# EDGAR Filing Document

**Accession Number:** 0001581835
**File Stem:** 0001104659-23-021198
**Filing Date:** 2023-2
**Character Count:** 37028
**Document Hash:** 078cc293535340e0068d2da83bfa23dc
**Contains OCR:** False
**Source Format:** 

## Filing Content

## Filing Summary
**0001104659-23-021198.hdr.sgml**: 20230214

**ACCESSION NUMBER**: 0001104659-23-021198

**CONFORMED SUBMISSION TYPE**: SC 13D/A

**PUBLIC DOCUMENT COUNT**: 1

**FILED AS OF DATE**: 20230214

**DATE AS OF CHANGE**: 20230214

**GROUP MEMBERS**: BPIFRANCE S.A.

**GROUP MEMBERS**: CAISSE DES DEPOTS ET CONSIGNATIONS

**GROUP MEMBERS**: EPIC BPIFRANCE

**SUBJECT COMPANY**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** Cellectis S.A.
- **CENTRAL INDEX KEY:** 0001627281
- **STANDARD INDUSTRIAL CLASSIFICATION:** BIOLOGICAL PRODUCTS (NO DIAGNOSTIC SUBSTANCES) [2836]
- **IRS NUMBER:** 000000000
- **STATE OF INCORPORATION:** I0
- **FISCAL YEAR END:** 1231

**FILING VALUES:**
- **FORM TYPE:** SC 13D/A
- **SEC ACT:** 1934 Act
- **SEC FILE NUMBER:** 005-89356
- **FILM NUMBER:** 23625661

**BUSINESS ADDRESS:**
- **STREET 1:** 8 RUE DE LA CROIX JARRY
- **CITY:** PARIS, LLE-DE-FRANCE
- **STATE:** I0
- **ZIP:** 75013
- **BUSINESS PHONE:** 33181691600

**MAIL ADDRESS:**
- **STREET 1:** 8 RUE DE LA CROIX JARRY
- **CITY:** PARIS, LLE-DE-FRANCE
- **STATE:** I0
- **ZIP:** 75013
**FILED BY**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** Bpifrance Participations SA
- **CENTRAL INDEX KEY:** 0001581835
- **IRS NUMBER:** 000000000
- **FISCAL YEAR END:** 1231

**FILING VALUES:**
- **FORM TYPE:** SC 13D/A

**BUSINESS ADDRESS:**
- **STREET 1:** 27-31 AVENUE DU GENERAL LECLERC
- **CITY:** MAISONS ALFORT, CEDEX
- **STATE:** I0
- **ZIP:** 94710
- **BUSINESS PHONE:** 33153898789

**MAIL ADDRESS:**
- **STREET 1:** 27-31 AVENUE DU GENERAL LECLERC
- **CITY:** MAISONS ALFORT, CEDEX
- **STATE:** I0
- **ZIP:** 94710

---

| |
|:---|
| **UNITED STATES** |
| **SECURITIES AND EXCHANGE COMMISSION** |
| **Washington, D.C. 20549** |

---

**SCHEDULE 13D**

**Under the Securities Exchange Act of 1934<br> (Amendment No. 2)\***

**Cellectis S.A.**

(Name of Issuer)

**Ordinary Shares, nominal value €0.05 per share**

(Title of Class of Securities)

**15117K103 (American Depositary Shares, each representing one Ordinary Share)**

(CUSIP Number)

**Sophie Paquin**

**Bpifrance Participations S.A.**

**6-8, boulevard Haussmann**

**75009 Paris**

**France**

**+33 1 53 89 55 73**

**With copy to:**

 **John C. Partigan**

**Nixon Peabody LLP**

**799 9th Street NW, Suite 500**

**Washington, DC 20001**

**(202) 585-8000**

((Name, Address and Telephone Number of Person<br> Authorized to Receive Notices and Communications)

**February 7, 2023**

(Date of Event Which Requires Filing of this Statement)

If the filing person has previously filed a statement on Schedule 13G to report the acquisition that is the subject of this Schedule 13D, and is filing this schedule because of §§ 240.13d-1(e), 240.13d-1(f) or 240.13d-1(g), check the following box ⌧.

Note: Schedules filed in paper format shall include a signed original and five copies of the schedule, including all exhibits. See Rule 13d-7 for other parties to whom copies are to be sent.

\*The remainder of this cover page shall be filled out for a reporting person's initial filing on this form with respect to the subject class of securities, and for any subsequent amendment containing information which would alter disclosures provided in a prior cover page.

The information required on the remainder of this cover page shall not be deemed to be "filed" for the purpose of Section 18 of the Securities Exchange Act of 1934 ("Act") or otherwise subject to the liabilities of that section of the Act but shall be subject to all other provisions of the Act.

---

| | | | | |
|:---|:---|:---|:---|:---|
| CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) |
| 1 | Name of Reporting Person<br> Bpifrance Participations S.A. | Name of Reporting Person<br> Bpifrance Participations S.A. | Name of Reporting Person<br> Bpifrance Participations S.A. | Name of Reporting Person<br> Bpifrance Participations S.A. |
| 2 | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) |
|  | (a) | (a) | ◻ | ◻ |
|  | (b) | (b) | ◻ | ◻ |
| 3 | SEC Use Only | SEC Use Only | SEC Use Only | SEC Use Only |
| 4 | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC |
| 5 | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ |
| 6 | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 7 | 7 | Sole Voting Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 8 | 8 | Shared Voting Power <br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 9 | 9 | Sole Dispositive Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 10 | 10 | Shared Dispositive Power <br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| 11 | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| 12 | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ |
| 13 | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>1</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>1</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>1</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>1</sup>; 14.2% of the voting rights |
| 14 | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO |

---

<sup>1</sup> Percentage is calculated based on 55,475,610 Ordinary Shares, as reported by the Issuer in its Form 6-K filed with the SEC on February 8, 2023.

Page 2 of 7 Pages

---

| | | | | |
|:---|:---|:---|:---|:---|
| CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) |
| 1 | Name of Reporting Person<br> Caisse des dépôts et consignations | Name of Reporting Person<br> Caisse des dépôts et consignations | Name of Reporting Person<br> Caisse des dépôts et consignations | Name of Reporting Person<br> Caisse des dépôts et consignations |
| 2 | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) |
|  | (a) | (a) | ◻ | ◻ |
|  | (b) | (b) | ◻ | ◻ |
| 3 | SEC Use Only | SEC Use Only | SEC Use Only | SEC Use Only |
| 4 | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC |
| 5 | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ |
| 6 | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 7 | 7 | Sole Voting Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 8 | 8 | Shared Voting Power <br> 6,427,497 Ordinary Shares; 9,306,997 voting rights |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 9 | 9 | Sole Dispositive Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 10 | 10 | Shared Dispositive Power <br> 6,427,497 Ordinary Shares; 9,306,997 voting rights |
| 11 | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 6,427,497 Ordinary Shares; 9,306,997 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 6,427,497 Ordinary Shares; 9,306,997 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 6,427,497 Ordinary Shares; 9,306,997 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 6,427,497 Ordinary Shares; 9,306,997 voting rights |
| 12 | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ |
| 13 | Percent of Class Represented by Amount in Row (11)<br> 11.6% of the Ordinary Shares<sup>2</sup>; 15.1% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 11.6% of the Ordinary Shares<sup>2</sup>; 15.1% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 11.6% of the Ordinary Shares<sup>2</sup>; 15.1% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 11.6% of the Ordinary Shares<sup>2</sup>; 15.1% of the voting rights |
| 14 | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO |

---

<sup>2</sup> Percentage is calculated based on 55,475,610 Ordinary Shares, as reported by the Issuer in its Form 6-K filed with the SEC on February 8, 2023.

Page 3 of 7 Pages

---

| | | | | |
|:---|:---|:---|:---|:---|
| CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) |
| 1 | Name of Reporting Person<br> EPIC Bpifrance | Name of Reporting Person<br> EPIC Bpifrance | Name of Reporting Person<br> EPIC Bpifrance | Name of Reporting Person<br> EPIC Bpifrance |
| 2 | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) |
|  | (a) | (a) | ◻ | ◻ |
|  | (b) | (b) | ◻ | ◻ |
| 3 | SEC Use Only | SEC Use Only | SEC Use Only | SEC Use Only |
| 4 | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC |
| 5 | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ |
| 6 | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 7 | 7 | Sole Voting Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 8 | 8 | Shared Voting Power <br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 9 | 9 | Sole Dispositive Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 10 | 10 | Shared Dispositive Power <br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| 11 | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| 12 | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ |
| 13 | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>3</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>3</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>3</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>3</sup>; 14.2% of the voting rights |
| 14 | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO |

---

<sup>3</sup> Percentage is calculated based on 55,475,610 Ordinary Shares, as reported by the Issuer in its Form 6-K filed with the SEC on February 8, 2023.

Page 4 of 7 Pages

---

| | | | | |
|:---|:---|:---|:---|:---|
| CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) | CUSIP No. 15117K103 (American Depositary Shares, each representing one Ordinary Share) |
| 1 | Name of Reporting Person<br> Bpifrance S.A. | Name of Reporting Person<br> Bpifrance S.A. | Name of Reporting Person<br> Bpifrance S.A. | Name of Reporting Person<br> Bpifrance S.A. |
| 2 | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) | Check the Appropriate Box if a Member of a Group (See Instructions) |
|  | (a) | (a) | ◻ | ◻ |
|  | (b) | (b) | ◻ | ◻ |
| 3 | SEC Use Only | SEC Use Only | SEC Use Only | SEC Use Only |
| 4 | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC | Source of Funds <br> WC |
| 5 | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ | Check Box if Disclosure of Legal Proceedings Is Required Pursuant to Items 2(d) or 2(e) ◻ |
| 6 | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France | Citizenship or Place of Organization <br> France |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 7 | 7 | Sole Voting Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 8 | 8 | Shared Voting Power <br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 9 | 9 | Sole Dispositive Power <br> 0 |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With |  |  |  |
| Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | Number of<br> Shares<br> Beneficially<br> Owned by<br> Each<br> Reporting<br> Person With | 10 | 10 | Shared Dispositive Power <br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| 11 | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights | Aggregate Amount Beneficially Owned by Each Reporting Person<br> 5,873,247 Ordinary Shares; 8,752,747 voting rights |
| 12 | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ | Check Box if the Aggregate Amount in Row (11) Excludes Certain Shares (See Instructions) ◻ |
| 13 | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>4</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>4</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>4</sup>; 14.2% of the voting rights | Percent of Class Represented by Amount in Row (11)<br> 10.6% of the Ordinary Shares<sup>4</sup>; 14.2% of the voting rights |
| 14 | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO | Type of Reporting Person (See Instructions)<br> OO |

---

<sup>4</sup> Percentage is calculated based on 55,475,610 Ordinary Shares, as reported by the Issuer in its Form 6-K filed with the SEC on February 8, 2023.

Page 5 of 7 Pages

**Amendment No. 2 to Schedule 13D**

This Amendment No. 2 ("Amendment No. 2") amends and supplements the Schedule 13D filed with the Securities and Exchange Commission (the "SEC") on August 4, 2020, as amended by Amendment No. 1 filed with the SEC on February 12, 2021 (collectively, the "Schedule 13D"). Except as specifically provided herein, this Amendment No. 2 does not modify any of the information previously reported in the Schedule 13D. Capitalized terms used herein shall have the meanings ascribed to them in the Schedule 13D.

**Item 4. Purpose of Transaction.**

Item 4 of the Schedule 13D is hereby amended and supplemented as follows:

The percentage ownership of Ordinary Shares is based on 54,367,810 Ordinary Shares outstanding after the offering, as reported by the Issuer in the Prospectus Supplement filed pursuant to Rule 424(b)(5) filed on February 2, 2023, and an additional 1,107,800 Ordinary Shares as a result of the exercise by the underwriters of their option to purchase additional shares, as reported by the Issuer in its Form 6-K filed with the SEC on February 8, 2023.

Bpifrance Participations participated in the offering and purchased 2,186,960 American Depositary Shares, or ADSs, at the offering price of $2.50 per ADS. Each ADS represents one Ordinary Share.

All of the Ordinary Shares that are held of record by the Reporting Persons as reported herein were acquired for investment purposes. The Reporting Persons retain the right to change their investment intent, from time to time to acquire additional Ordinary Shares or other securities of the Issuer, or to sell or otherwise dispose of all or part of the Ordinary Shares or other securities of the Issuer, if any, beneficially owned by them, in any manner permitted by law. The Reporting Persons may engage from time to time in ordinary course transactions with financial institutions with respect to the securities described herein.

Except as set forth above, none of the Reporting Persons currently has any plans or proposals which would be related to or would result in any of the matters described in Items 4(a)-(j) of the Instructions to Schedule 13D. However, as part of the ongoing evaluation of investment and investment alternatives, the Reporting Persons may consider such matters and, subject to applicable law, may formulate a plan with respect to such matters, and, from time to time, may hold discussions with or make formal proposals to management or the board of directors of the Issuer or other third parties regarding such matters.

**Item 5. Interest in Securities of the Issuer.**

Item 5(c) of the Schedule 13D is hereby amended and supplemented as follows:

Except as described above, there have been no reportable transactions with respect to the Ordinary Shares of the Issuer within the last 60 days by the Reporting Persons.

 **Item 7. Material to Be Filed as Exhibits.**

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| | |
|:---|:---|
| **Exhibit** | **Description** |
| [99.1](https://www.sec.gov/Archives/edgar/data/1627281/000110465921021688/tm216133d3_ex99-1.htm) | [Joint Filing Agreement, dated as of February 12, 2021, by and among the Reporting Persons (incorporate by reference to Exhibit 99.1 to Amendment No. 1 to Schedule 13D filed on February 12, 2021).](https://www.sec.gov/Archives/edgar/data/1627281/000110465921021688/tm216133d3_ex99-1.htm) |

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Page 6 of 7 Pages

**SIGNATURE**

After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.

Dated: February 14, 2023

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| | |
|:---|:---|
| **Bpifrance Participations S.A.** | **Bpifrance Participations S.A.** |
| By: | /s/ Sophie Paquin |
| Name: | Sophie Paquin |
| Title: | Director of Legal Affairs |
| **Caisse des dépôts et consignations** | **Caisse des dépôts et consignations** |
| By: | /s/ Laurence Giraudon |
| Name: | Laurence Giraudon |
| Title: | Chief Operating Officer, Finance and Operations Department, Asset Management Division |
| **EPIC Bpifrance** | **EPIC Bpifrance** |
| By: | /s/ Sophie Paquin |
| Name: | Sophie Paquin |
| Title: | Director of Legal Affairs |
| **Bpifrance S.A.** | **Bpifrance S.A.** |
| By: | /s/ Boubakar Dione |
| Name: | Boubakar Dione |
| Title: | Group Director of Legal Affairs |

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Page 7 of 7 Pages

**APPENDIX A**

**Name, business address, present principal occupation or employment and place of citizenship of the directors and executive officers of**

**BPIFRANCE PARTICIPATIONS S.A.**

The name, business address and present principal occupation or employment of each of the directors and executive officers of Bpifrance Participations S.A. are set forth below. The business address of each director and executive officer is Bpifrance Participations S.A., 27-31, avenue du Général Leclerc, 94710 Maisons-Alfort Cedex, France. Unless otherwise indicated, each director and executive officer is a citizen of France.

**BOARD OF DIRECTORS**

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| | |
|:---|:---|
| **Name** | **Present Principal Occupation or Employment** |
| NICOLAS DUFOURCQ | **Director, Chairman, Chief Executive Officer of Bpifrance Participations**, and Chief Executive Officer of Bpifrance |
| CAROLE ABBEY DUVAL | **Director**, Head of the Steering Department at Caisse des Dépôts |
| ANTOINE SAINTOYANT | **Director,** Director of strategic holdings at Caisse des Dépôts |
| FREDERIC SAINT-GEOURS | **Director**, Director of Société nationale SNCF |
| FRENCH STATE, represented by CHARLES SARRAZIN | **Director**, in charge of shareholding interests in the Service & Finance sectors, Agence des Participations de l'Etat (French State Shareholding Agency) |
| CONSTANCE VALIGNY | **Director**, Assistant Director for macroeconomic policies, Direction Générale du Trésor (French Treasury) |
| CHLOE MAYENOBE | **Director**, deputy CEO Natixis Payment |
| VICTOIRE AUBRY | **Director**, Chief Financial Officer of Icade |
| SOPHIE STABILE | **Director**, Founder and CEO of Révérence |
| ROMAIN BONENFANT | **Director,** Chief, Service of Industry, General Directorate for Companies of the Ministry of Economy and Finance |

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**EXECUTIVE OFFICERS**

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| | |
|:---|:---|
| **Name** | **Present Principal Occupation or Employment** |
| NICOLAS DUFOURCQ | **Chief Executive Officer** |
| JOSÉ GONZALO | **Executive Director** |
| PIERRE BENEDETTI | **Chief Financial Officer** |

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**APPENDIX B**

**Name, business address, present principal occupation or employment and place of citizenship of the directors and executive officers of**

**CDC CROISSANCE S.A.**

The name, business address and present principal occupation or employment of each of each of the directors and executive officers of CDC Croissance S.A. are set forth below. The business address of each director and executive officer is CDC Croissance S.A., c/o 209, rue de l'Université, 75007 Paris, France. Unless otherwise indicated, each such person is a citizen of France.

**BOARD OF DIRECTORS**

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| | |
|:---|:---|
| **Name** | **Present Principal Occupation or Employment** |
| Olivier Mareuse | **Director of Asset Management and Savings Funds at Caisse des Dépôts. Deputy Director General** |
| Virginie Chapron-DU JEU | **Director of Risk at the Caisse des Dépôts Group** |
| LAURENT DEBORDE |  |
| françois-louis ricard |  |
| nicolas fourt |  |

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**EXECUTIVE OFFICERS**

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| | |
|:---|:---|
| **Name** | **Present Principal Occupation or Employment** |
| Christophe Bourdillon |  |

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**APPENDIX C**

**Name, business address, present principal occupation or employment and place of citizenship of the directors and executive officers of**

**CAISSE DES DÉPÔTS**

The name, business address and present principal occupation or employment of each of the members of the Management Committee of Caisse des Dépôts are set forth below. The business address of each director and executive officer is Caisse des Dépôts, c/o 56, rue de Lille, 75007 Paris, France. Unless otherwise indicated, each such person is a citizen of France.

**MANAGEMENT COMMITTEE**

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| | |
|:---|:---|
| **Name** | **Present Principal Occupation or Employment** |
| ERIC LOMBARD | **Chief Executive Officer of Caisse des Dépôts** |
| OLIVIER SICHEL | **Deputy Chief Executive Officer of Caisse des Dépôts** |
| NICOLAS DUFOURCQ | **Chief Executive Officer of Bpifrance** |
| VIRGINIE CHAPRON-DU JEU | **Director of Risk at the Caisse des Dépôts Group** |
| PIERRE CHEVALIER | **Head of Legal and Tax Department** |
| NATHALIE TUBIANA | **Director of Finance at the Caisse des Dépôts Group** |
| OLIVIER MAREUSE | **Director of Asset Management and Savings Funds at Caisse des Dépôts. Deputy Director General** |
| CATHERINE MAYENOBE | **Director of Operations and Business Transformation Management. Deputy Managing Director** |
| AURELIE ROBINEAU-ISRAEL | **Caisse des Dépôts Group Human Resources Director** |
| SOPHIE QUATREHOMME | **Group Corporate Communications Director** |
| MICHEL YAHIEL | **Pensions and Solidarity Director** |
| ANTOINE SAINTOYANT | **Director of strategic holdings at Caisse des Dépôts** |

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| | |
|:---|:---|
| MARIE-LAURE GADRAT | **Chief of Staff to the Director Général** |

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**APPENDIX D**

**Name, business address, present principal occupation or employment and place of citizenship of the directors and executive officers of**

**EPIC BPIFRANCE**

The name, business address and present principal occupation or employment of each of the directors and executive officers of EPIC Bpifrance are set forth below. The business address of each director and executive officer is EPIC Bpifrance, 27-31, avenue du Général Leclerc, 94710 Maisons-Alfort Cedex, France. Unless otherwise indicated, each director and executive officer is a citizen of France.

**DIRECTORS**

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| | |
|:---|:---|
| **Name** | **Present Principal Occupation or Employment** |
| CHRISTIAN BODIN | **Director, Chairman, Chief Executive Officer of EPIC Bpifrance** |
| PIERRE-LOUIS AUTIN | **Director**, Head of department SITTAR at the Higher Education, Research and Innovation Ministry |
| YANN POUËZAT | **Director**, Deputy Director at the Directorate of Financing, Industry and Market of the Ministry of Economy and Finance |
| ALBAN HAUTIER | **Director**, Deputy Director at the General Directorate for Budget of the Ministry of Economy and Finance |
| VINCENT TEJEDOR | **Director**, Assistant Director in charge of the development of enterprises, General Directorate for Companies of the Ministry of Economy and Finance |
| NAOMI PERES | **Director**, Deputy General Secretary for Public Investment |

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**APPENDIX E**

**Name, business address, present principal occupation or employment and place of citizenship of the directors and executive officers of**

**BPIFRANCE S.A.**

The name, business address and present principal occupation or employment of each of the directors and executive officers of Bpifrance S.A. are set forth below. The business address of each director and executive officer is Bpifrance S.A., 27-31, avenue du Général Leclerc, 94710 Maisons-Alfort Cedex, France. Unless otherwise indicated, each director and executive officer is a citizen of France.

**DIRECTORS**

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| | |
|:---|:---|
| **Name** | **Present Principal Occupation or Employment** |
| ERIC LOMBARD | **Director, Chairman,** Chief Executive Officer of the Caisse des Dépôts |
| NICOLAS DUFOURCQ | **Director, Chief Executive Officer of Bpifrance** and Director, Chairman and Chief Executive Officer of Bpifrance Participations |
| MARTIN VIAL | **Director**, Chairman of the Agence des Participations de l'Etat (French State Shareholding Agency) |
| MARIE DELEAGE | **Director representing the employees of Bpifrance** |
| PHILIPPE BAYEUX | **Director representing the employees of Bpifrance** |
| VIRGINIE CHAPRON-DU JEU | **Director**, **Director of Finance for the Caisse des Dépôts Group** |
| CLAIRE DUMAS | **Director**, Finance Director at Société Générale |

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| | |
|:---|:---|
| CATHERINE LAGNEAU | **Director**, Deputy Director of the General council of economy |
| ANTOINE SAINTOYANT | **Director**, Director of strategic holdings at Caisse des Dépôts |
| BERNARD DELPIT | **Director**, Chief Financial Officer of Safran |
| ANNABEL ANDRE | **Director**, Vice-president of the Regional Council of Auvergne-Rhone-Alpes, delegate for the economy and companies |
| HAROLD HUWART | **Director**, Vice-president of the Regional Council of Centre-Val-de-Loire, in charge of the economy, farming and associations |
| CAROLE ABBEY DUVAL | **Director**, Head of the Steering Department at Caisse des Dépôts |
| DIANE SIMIU | **Director**, deputy of the general commissioner for sustainable development at the Ministry for economic and solidarity<br> transition |

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