# EDGAR Filing Document

**Accession Number:** 0001350290
**File Stem:** 0001104659-26-062488
**Filing Date:** 2026-5
**Character Count:** 17953
**Document Hash:** c5b513c8d0ff9cf53acf5bd601e69aff
**Contains OCR:** False
**Source Format:** 

## Filing Content

## Filing Summary
**0001104659-26-062488.hdr.sgml**: 20260515

**ACCESSION NUMBER**: 0001104659-26-062488

**CONFORMED SUBMISSION TYPE**: 13F-HR

**PUBLIC DOCUMENT COUNT**: 2

**CONFORMED PERIOD OF REPORT**: 20260331

**FILED AS OF DATE**: 20260515

**DATE AS OF CHANGE**: 20260515

**EFFECTIVENESS DATE**: 20260515

**FILER**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** Portolan Capital Management, LLC
- **CENTRAL INDEX KEY:** 0001350290

**ORGANIZATION NAME:**
- **EIN:** 201926326
- **STATE OF INCORPORATION:** DE
- **FISCAL YEAR END:** 1231

**FILING VALUES:**
- **FORM TYPE:** 13F-HR
- **SEC ACT:** 1934 Act
- **SEC FILE NUMBER:** 028-11602
- **FILM NUMBER:** 26986885

**BUSINESS ADDRESS:**
- **STREET 1:** 2 INTERNATIONAL PLACE, 26TH FLOOR
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02110
- **BUSINESS PHONE:** 617.753.6400

**MAIL ADDRESS:**
- **STREET 1:** 2 INTERNATIONAL PLACE, 26TH FLOOR
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02110

### FORM 13F COVER PAGE

**Report for the Calendar Year or Quarter Ended:** 03-31-2026

**Check here if Amendment** [ ] **Amendment Number:** 

**This Amendment (Check only one.):** [ ] is a restatement.

[ ] adds new holdings entries.

**Institutional Investment Manager Filing this Report:**

**Name:** Portolan Capital Management LLC<br>**Address:** 2 INTERNATIONAL PLACE 26TH FLOOR<br>BOSTON, MA 02110

**Form 13F File Number:** 028-11602

The institutional investment manager filing this report and the person by whom it is signed hereby represent that the person signing the report is authorized to submit it, that all information contained herein is true, correct and complete, and that it is understood that all required items, statements, schedules, lists, and tables, are considered integral parts of this form.

**Person Signing this Report on Behalf of Reporting Manager:**

**Name:** Betsy Flaherty<br>**Title:** Chief Compliance Officer<br>**Phone:** 617-753-6400

**Signature, Place, and Date of Signing:**

/s/Betsy Flaherty  Boston, MA  05-15-2026<br>[Signature]  [City, State]  [Date]

**Report Type (Check only one.):**

[x] **13F HOLDINGS REPORT.** (Check here if all holdings of this reporting manager are reported in this report.)

[ ] **13F NOTICE.** (Check here if no holdings are reported in this report, and all holdings are reported by other reporting manager(s).)

[ ] **13F COMBINATION REPORT.** (Check here if a portion of the holdings for this reporting manager are reported in this report and a portion are reported by other reporting manager(s).)

### Form 13F Summary Page

**Report Summary:**

**Number of Other Included Managers:** 0

**Form 13F Information Table Entry Total:** 144

**Form 13F Information Table Value Total:** $1878962983

      (round to nearest dollar)

**List of Other Included Managers:**

Provide a numbered list of the name(s) and Form 13F file number(s) of all institutional investment managers with respect to which this report is filed, other than the manager filing this report.

[If there are no entries in this list, state “NONE” and omit the column headings and list entries.]

**NONE**

### FORM 13F INFORMATION TABLE

---

|  |  |  |  |  |  |  |  |  |  |  |  |  |
| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
| NAME OF ISSUER | TITLE OF CLASS | CUSIP | FIGI | VALUE (x$1000) | SHRS OR PRN AMT | SH/PRN | PUT/CALL | INVESTMENT DISCRETION | OTHER MANAGER | VOTING AUTHORITY | VOTING AUTHORITY | VOTING AUTHORITY |
| NAME OF ISSUER | TITLE OF CLASS | CUSIP | FIGI | VALUE (x$1000) | SHRS OR PRN AMT | SH/PRN | PUT/CALL | INVESTMENT DISCRETION | OTHER MANAGER | SOLE | SHARED | NONE |
| ACM RESH INC | COM CL A | 00108J109 |  | 3276124 | 83256 | SH |  | SOLE |  | 83256 | 0 | 0 |
| ADEIA INC | COM | 00676P107 |  | 17528948 | 729461 | SH |  | SOLE |  | 729461 | 0 | 0 |
| AEROVIRONMENT INC | COM | 008073108 |  | 4920384 | 26880 | SH |  | SOLE |  | 26880 | 0 | 0 |
| AKAMAI TECHNOLOGIES INC | COM | 00971T101 |  | 48632314 | 423442 | SH |  | SOLE |  | 423442 | 0 | 0 |
| ALLIENT INC | COM | 019330109 |  | 19112838 | 323453 | SH |  | SOLE |  | 323453 | 0 | 0 |
| ALPHABET INC | CAP STK CL C | 02079K107 |  | 1637971 | 5710 | SH |  | SOLE |  | 5710 | 0 | 0 |
| AMBARELLA INC | SHS | G037AX101 |  | 5597855 | 108749 | SH |  | SOLE |  | 108749 | 0 | 0 |
| AMER SPORTS INC | COM SHS | G0260P102 |  | 11779632 | 357826 | SH |  | SOLE |  | 357826 | 0 | 0 |
| AMERICAN SUPERCONDUCTOR CORP | SHS NEW | 030111207 |  | 10007279 | 295636 | SH |  | SOLE |  | 295636 | 0 | 0 |
| AMPLIFY ENERGY CORP NEW | COM | 03212B103 |  | 462509 | 74120 | SH |  | SOLE |  | 74120 | 0 | 0 |
| AMPLITUDE INC | COM CL A | 03213A104 |  | 2452424 | 359593 | SH |  | SOLE |  | 359593 | 0 | 0 |
| APPLIED OPTOELECTRONICS INC | COM | 03823U102 |  | 2523573 | 29833 | SH |  | SOLE |  | 29833 | 0 | 0 |
| AXCELIS TECHNOLOGIES INC | COM NEW | 054540208 |  | 7182797 | 77168 | SH |  | SOLE |  | 77168 | 0 | 0 |
| BAIDU INC | SPON ADR REP A | 056752108 |  | 40827519 | 366429 | SH |  | SOLE |  | 366429 | 0 | 0 |
| BEL FUSE INC | CL B | 077347300 |  | 10526597 | 53170 | SH |  | SOLE |  | 53170 | 0 | 0 |
| BIO RAD LABS INC | CL A | 090572207 |  | 3950724 | 14173 | SH |  | SOLE |  | 14173 | 0 | 0 |
| BIOGEN INC | COM | 09062X103 |  | 21721122 | 118481 | SH |  | SOLE |  | 118481 | 0 | 0 |
| BLACKLINE INC | COM | 09239B109 |  | 5883962 | 159026 | SH |  | SOLE |  | 159026 | 0 | 0 |
| CANADA GOOSE HLDGS INC | SHS SUB VTG | 135086106 |  | 35304499 | 3218277 | SH |  | SOLE |  | 3218277 | 0 | 0 |
| CANDEL THERAPEUTICS INC | COM | 137404109 |  | 14182193 | 2894325 | SH |  | SOLE |  | 2894325 | 0 | 0 |
| CAREDX INC | COM | 14167L103 |  | 5908511 | 340352 | SH |  | SOLE |  | 340352 | 0 | 0 |
| CASELLA WASTE SYS INC | CL A | 147448104 |  | 1198034 | 15100 | SH |  | SOLE |  | 15100 | 0 | 0 |
| CASTLE BIOSCIENCES INC | COM | 14843C105 |  | 33149448 | 1350283 | SH |  | SOLE |  | 1350283 | 0 | 0 |
| CELESTICA INC | COM | 15101Q207 |  | 55734050 | 197863 | SH |  | SOLE |  | 197863 | 0 | 0 |
| CELLEBRITE DI LTD | ORDINARY SHARES | M2197Q107 |  | 30053133 | 2180924 | SH |  | SOLE |  | 2180924 | 0 | 0 |
| CELSIUS HLDGS INC | COM NEW | 15118V207 |  | 5839902 | 164597 | SH |  | SOLE |  | 164597 | 0 | 0 |
| CENTURY ALUM CO | COM | 156431108 |  | 5577839 | 95039 | SH |  | SOLE |  | 95039 | 0 | 0 |
| CIENA CORP | COM NEW | 171779309 |  | 7169055 | 18466 | SH |  | SOLE |  | 18466 | 0 | 0 |
| CIRCLE INTERNET GROUP INC | COM CL A | 172573107 |  | 20398276 | 213796 | SH |  | SOLE |  | 213796 | 0 | 0 |
| CORE NATURAL RESOURCES INC | COM SHS | 218937100 |  | 24824780 | 237036 | SH |  | SOLE |  | 237036 | 0 | 0 |
| CRESCENT ENERGY COMPANY | CL A COM | 44952J104 |  | 35115417 | 2601142 | SH |  | SOLE |  | 2601142 | 0 | 0 |
| DANA INC | COM | 235825205 |  | 1152075 | 34237 | SH |  | SOLE |  | 34237 | 0 | 0 |
| DAVE INC | CLASS A COM NEW | 23834J201 |  | 18356398 | 105442 | SH |  | SOLE |  | 105442 | 0 | 0 |
| DHT HOLDINGS INC | SHS NEW | Y2065G121 |  | 19946967 | 1091788 | SH |  | SOLE |  | 1091788 | 0 | 0 |
| DIGI INTL INC | COM | 253798102 |  | 8206339 | 170256 | SH |  | SOLE |  | 170256 | 0 | 0 |
| DIGITALBRIDGE GROUP INC | CL A NEW | 25401T603 |  | 5753665 | 373130 | SH |  | SOLE |  | 373130 | 0 | 0 |
| DOMO INC | COM CL B | 257554105 |  | 603138 | 197104 | SH |  | SOLE |  | 197104 | 0 | 0 |
| DOUGLAS ELLIMAN INC | COM | 25961D105 |  | 4377209 | 2669030 | SH |  | SOLE |  | 2669030 | 0 | 0 |
| DRAGANFLY INC. | COM | 26142Q304 |  | 3352204 | 682730 | SH |  | SOLE |  | 682730 | 0 | 0 |
| E L F BEAUTY INC | COM | 26856L103 |  | 8285811 | 136707 | SH |  | SOLE |  | 136707 | 0 | 0 |
| EBAY INC. | COM | 278642103 |  | 11477258 | 126096 | SH |  | SOLE |  | 126096 | 0 | 0 |
| ELEMENT SOLUTIONS INC | COM | 28618M106 |  | 7552860 | 221232 | SH |  | SOLE |  | 221232 | 0 | 0 |
| ENOVA INTL INC | COM | 29357K103 |  | 10991771 | 80923 | SH |  | SOLE |  | 80923 | 0 | 0 |
| ERMENEGILDO ZEGNA N V | ORD SHS | N30577105 |  | 7404723 | 710626 | SH |  | SOLE |  | 710626 | 0 | 0 |
| ESTABLISHMENT LABS HLDGS INC | COM | G31249108 |  | 543555 | 9573 | SH |  | SOLE |  | 9573 | 0 | 0 |
| EVERUS CONSTR GROUP | COM | 300426103 |  | 1030074 | 8725 | SH |  | SOLE |  | 8725 | 0 | 0 |
| EVOLV TECHNOLOGIES HLDNGS IN | COM CL A | 30049H102 |  | 10863912 | 1795688 | SH |  | SOLE |  | 1795688 | 0 | 0 |
| EXZEO GROUP INC | COM SHS | 30234F101 |  | 3079013 | 209885 | SH |  | SOLE |  | 209885 | 0 | 0 |
| FABRINET | SHS | G3323L100 |  | 28560000 | 54763 | SH |  | SOLE |  | 54763 | 0 | 0 |
| FLEXTRONICS INTL LTD | ORD | Y2573F102 |  | 29007159 | 443128 | SH |  | SOLE |  | 443128 | 0 | 0 |
| FORGENT POWER SOLUTIONS INC | COM SHS CL A | 34631F102 |  | 3451899 | 117933 | SH |  | SOLE |  | 117933 | 0 | 0 |
| FORRESTER RESH INC | COM | 346563109 |  | 8723622 | 1541276 | SH |  | SOLE |  | 1541276 | 0 | 0 |
| FREQUENCY ELECTRS INC | COM | 358010106 |  | 6645241 | 150141 | SH |  | SOLE |  | 150141 | 0 | 0 |
| FULGENT GENETICS INC | COM | 359664109 |  | 1695608 | 106642 | SH |  | SOLE |  | 106642 | 0 | 0 |
| GDS HLDGS LTD | SPONSORED ADS | 36165L108 |  | 615671 | 15281 | SH |  | SOLE |  | 15281 | 0 | 0 |
| GENEDX HOLDINGS CORP | COM CL A | 81663L200 |  | 6587367 | 102575 | SH |  | SOLE |  | 102575 | 0 | 0 |
| GENIUS SPORTS LIMITED | SHARES CL A | G3934V109 |  | 12364210 | 2791018 | SH |  | SOLE |  | 2791018 | 0 | 0 |
| GIGACLOUD TECHNOLOGY INC | CLASS A ORD | G38644103 |  | 2360395 | 52014 | SH |  | SOLE |  | 52014 | 0 | 0 |
| GRAHAM CORP | COM | 384556106 |  | 8172640 | 103556 | SH |  | SOLE |  | 103556 | 0 | 0 |
| GREEN DOT CORP | CL A | 39304D102 |  | 3351380 | 298697 | SH |  | SOLE |  | 298697 | 0 | 0 |
| GUARDANT HEALTH INC | COM | 40131M109 |  | 1307497 | 14155 | SH |  | SOLE |  | 14155 | 0 | 0 |
| HAFNIA LTD | SHS | Y2990R101 |  | 14810249 | 1948717 | SH |  | SOLE |  | 1948717 | 0 | 0 |
| HEALTHEQUITY INC | COM | 42226A107 |  | 8494556 | 101646 | SH |  | SOLE |  | 101646 | 0 | 0 |
| HONEST CO INC | COM | 438333106 |  | 8954752 | 3045834 | SH |  | SOLE |  | 3045834 | 0 | 0 |
| I3 VERTICALS INC | COM CL A | 46571Y107 |  | 4634647 | 207274 | SH |  | SOLE |  | 207274 | 0 | 0 |
| IES HOLDINGS INC | COM | 44951W106 |  | 5655222 | 11869 | SH |  | SOLE |  | 11869 | 0 | 0 |
| JABIL INC | COM | 466313103 |  | 20505839 | 77197 | SH |  | SOLE |  | 77197 | 0 | 0 |
| KOSMOS ENERGY LTD | COM | 500688106 |  | 1675067 | 602542 | SH |  | SOLE |  | 602542 | 0 | 0 |
| KRATOS DEFENSE  SEC SOLUTIO | COM NEW | 50077B207 |  | 18319274 | 259811 | SH |  | SOLE |  | 259811 | 0 | 0 |
| KURA SUSHI USA INC | CL A COM | 501270102 |  | 21021585 | 301212 | SH |  | SOLE |  | 301212 | 0 | 0 |
| L3HARRIS TECHNOLOGIES INC | COM | 502431109 |  | 744489 | 2157 | SH |  | SOLE |  | 2157 | 0 | 0 |
| LANTHEUS HLDGS INC | COM | 516544103 |  | 31659032 | 417390 | SH |  | SOLE |  | 417390 | 0 | 0 |
| LIGHTPATH TECHNOLOGIES INC | COM CL A | 532257805 |  | 10435834 | 1040462 | SH |  | SOLE |  | 1040462 | 0 | 0 |
| LINCOLN EDL SVCS CORP | COM | 533535100 |  | 569805 | 14007 | SH |  | SOLE |  | 14007 | 0 | 0 |
| LUMENTUM HLDGS INC | COM | 55024U109 |  | 4091469 | 5822 | SH |  | SOLE |  | 5822 | 0 | 0 |
| LUXEXPERIENCE BV | SPONSORED ADS | 55406W103 |  | 29871544 | 3733943 | SH |  | SOLE |  | 3733943 | 0 | 0 |
| MAMAS CREATIONS INC | COM | 56146T103 |  | 18054198 | 1176936 | SH |  | SOLE |  | 1176936 | 0 | 0 |
| MAYVILLE ENGR CO INC | COM | 578605107 |  | 14739391 | 821136 | SH |  | SOLE |  | 821136 | 0 | 0 |
| META PLATFORMS INC | CL A | 30303M102 |  | 5870626 | 10261 | SH |  | SOLE |  | 10261 | 0 | 0 |
| MIAMI INTL HLDGS INC | COM | 59356Q108 |  | 5013635 | 128819 | SH |  | SOLE |  | 128819 | 0 | 0 |
| MOBILEYE GLOBAL INC | COMMON CLASS A | 60741F104 |  | 4658787 | 678135 | SH |  | SOLE |  | 678135 | 0 | 0 |
| MODINE MFG CO | COM | 607828100 |  | 89304457 | 412092 | SH |  | SOLE |  | 412092 | 0 | 0 |
| MOOG INC | CL A | 615394202 |  | 8614444 | 29437 | SH |  | SOLE |  | 29437 | 0 | 0 |
| MURPHY USA INC | COM | 626755102 |  | 23763415 | 48107 | SH |  | SOLE |  | 48107 | 0 | 0 |
| NATERA INC | COM | 632307104 |  | 2283486 | 11418 | SH |  | SOLE |  | 11418 | 0 | 0 |
| NEXTPOWER INC | CLASS A COM | 65290E101 |  | 6840851 | 56747 | SH |  | SOLE |  | 56747 | 0 | 0 |
| NLIGHT INC | COM | 65487K100 |  | 12862344 | 225576 | SH |  | SOLE |  | 225576 | 0 | 0 |
| NOKIA CORP | SPONSORED ADR | 654902204 |  | 19247519 | 2393970 | SH |  | SOLE |  | 2393970 | 0 | 0 |
| NORTHERN OIL  GAS INC | COM | 665531307 |  | 5970052 | 204244 | SH |  | SOLE |  | 204244 | 0 | 0 |
| NOVANTA INC | COM | 67000B104 |  | 4429952 | 37507 | SH |  | SOLE |  | 37507 | 0 | 0 |
| OKEANIS ECO TANKERS COR | SHS | Y64177101 |  | 27260927 | 538860 | SH |  | SOLE |  | 538860 | 0 | 0 |
| ON HLDG AG | NAMEN AKT A | H5919C104 |  | 6887145 | 202444 | SH |  | SOLE |  | 202444 | 0 | 0 |
| ONTO INNOVATION INC | COM | 683344105 |  | 4565473 | 22263 | SH |  | SOLE |  | 22263 | 0 | 0 |
| OPEN LENDING CORP | COM | 68373J104 |  | 8628610 | 6902888 | SH |  | SOLE |  | 6902888 | 0 | 0 |
| OSHKOSH CORP | COM | 688239201 |  | 2677014 | 18185 | SH |  | SOLE |  | 18185 | 0 | 0 |
| PAR PAC HOLDINGS INC | COM NEW | 69888T207 |  | 43461762 | 693834 | SH |  | SOLE |  | 693834 | 0 | 0 |
| PARK AEROSPACE CORP | COM | 70014A104 |  | 11243734 | 410655 | SH |  | SOLE |  | 410655 | 0 | 0 |
| PBF ENERGY INC | CL A | 69318G106 |  | 38095905 | 799998 | SH |  | SOLE |  | 799998 | 0 | 0 |
| PERSONALIS INC | COM | 71535D106 |  | 4692862 | 736713 | SH |  | SOLE |  | 736713 | 0 | 0 |
| PLANET LABS PBC | COM CL A | 72703X106 |  | 1321700 | 47288 | SH |  | SOLE |  | 47288 | 0 | 0 |
| PORCH GROUP INC | COM | 733245104 |  | 15378861 | 2144890 | SH |  | SOLE |  | 2144890 | 0 | 0 |
| POWELL INDS INC | COM | 739128106 |  | 41030096 | 75830 | SH |  | SOLE |  | 75830 | 0 | 0 |
| PRIMORIS SVCS CORP | COM | 74164F103 |  | 18684743 | 130626 | SH |  | SOLE |  | 130626 | 0 | 0 |
| PROCEPT BIOROBOTICS CORP | COM | 74276L105 |  | 6295417 | 251716 | SH |  | SOLE |  | 251716 | 0 | 0 |
| PROPETRO HLDG CORP | COM | 74347M108 |  | 4536484 | 314815 | SH |  | SOLE |  | 314815 | 0 | 0 |
| PULSE BIOSCIENCES INC | COM | 74587B101 |  | 15833501 | 733372 | SH |  | SOLE |  | 733372 | 0 | 0 |
| QUALYS INC | COM | 74758T303 |  | 20998610 | 239028 | SH |  | SOLE |  | 239028 | 0 | 0 |
| QUANTERIX CORP | COM | 74766Q101 |  | 10369297 | 2945823 | SH |  | SOLE |  | 2945823 | 0 | 0 |
| RADNET INC | COM | 750491102 |  | 1326158 | 23728 | SH |  | SOLE |  | 23728 | 0 | 0 |
| RANPAK HOLDINGS CORP | COM CL A | 75321W103 |  | 6953221 | 1947681 | SH |  | SOLE |  | 1947681 | 0 | 0 |
| REGAL REXNORD CORPORATION | COM | 758750103 |  | 10107359 | 53975 | SH |  | SOLE |  | 53975 | 0 | 0 |
| REMITLY GLOBAL INC | COM | 75960P104 |  | 10940261 | 698166 | SH |  | SOLE |  | 698166 | 0 | 0 |
| REPAY HLDGS CORP | COM CL A | 76029L100 |  | 7426063 | 2856178 | SH |  | SOLE |  | 2856178 | 0 | 0 |
| SANDRIDGE ENERGY INC | COM NEW | 80007P869 |  | 2015769 | 123591 | SH |  | SOLE |  | 123591 | 0 | 0 |
| SANMINA CORP | COM | 801056102 |  | 3543320 | 27332 | SH |  | SOLE |  | 27332 | 0 | 0 |
| SEMTECH CORP | COM | 816850101 |  | 25552623 | 332327 | SH |  | SOLE |  | 332327 | 0 | 0 |
| SEZZLE INC | COM | 78435P105 |  | 4433211 | 70046 | SH |  | SOLE |  | 70046 | 0 | 0 |
| SHIFT4 PMTS INC | CL A | 82452J109 |  | 9370639 | 214284 | SH |  | SOLE |  | 214284 | 0 | 0 |
| SOLARIS ENERGY INFRAS INC | COM CL A | 83418M103 |  | 1578324 | 27930 | SH |  | SOLE |  | 27930 | 0 | 0 |
| SPROUT SOCIAL INC | COM CL A | 85209W109 |  | 3182595 | 558350 | SH |  | SOLE |  | 558350 | 0 | 0 |
| TAKE-TWO INTERACTIVE SOFTWAR | COM | 874054109 |  | 6363253 | 32219 | SH |  | SOLE |  | 32219 | 0 | 0 |
| TALEN ENERGY CORP | COM | 87422Q109 |  | 26040868 | 81574 | SH |  | SOLE |  | 81574 | 0 | 0 |
| TALOS ENERGY INC | COM | 87484T108 |  | 5430092 | 344549 | SH |  | SOLE |  | 344549 | 0 | 0 |
| TAPESTRY INC | COM | 876030107 |  | 2404232 | 17038 | SH |  | SOLE |  | 17038 | 0 | 0 |
| TECOGEN INC NEW | COM NEW | 87876P201 |  | 1135501 | 443555 | SH |  | SOLE |  | 443555 | 0 | 0 |
| TELIX PHARMACEUTICAL LTD | SPONSORED ADS | 87961M105 |  | 5179925 | 541267 | SH |  | SOLE |  | 541267 | 0 | 0 |
| THE REAL BROKERAGE INC | COM NEW | 75585H206 |  | 36299620 | 14519848 | SH |  | SOLE |  | 14519848 | 0 | 0 |
| TOWER SEMICONDUCTOR LTD | SHS NEW | M87915274 |  | 5622906 | 32043 | SH |  | SOLE |  | 32043 | 0 | 0 |
| TRANSMEDICS GROUP INC | COM | 89377M109 |  | 1062494 | 10688 | SH |  | SOLE |  | 10688 | 0 | 0 |
| TTM TECHNOLOGIES INC | COM | 87305R109 |  | 70872465 | 727494 | SH |  | SOLE |  | 727494 | 0 | 0 |
| TWILIO INC | CL A | 90138F102 |  | 9407436 | 74769 | SH |  | SOLE |  | 74769 | 0 | 0 |
| ULTRA CLEAN HLDGS INC | COM | 90385V107 |  | 2095466 | 33700 | SH |  | SOLE |  | 33700 | 0 | 0 |
| UNITED THERAPEUTICS CORP DEL | COM | 91307C102 |  | 37615686 | 63435 | SH |  | SOLE |  | 63435 | 0 | 0 |
| UPWORK INC | COM | 91688F104 |  | 17213338 | 1570560 | SH |  | SOLE |  | 1570560 | 0 | 0 |
| VEECO INSTRS INC DEL | COM | 922417100 |  | 8579413 | 253379 | SH |  | SOLE |  | 253379 | 0 | 0 |
| VIAVI SOLUTIONS INC | COM | 925550105 |  | 4853688 | 145844 | SH |  | SOLE |  | 145844 | 0 | 0 |
| VICOR CORP | COM | 925815102 |  | 4293870 | 26670 | SH |  | SOLE |  | 26670 | 0 | 0 |
| VIRTU FINL INC | CL A | 928254101 |  | 53298790 | 1211887 | SH |  | SOLE |  | 1211887 | 0 | 0 |
| VISHAY PRECISION GROUP INC | COM | 92835K103 |  | 44282669 | 1019868 | SH |  | SOLE |  | 1019868 | 0 | 0 |
| VNET GROUP INC | SPONSORED ADS A | 90138A103 |  | 6925861 | 825490 | SH |  | SOLE |  | 825490 | 0 | 0 |
| WARBY PARKER INC | CL A COM | 93403J106 |  | 10549875 | 500706 | SH |  | SOLE |  | 500706 | 0 | 0 |
| WILLDAN GROUP INC | COM | 96924N100 |  | 7193884 | 93964 | SH |  | SOLE |  | 93964 | 0 | 0 |
| XPENG INC | ADS | 98422D105 |  | 4712642 | 275432 | SH |  | SOLE |  | 275432 | 0 | 0 |
| ZETA GLOBAL HOLDINGS CORP | CL A | 98956A105 |  | 7741434 | 486271 | SH |  | SOLE |  | 486271 | 0 | 0 |

---