      |
| Kratos Defense & Security Solutions Inc      | Kratos Defense & Security Solutions Inc      | CUSIP: 50077B207<br>LEI: 5493000EKBN0UY1F3F88 | Long             | EC               | CORP              | US        |    233250 | NS      | $16446457.50  | 0.73%             |  |  |  | No            |                  1 | On Loan: No      |
| Lakeland Financial Corp                      | Lakeland Financial Corp                      | CUSIP: 511656100<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    214229 | NS      | $12292460.02  | 0.55%             |  |  |  | No            |                  1 | On Loan: No      |
| Lattice Semiconductor Corp                   | Lattice Semiconductor Corp                   | CUSIP: 518415104<br>LEI: 549300LRBBPZDQ2II567 | Long             | EC               | CORP              | US        |    184881 | NS      | $17149561.56  | 0.76%             |  |  |  | No            |                  1 | On Loan: No      |
| Madrigal Pharmaceuticals Inc                 | Madrigal Pharmaceuticals Inc                 | CUSIP: 558868105<br>LEI: 549300CHIMT2EUSNJB60 | Long             | EC               | CORP              | US        |     26454 | NS      | $13847875.38  | 0.62%             |  |  |  | No            |                  1 | On Loan: No      |
| Marvell Technology Inc                       | Marvell Technology Inc                       | CUSIP: 573874104<br>LEI: 254900WVU0BM7ZCJ9E93 | Long             | EC               | CORP              | US        |    298398 | NS      | $29556321.90  | 1.32%             |  |  |  | No            |                  1 | On Loan: No      |
| MasTec Inc                                   | MasTec Inc                                   | CUSIP: 576323109<br>LEI: 549300TMDIT8WREDOF45 | Long             | EC               | CORP              | US        |    131179 | NS      | $42205531.46  | 1.88%             |  |  |  | No            |                  1 | On Loan: No      |
| McGrath RentCorp                             | McGrath RentCorp                             | CUSIP: 580589109<br>LEI: 254900EUH6IPWFQ83445 | Long             | EC               | CORP              | US        |    136694 | NS      | $15074614.32  | 0.67%             |  |  |  | No            |                  1 | On Loan: No      |
| Miami International Holdings Inc             | Miami International Holdings Inc             | CUSIP: 59356Q108<br>LEI: 254900TREUDBWGLAP788 | Long             | EC               | CORP              | US        |    426477 | NS      | $16598484.84  | 0.74%             |  |  |  | No            |                  1 | On Loan: No      |
| Modine Manufacturing Co                      | Modine Manufacturing Co                      | CUSIP: 607828100<br>LEI: Z35SZFYJR8GZZRSIUU09 | Long             | EC               | CORP              | US        |    189531 | NS      | $41073263.01  | 1.83%             |  |  |  | No            |                  1 | On Loan: No      |
| Mueller Water Products Inc                   | Mueller Water Products Inc                   | CUSIP: 624758108<br>LEI: 549300F7ZU4P1F8V8L60 | Long             | EC               | CORP              | US        |    483160 | NS      | $13282068.40  | 0.59%             |  |  |  | No            |                  1 | On Loan: No      |
| Neurocrine Biosciences Inc                   | Neurocrine Biosciences Inc                   | CUSIP: 64125C109<br>LEI: 549300FECER0XBN49756 | Long             | EC               | CORP              | US        |    116190 | NS      | $15306870.60  | 0.68%             |  |  |  | No            |                  1 | On Loan: No      |
| Nextpower Inc                                | Nextpower Inc                                | CUSIP: 65290E101<br>LEI: N/A                  | Long             | EC               | CORP              | US        |     71632 | NS      | $8635237.60   | 0.38%             |  |  |  | No            |                  1 | On Loan: No      |
| Nuvalent Inc                                 | Nuvalent Inc                                 | CUSIP: 670703107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    174598 | NS      | $17887565.10  | 0.80%             |  |  |  | No            |                  1 | On Loan: No      |
| Parsons Corp                                 | Parsons Corp                                 | CUSIP: 70202L102<br>LEI: 549300ZXH0VRBSEPX752 | Long             | EC               | CORP              | US        |    486820 | NS      | $26371039.40  | 1.17%             |  |  |  | No            |                  1 | On Loan: No      |
| Paylocity Holding Corp                       | Paylocity Holding Corp                       | CUSIP: 70438V106<br>LEI: 549300VH75FTB4DIEE07 | Long             | EC               | CORP              | US        |    167076 | NS      | $18050891.04  | 0.80%             |  |  |  | No            |                  1 | On Loan: No      |
| Pennant Group Inc/The                        | Pennant Group Inc/The                        | CUSIP: 70805E109<br>LEI: 5493007H2OYR6BYLDA64 | Long             | EC               | CORP              | US        |    699307 | NS      | $21314877.36  | 0.95%             |  |  |  | No            |                  1 | On Loan: No      |
| Repligen Corp                                | Repligen Corp                                | CUSIP: 759916109<br>LEI: 54930006QJ0T5AUO1E71 | Long             | EC               | CORP              | US        |    159623 | NS      | $18806781.86  | 0.84%             |  |  |  | No            |                  1 | On Loan: No      |
| Revolution Medicines Inc                     | Revolution Medicines Inc                     | CUSIP: 76155X100<br>LEI: 549300JU3X0LBCQ65892 | Long             | EC               | CORP              | US        |    135489 | NS      | $13176305.25  | 0.59%             |  |  |  | No            |                  1 | On Loan: No      |
| Rhythm Pharmaceuticals Inc                   | Rhythm Pharmaceuticals Inc                   | CUSIP: 76243J105<br>LEI: 529900JPNWXQ8DPLA551 | Long             | EC               | CORP              | US        |    124824 | NS      | $10855943.28  | 0.48%             |  |  |  | No            |                  1 | On Loan: No      |
| Rush Street Interactive Inc                  | Rush Street Interactive Inc                  | CUSIP: 782011100<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    631555 | NS      | $13736321.25  | 0.61%             |  |  |  | No            |                  1 | On Loan: No      |
| Ryan Specialty Holdings Inc                  | Ryan Specialty Holdings Inc                  | CUSIP: 78351F107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    647394 | NS      | $21843073.56  | 0.97%             |  |  |  | No            |                  1 | On Loan: No      |
| State Street SPDR S&P Biotech ETF            | State Street SPDR S&P Biotech ETF            | CUSIP: 78464A870<br>LEI: 549300Y3ODFMPD5XXL58 | Long             | EC               | RF                | US        |    278004 | NS      | $35509450.92  | 1.58%             |  |  |  | No            |                  1 | On Loan: No      |
| SPX Technologies Inc                         | SPX Technologies Inc                         | CUSIP: 78473E103<br>LEI: P761FCI0CPOG24I8JE10 | Long             | EC               | CORP              | US        |    279748 | NS      | $55932815.12  | 2.49%             |  |  |  | No            |                  1 | On Loan: No      |
| SailPoint Inc                                | SailPoint Inc                                | CUSIP: 78781J109<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    944155 | NS      | $12500612.20  | 0.56%             |  |  |  | No            |                  1 | On Loan: No      |
| Semtech Corp                                 | Semtech Corp                                 | CUSIP: 816850101<br>LEI: 549300J7WUY317SW6O61 | Long             | EC               | CORP              | US        |    421296 | NS      | $32393449.44  | 1.44%             |  |  |  | No            |                  1 | On Loan: No      |
| SiTime Corp                                  | SiTime Corp                                  | CUSIP: 82982T106<br>LEI: N/A                  | Long             | EC               | CORP              | US        |     48846 | NS      | $16868966.10  | 0.75%             |  |  |  | No            |                  1 | On Loan: No      |
| Sprouts Farmers Market Inc                   | Sprouts Farmers Market Inc                   | CUSIP: 85208M102<br>LEI: 529900P8F4G77Q52XE24 | Long             | EC               | CORP              | US        |    248151 | NS      | $19139886.63  | 0.85%             |  |  |  | No            |                  1 | On Loan: No      |
| Sterling Infrastructure Inc                  | Sterling Infrastructure Inc                  | CUSIP: 859241101<br>LEI: 549300RZIK6K1215XU51 | Long             | EC               | CORP              | US        |    107856 | NS      | $43926513.12  | 1.96%             |  |  |  | No            |                  1 | On Loan: No      |
| StoneX Group Inc                             | StoneX Group Inc                             | CUSIP: 861896108<br>LEI: 549300JUF07L8VF02M60 | Long             | EC               | CORP              | US        |    413430 | NS      | $33343129.50  | 1.48%             |  |  |  | No            |                  1 | On Loan: No      |
| TopBuild Corp                                | TopBuild Corp                                | CUSIP: 89055F103<br>LEI: 5493007YSKHOL4RWII46 | Long             | EC               | CORP              | US        |     92442 | NS      | $32474874.60  | 1.45%             |  |  |  | No            |                  1 | On Loan: No      |
| VSE Corp                                     | VSE Corp                                     | CUSIP: 918284100<br>LEI: 549300QWJYQDH4U5XI15 | Long             | EC               | CORP              | US        |    223919 | NS      | $41290663.60  | 1.84%             |  |  |  | No            |                  1 | On Loan: No      |
| Viavi Solutions Inc                          | Viavi Solutions Inc                          | CUSIP: 925550105<br>LEI: 5493006K3E3N1QOBF092 | Long             | EC               | CORP              | US        |   1333081 | NS      | $44364935.68  | 1.98%             |  |  |  | No            |                  1 | On Loan: No      |
| WD-40 Co                                     | WD-40 Co                                     | CUSIP: 929236107<br>LEI: 549300UZHTN0CNKBOB19 | Long             | EC               | CORP              | US        |    146352 | NS      | $29847026.88  | 1.33%             |  |  |  | No            |                  1 | On Loan: No      |
| Western Alliance Bancorp                     | Western Alliance Bancorp                     | CUSIP: 957638109<br>LEI: 5493003VJXZ5JXT9S762 | Long             | EC               | CORP              | US        |    419941 | NS      | $29752819.85  | 1.33%             |  |  |  | No            |                  1 | On Loan: No      |
| Willdan Group Inc                            | Willdan Group Inc                            | CUSIP: 96924N100<br>LEI: 529900YQO0LF5VAEKN05 | Long             | EC               | CORP              | US        |    206119 | NS      | $15780470.64  | 0.70%             |  |  |  | No            |                  1 | On Loan: No      |
| Wingstop Inc                                 | Wingstop Inc                                 | CUSIP: 974155103<br>LEI: 549300SSH3F8HPIM3Q47 | Long             | EC               | CORP              | US        |    242143 | NS      | $37524900.71  | 1.67%             |  |  |  | No            |                  1 | On Loan: No      |
| Workiva Inc                                  | Workiva Inc                                  | CUSIP: 98139A105<br>LEI: 529900RTOQ8GRWHY1O58 | Long             | EC               | CORP              | US        |    184993 | NS      | $11031132.59  | 0.49%             |  |  |  | No            |                  1 | On Loan: No      |
| Zeta Global Holdings Corp                    | Zeta Global Holdings Corp                    | CUSIP: 98956A105<br>LEI: 2549008P9XSJQHGQ9S27 | Long             | EC               | CORP              | US        |    895992 | NS      | $14264192.64  | 0.64%             |  |  |  | No            |                  1 | On Loan: No      |
| Zscaler Inc                                  | Zscaler Inc                                  | CUSIP: 98980G102<br>LEI: 529900MZ0RTK1BWRNF46 | Long             | EC               | CORP              | US        |     83570 | NS      | $11724035.30  | 0.52%             |  |  |  | No            |                  1 | On Loan: No      |
| DNA SCRIPT SAS                               | DNA SCRIPT SAS                               | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | FR        |      1139 | NS      | $94709.96     | 0.00%             |  |  |  | Yes           |                  3 | On Loan: No      |
| Credo Technology Group Holding Ltd           | Credo Technology Group Holding Ltd           | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | KY        |    147591 | NS      | $13854367.17  | 0.62%             |  |  |  | No            |                  1 | On Loan: No      |
| FTAI Aviation Ltd                            | FTAI Aviation Ltd                            | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | KY        |    109872 | NS      | $26918640.00  | 1.20%             |  |  |  | No            |                  1 | On Loan: No      |
| NIQ Global Intelligence Plc                  | NIQ Global Intelligence Plc                  | CUSIP: 000000000<br>LEI: 529900Z0P0XAM4O3H416 | Long             | EC               | CORP              | IE        |   1232928 | NS      | $14018391.36  | 0.62%             |  |  |  | No            |                  1 | On Loan: No      |
| SharkNinja Inc                               | SharkNinja Inc                               | CUSIP: 000000000<br>LEI: N/A                  | Long             | EC               | CORP              | KY        |    238583 | NS      | $25265939.70  | 1.13%             |  |  |  | No            |                  1 | On Loan: No      |
| Monday.com Ltd                               | Monday.com Ltd                               | CUSIP: 000000000<br>LEI: 984500MDAFVD86E3B328 | Long             | EC               | CORP              | IL        |    106304 | NS      | $7346669.44   | 0.33%             |  |  |  | No            |                  1 | On Loan: No      |
| Newamsterdam Pharma Co NV                    | Newamsterdam Pharma Co NV                    | CUSIP: 000000000<br>LEI: 984500B6549TF6E14D36 | Long             | EC               | CORP              | NL        |    363308 | NS      | $11629489.08  | 0.52%             |  |  |  | No            |                  1 | On Loan: No      |

### Signature

**Date Signed:** 2026-05-04

**Name of Applicant:** COLUMBIA ACORN TRUST

**Signature:** Jennifer Borges

**Name of Signer:** Jennifer Borges

**Title:** Senior Manager