# EDGAR Filing Document

**Accession Number:** 0001097519
**File Stem:** 0001193125-25-252371
**Filing Date:** 2025-10
**Character Count:** 40961
**Document Hash:** a7aa61180d9692374952712563b2150f
**Contains OCR:** False
**Source Format:** 

## Filing Content

## Filing Summary
**0001193125-25-252371.hdr.sgml**: 20251028

**ACCESSION NUMBER**: 0001193125-25-252371

**CONFORMED SUBMISSION TYPE**: NPORT-P

**PUBLIC DOCUMENT COUNT**: 1

**CONFORMED PERIOD OF REPORT**: 20250831

**FILED AS OF DATE**: 20251028

**DATE AS OF CHANGE**: 20251028

**PERIOD START**: 20260228

**FILER**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** COLUMBIA FUNDS SERIES TRUST
- **CENTRAL INDEX KEY:** 0001097519

**ORGANIZATION NAME:**
- **EIN:** 000000000
- **STATE OF INCORPORATION:** DE
- **FISCAL YEAR END:** 0331

**FILING VALUES:**
- **FORM TYPE:** NPORT-P
- **SEC ACT:** 1940 Act
- **SEC FILE NUMBER:** 811-09645
- **FILM NUMBER:** 251421427

**BUSINESS ADDRESS:**
- **STREET 1:** 290 CONGRESS STREET
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02210
- **BUSINESS PHONE:** 800-345-6611

**MAIL ADDRESS:**
- **STREET 1:** 290 CONGRESS STREET
- **CITY:** BOSTON
- **STATE:** MA
- **ZIP:** 02210

**FORMER COMPANY:**
- **FORMER CONFORMED NAME:** NATIONS FUNDS TRUST
- **DATE OF NAME CHANGE:** 19991022

## Series and Classes Contracts Data

### Columbia Small Cap Value and Inflection Fund (Series ID: S000010793)

| Class ID   | Class Name                                                         | Ticker Symbol   |
|:---|:---|:---|
| C000029855 | Columbia Small Cap Value and Inflection Fund Class A               | COVAX           |
| C000029858 | Columbia Small Cap Value and Inflection Fund Class R               | CCTRX           |
| C000029859 | Columbia Small Cap Value and Inflection Fund Institutional Class   | NSVAX           |
| C000122579 | Columbia Small Cap Value and Inflection Fund Institutional 2 Class | CRRRX           |
| C000122580 | Columbia Small Cap Value and Inflection Fund Institutional 3 Class | CRRYX           |
| C000253705 | Columbia Small Cap Value and Inflection Fund Class S               | NSVBX           |

## Form NPORT-P: Monthly Portfolio Investments Report

### NPORT-P: Part A: General Information

**Item A.1. Information about the Registrant.**

- **a. Name of Registrant:** Columbia Funds Series Trust

- **b. Investment Company Act file number:** 811-09645

- **c. CIK number of Registrant:** 0001097519

- **d. LEI of Registrant:** 549300712V4UL3XCAB43

- **e. Address and telephone number of Registrant.**

  - **Street Address 1:** 290 Congress Street

  - **City:** Boston

  - **State:** MA

  - **Foreign country:** US

  - **Zip / Postal Code:** 02110

  - **Telephone number:** 800-345-6611

**Item A.2. Information about the Series.**

- **a. Name of Series:** Columbia Small Cap Value and Inflection Fund

- **b. EDGAR series identifier (if any):** S000010793

- **c. LEI of Series:** CMWWRPUO7POPOTWW1R92

**Item A.3. Reporting period.**

- **a. Date of fiscal year-end:** 2026-02-28

- **b. Date as of which information is reported:** 2025-08-31

**Item A.4. Final filing**

Does the Fund anticipate that this will be its final filing on Form N-PORT? **No**

### Fund Information

**Total Assets:** $887330586.95

**Total Liabilities:** $377315.43

**Net Assets:** $886953271.52

**Cash Not Reported:** $911.00

**Monthly Return Information**

| Class               | Month 1 Return (%)   | Month 2 Return (%)   | Month 3 Return (%)   |
|:---|:---|:---|:---|
| Class ID C000029855 | 4.86%                | 1.25%                | 8.24%                |
| Class ID C000029858 | 4.82%                | 1.22%                | 8.21%                |
| Class ID C000029859 | 4.93%                | 1.20%                | 8.25%                |
| Class ID C000122579 | 4.91%                | 1.29%                | 8.26%                |
| Class ID C000122580 | 4.95%                | 1.21%                | 8.27%                |
| Class ID C000253705 | 4.93%                | 1.20%                | 8.25%                |

**Monthly Gains & Losses**

| Period   | Net Realized Gain/Loss   | Net Unrealized Appreciation/Depreciation   |
|:---|:---|:---|
| Month 1  | $14117506.87             | $24599820.24                               |
| Month 2  | $26461933.50             | $-15851700.66                              |
| Month 3  | $14258923.55             | $53138134.68                               |

**Designated Index Information**

- **Index Name:** N/A

- **Index Identifier:** N/A

### Schedule of Portfolio Investments

| Name                                    | Title                                         | Identifiers                                   | Payoff Profile   | Asset Category   | Issuer Category   | Country   |   Balance | Units   | Value (USD)   | % of Net Assets   | Maturity Date   | Coupon Type   | Annualized Rate (%)   | Restricted?   |   Fair Value Level | Lending Status   |
|:---|:---|:---|:---|:---|:---|:---|---:|:---|:---|:---|:---|:---|:---|:---|---:|:---|
| ANI PHARMACEUTICALS INC                 | ANI Pharmaceuticals Inc                       | CUSIP: 00182C103<br>LEI: 5493000T6CXTND40YH31 | Long             | EC               | CORP              | US        |     71360 | NS      | $6672160.00   | 0.75%             |  |  |  | No            |                  1 | On Loan: No      |
| ADAPTIVE BIOTECHNOLOGIES CORP           | Adaptive Biotechnologies Corp                 | CUSIP: 00650F109<br>LEI: 549300ZTF7OT1FW66Q96 | Long             | EC               | CORP              | US        |    304540 | NS      | $4013837.20   | 0.45%             |  |  |  | No            |                  1 | On Loan: No      |
| CTS CORP                                | CTS Corp                                      | CUSIP: 126501105<br>LEI: 549300YV1HQLBVHOI649 | Long             | EC               | CORP              | US        |    115100 | NS      | $4890599.00   | 0.55%             |  |  |  | No            |                  1 | On Loan: No      |
| CNX RESOURCES CORP                      | CNX Resources Corp                            | CUSIP: 12653C108<br>LEI: 549300JK3KH8PWM3B226 | Long             | EC               | CORP              | US        |    356520 | NS      | $10410384.00  | 1.17%             |  |  |  | No            |                  1 | On Loan: No      |
| ADAPTHEALTH CORP                        | AdaptHealth Corp                              | CUSIP: 00653Q102<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    542790 | NS      | $5151077.10   | 0.58%             |  |  |  | No            |                  1 | On Loan: No      |
| ALASKA AIR GRP INC.                     | Alaska Air Group Inc                          | CUSIP: 011659109<br>LEI: 529900G48RT4KWLC6C27 | Long             | EC               | CORP              | US        |    146115 | NS      | $9173099.70   | 1.03%             |  |  |  | No            |                  1 | On Loan: No      |
| AMERICAN ASSETS TRUST INC               | American Assets Trust Inc                     | CUSIP: 024013104<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    224267 | NS      | $4687180.30   | 0.53%             |  |  |  | No            |                  1 | On Loan: No      |
| AMICUS THERAPEUTICS INC                 | Amicus Therapeutics Inc                       | CUSIP: 03152W109<br>LEI: 549300MRD1SMUTNBPQ53 | Long             | EC               | CORP              | US        |    409640 | NS      | $3109167.60   | 0.35%             |  |  |  | No            |                  1 | On Loan: No      |
| CABOT CORP                              | Cabot Corp                                    | CUSIP: 127055101<br>LEI: K05C0SER542GQ6VLRO68 | Long             | EC               | CORP              | US        |    112427 | NS      | $9169546.12   | 1.03%             |  |  |  | No            |                  1 | On Loan: No      |
| CAVCO INDUSTRIES INC                    | Cavco Industries Inc                          | CUSIP: 149568107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |     12470 | NS      | $6615210.30   | 0.75%             |  |  |  | No            |                  1 | On Loan: No      |
| AMKOR TECH INC                          | Amkor Technology Inc                          | CUSIP: 031652100<br>LEI: 529900VHLRTKPWZJBM84 | Long             | EC               | CORP              | US        |    388930 | NS      | $9408216.70   | 1.06%             |  |  |  | No            |                  1 | On Loan: No      |
| CENTRAL GARDEN & PET CO                 | Central Garden & Pet Co                       | CUSIP: 153527205<br>LEI: 5493004DVZLD37DJ0H83 | Long             | EC               | CORP              | US        |     60518 | NS      | $1998909.54   | 0.23%             |  |  |  | No            |                  1 | On Loan: No      |
| CLEAR SECURE INC                        | Clear Secure Inc                              | CUSIP: 18467V109<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    257403 | NS      | $9346302.93   | 1.05%             |  |  |  | No            |                  1 | On Loan: No      |
| COLUMBIA BANKING SYST INC.              | Columbia Banking System Inc                   | CUSIP: 197236102<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    353110 | NS      | $9452754.70   | 1.07%             |  |  |  | No            |                  1 | On Loan: No      |
| COLUMBIA SHORT TERM CASH FUND           | COLUMBIA SHORT TERM CASH FUND                 | CUSIP: 19766H239<br>LEI: N/A                  | Long             | STIV             | RF                | US        |    586756 | NS      | $586579.79    | 0.07%             |  |  |  | No            |                  1 | On Loan: No      |
| COMMERCIAL METALS COMPANY               | Commercial Metals Co                          | CUSIP: 201723103<br>LEI: 549300OQS2LO07ZJ7N73 | Long             | EC               | CORP              | US        |    242930 | NS      | $14009773.10  | 1.58%             |  |  |  | No            |                  1 | On Loan: No      |
| COUSINS PROPERTIES INC                  | Cousins Properties Inc                        | CUSIP: 222795502<br>LEI: 5493007XPYD5EJABN062 | Long             | EC               | CORP              | US        |    298901 | NS      | $8814590.49   | 0.99%             |  |  |  | No            |                  1 | On Loan: No      |
| CRANE NXT CO                            | Crane NXT Co                                  | CUSIP: 224441105<br>LEI: 549300EXNPSXEWUD3N21 | Long             | EC               | CORP              | US        |    181884 | NS      | $10863931.32  | 1.22%             |  |  |  | No            |                  1 | On Loan: No      |
| CYTOKINETICS INCORPORATED               | Cytokinetics Inc                              | CUSIP: 23282W605<br>LEI: 549300BE5DY1HW7IHA64 | Long             | EC               | CORP              | US        |    157900 | NS      | $5578607.00   | 0.63%             |  |  |  | No            |                  1 | On Loan: No      |
| DIGI INTL INC.                          | Digi International Inc                        | CUSIP: 253798102<br>LEI: 529900KG656HGROI4N27 | Long             | EC               | CORP              | US        |    139590 | NS      | $4846564.80   | 0.55%             |  |  |  | No            |                  1 | On Loan: No      |
| DIODES INC                              | Diodes Inc                                    | CUSIP: 254543101<br>LEI: 549300XP2X1ARVH3BX23 | Long             | EC               | CORP              | US        |    173410 | NS      | $9439573.35   | 1.06%             |  |  |  | No            |                  1 | On Loan: No      |
| DOUBLEVERIFY HOLDINGS INC               | DoubleVerify Holdings Inc                     | CUSIP: 25862V105<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    375570 | NS      | $6110523.90   | 0.69%             |  |  |  | No            |                  1 | On Loan: No      |
| DYCOM INDUSTRIES INC                    | Dycom Industries Inc                          | CUSIP: 267475101<br>LEI: 549300F56EKTHQI3WT60 | Long             | EC               | CORP              | US        |     21055 | NS      | $5315755.85   | 0.60%             |  |  |  | No            |                  1 | On Loan: No      |
| DYNEX CAPITAL INC                       | Dynex Capital Inc                             | CUSIP: 26817Q886<br>LEI: 5CIJ4S9HAEBWILA19B65 | Long             | EC               | CORP              | US        |    499260 | NS      | $6300661.20   | 0.71%             |  |  |  | No            |                  1 | On Loan: No      |
| ECOVYST INC                             | Ecovyst Inc                                   | CUSIP: 27923Q109<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    365640 | NS      | $3323667.60   | 0.37%             |  |  |  | No            |                  1 | On Loan: No      |
| EPLUS INC                               | ePlus Inc                                     | CUSIP: 294268107<br>LEI: 529900FFY8AAFJ21NE91 | Long             | EC               | CORP              | US        |     90740 | NS      | $6566853.80   | 0.74%             |  |  |  | No            |                  1 | On Loan: No      |
| EVERTEC INC                             | EVERTEC Inc                                   | CUSIP: 30040P103<br>LEI: N/A                  | Long             | EC               | CORP              | PR        |    238780 | NS      | $8519670.40   | 0.96%             |  |  |  | No            |                  1 | On Loan: No      |
| FIRSTCASH HOLDINGS INC                  | FirstCash Holdings Inc                        | CUSIP: 33768G107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |     76510 | NS      | $11267627.70  | 1.27%             |  |  |  | No            |                  1 | On Loan: No      |
| FULTON FINL CORP                        | Fulton Financial Corp                         | CUSIP: 360271100<br>LEI: 5493005QLZ6O1BNC0C11 | Long             | EC               | CORP              | US        |    702513 | NS      | $13811405.58  | 1.56%             |  |  |  | No            |                  1 | On Loan: No      |
| GENTHERM INC                            | Gentherm Inc                                  | CUSIP: 37253A103<br>LEI: HKM6YX1U5JS60TCUB219 | Long             | EC               | CORP              | US        |    110120 | NS      | $4049112.40   | 0.46%             |  |  |  | No            |                  1 | On Loan: No      |
| SHOALS TECHNOLOGIES GROUP INC           | Shoals Technologies Group Inc                 | CUSIP: 82489W107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    579310 | NS      | $3771308.10   | 0.43%             |  |  |  | No            |                  1 | On Loan: No      |
| SOLENO THERAPEUTICS INC                 | Soleno Therapeutics Inc                       | CUSIP: 834203309<br>LEI: 5493003LCY77N7FXK844 | Long             | EC               | CORP              | US        |     30680 | NS      | $2076115.60   | 0.23%             |  |  |  | No            |                  1 | On Loan: No      |
| STONEX GROUP INC                        | StoneX Group Inc                              | CUSIP: 861896108<br>LEI: 549300JUF07L8VF02M60 | Long             | EC               | CORP              | US        |     89005 | NS      | $9093640.85   | 1.03%             |  |  |  | No            |                  1 | On Loan: No      |
| SUNRUN INC                              | Sunrun Inc                                    | CUSIP: 86771W105<br>LEI: 54930007SJ77CI66U531 | Long             | EC               | CORP              | US        |    462040 | NS      | $7378778.80   | 0.83%             |  |  |  | No            |                  1 | On Loan: No      |
| TTM TECH INC                            | TTM Technologies Inc                          | CUSIP: 87305R109<br>LEI: 549300SC4BDOIS008048 | Long             | EC               | CORP              | US        |    189160 | NS      | $8430861.20   | 0.95%             |  |  |  | No            |                  1 | On Loan: No      |
| TAYLOR MORRISON HOME CORP               | Taylor Morrison Home Corp                     | CUSIP: 87724P106<br>LEI: 5299009UDLLGLKOOJE30 | Long             | EC               | CORP              | US        |    191903 | NS      | $12928505.11  | 1.46%             |  |  |  | No            |                  1 | On Loan: No      |
| TELADOC HEALTH INC                      | Teladoc Health Inc                            | CUSIP: 87918A105<br>LEI: 549300I2PVFBC9MFAC61 | Long             | EC               | CORP              | US        |    621920 | NS      | $4807441.60   | 0.54%             |  |  |  | No            |                  1 | On Loan: No      |
| TELEPHONE AND DATA SYSTEMS INC          | Telephone and Data Systems Inc                | CUSIP: 879433829<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    293360 | NS      | $11760802.40  | 1.33%             |  |  |  | No            |                  1 | On Loan: No      |
| TIDEWATER INC                           | Tidewater Inc                                 | CUSIP: 88642R109<br>LEI: 2549000JA5GRUPLMUH98 | Long             | EC               | CORP              | US        |    153030 | NS      | $9212406.00   | 1.04%             |  |  |  | No            |                  1 | On Loan: No      |
| TREEHOUSE FOODS INC                     | TreeHouse Foods Inc                           | CUSIP: 89469A104<br>LEI: 5493007M7EVHE470Q576 | Long             | EC               | CORP              | US        |    204130 | NS      | $3743744.20   | 0.42%             |  |  |  | No            |                  1 | On Loan: No      |
| UGI CORP                                | UGI Corp                                      | CUSIP: 902681105<br>LEI: DX6GCWD4Q1JO9CRE5I40 | Long             | EC               | CORP              | US        |    167730 | NS      | $5810167.20   | 0.66%             |  |  |  | No            |                  1 | On Loan: No      |
| ULTRA CLEAN HLD                         | Ultra Clean Holdings Inc                      | CUSIP: 90385V107<br>LEI: 5493007L6LS7QX18PE75 | Long             | EC               | CORP              | US        |    140920 | NS      | $3384898.40   | 0.38%             |  |  |  | No            |                  1 | On Loan: No      |
| URBAN OUTFITTERS INC                    | Urban Outfitters Inc                          | CUSIP: 917047102<br>LEI: 549300N3DUFR5U6NRN38 | Long             | EC               | CORP              | US        |     97860 | NS      | $6564448.80   | 0.74%             |  |  |  | No            |                  1 | On Loan: No      |
| VISTEON CORP                            | Visteon Corp                                  | CUSIP: 92839U206<br>LEI: 549300MOVLYHRW4GGW78 | Long             | EC               | CORP              | US        |     74025 | NS      | $9176139.00   | 1.03%             |  |  |  | No            |                  1 | On Loan: No      |
| VONTIER CORP                            | Vontier Corp                                  | CUSIP: 928881101<br>LEI: 549300VVSQLG4P5K4639 | Long             | EC               | CORP              | US        |    190144 | NS      | $8159079.04   | 0.92%             |  |  |  | No            |                  1 | On Loan: No      |
| WAFD INC                                | WaFd Inc                                      | CUSIP: 938824109<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    351345 | NS      | $11049800.25  | 1.25%             |  |  |  | No            |                  1 | On Loan: No      |
| WORLD KINECT CORPORATION                | World Kinect Corp                             | CUSIP: 981475106<br>LEI: D3W7PCXCBRQLL17DZ313 | Long             | EC               | CORP              | US        |    230180 | NS      | $6171125.80   | 0.70%             |  |  |  | No            |                  1 | On Loan: No      |
| WORTHINGTON ENTERPRISES INC             | Worthington Enterprises Inc                   | CUSIP: 981811102<br>LEI: 1WRCIANKYOIK6KYE5E82 | Long             | EC               | CORP              | US        |    184980 | NS      | $12171684.00  | 1.37%             |  |  |  | No            |                  1 | On Loan: No      |
| XENIA HOTELS & RESORTS INC              | Xenia Hotels & Resorts Inc                    | CUSIP: 984017103<br>LEI: 54930075IWIPHF5IQL62 | Long             | EC               | CORP              | US        |    328400 | NS      | $4640292.00   | 0.52%             |  |  |  | No            |                  1 | On Loan: No      |
| ASSURED GUARANTY LTD                    | Assured Guaranty Ltd                          | CUSIP: G0585R106<br>LEI: 549300BBOQUJEPH87M14 | Long             | EC               | CORP              | BM        |    136469 | NS      | $11217751.80  | 1.26%             |  |  |  | No            |                  1 | On Loan: No      |
| BANK OF N.T. BUTTERFIELD & SON LTD      | Bank of NT Butterfield & Son Ltd/The          | CUSIP: G0772R208<br>LEI: 549300V8RVM0EMTDPF28 | Long             | EC               | CORP              | BM        |    186990 | NS      | $8436988.80   | 0.95%             |  |  |  | No            |                  1 | On Loan: No      |
| CUSHMAN & WAKEFIELD PLC                 | Cushman & Wakefield PLC                       | CUSIP: G2717B108<br>LEI: N/A                  | Long             | EC               | CORP              | GB        |   1141610 | NS      | $18003189.70  | 2.03%             |  |  |  | No            |                  1 | On Loan: No      |
| DOLE PLC                                | Dole PLC                                      | CUSIP: G27907107<br>LEI: N/A                  | Long             | EC               | CORP              | IE        |    391170 | NS      | $5758022.40   | 0.65%             |  |  |  | No            |                  1 | On Loan: No      |
| MODINE MFG CO                           | Modine Manufacturing Co                       | CUSIP: 607828100<br>LEI: Z35SZFYJR8GZZRSIUU09 | Long             | EC               | CORP              | US        |     59360 | NS      | $8080676.80   | 0.91%             |  |  |  | No            |                  1 | On Loan: No      |
| RYDER SYSTEM INC                        | Ryder System Inc                              | CUSIP: 783549108<br>LEI: ZXJ6Y2OSPZXIVGRILD15 | Long             | EC               | CORP              | US        |     51680 | NS      | $9691033.60   | 1.09%             |  |  |  | No            |                  1 | On Loan: No      |
| Sandisk Corp/DE                         | Sandisk Corp/DE                               | CUSIP: 80004C200<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    127568 | NS      | $6693492.96   | 0.75%             |  |  |  | No            |                  1 | On Loan: No      |
| SAVERS VALUE VILLAGE INC                | Savers Value Village Inc                      | CUSIP: 80517M109<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    489070 | NS      | $5912856.30   | 0.67%             |  |  |  | No            |                  1 | On Loan: No      |
| SCOTTS MIRACLE-GRO CO/THE               | Scotts Miracle-Gro Co/The                     | CUSIP: 810186106<br>LEI: 2TH9E0JTOUXOEGI2R879 | Long             | EC               | CORP              | US        |    133487 | NS      | $8172074.14   | 0.92%             |  |  |  | No            |                  1 | On Loan: No      |
| ESSENT GROUP LTD                        | Essent Group Ltd                              | CUSIP: G3198U102<br>LEI: 254900QUO860OUT6ZF83 | Long             | EC               | CORP              | BM        |    248573 | NS      | $15595470.02  | 1.76%             |  |  |  | No            |                  1 | On Loan: No      |
| LIVANOVA PLC                            | LivaNova PLC                                  | CUSIP: G5509L101<br>LEI: 2138001K2FX6SHK2B339 | Long             | EC               | CORP              | GB        |    218280 | NS      | $12304443.60  | 1.39%             |  |  |  | No            |                  1 | On Loan: No      |
| TEEKAY TANKERS LTD                      | Teekay Tankers Ltd                            | CUSIP: G8726X106<br>LEI: 549300DE8ZAIYS2T6Q22 | Long             | EC               | CORP              | BM        |    149400 | NS      | $7345998.00   | 0.83%             |  |  |  | No            |                  1 | On Loan: No      |
| PERRIGO COMPANY PLC                     | Perrigo Co PLC                                | CUSIP: G97822103<br>LEI: 549300IFOQS6R1QLY196 | Long             | EC               | CORP              | IE        |    423550 | NS      | $10055077.00  | 1.13%             |  |  |  | No            |                  1 | On Loan: No      |
| FOREIGN TRADE BANK OF LATIN AMERICA INC | Banco Latinoamericano de Comercio Exterior SA | CUSIP: P16994132<br>LEI: 549300CN3134K4LC0651 | Long             | EC               | NUSS              | PA        |    145880 | NS      | $6729444.40   | 0.76%             |  |  |  | No            |                  1 | On Loan: No      |
| NAVIGATOR HOLDINGS LTD                  | Navigator Holdings Ltd                        | CUSIP: Y62132108<br>LEI: 213800H7RVOFYARK2W19 | Long             | EC               | CORP              | MH        |    270401 | NS      | $4350752.09   | 0.49%             |  |  |  | No            |                  1 | On Loan: No      |
| MOOG INC                                | Moog Inc                                      | CUSIP: 615394202<br>LEI: 549300WM2I4U7OBAHG07 | Long             | EC               | CORP              | US        |     41791 | NS      | $8184767.35   | 0.92%             |  |  |  | No            |                  1 | On Loan: No      |
| N-ABLE INC                              | N-able Inc/US                                 | CUSIP: 62878D100<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    660770 | NS      | $5325806.20   | 0.60%             |  |  |  | No            |                  1 | On Loan: No      |
| NATIONAL FUEL GAS CO                    | National Fuel Gas Co                          | CUSIP: 636180101<br>LEI: 8JB38FFW1Y3C1HM8E841 | Long             | EC               | CORP              | US        |     60490 | NS      | $5246902.60   | 0.59%             |  |  |  | No            |                  1 | On Loan: No      |
| NETSCOUT SYST INC                       | NetScout Systems Inc                          | CUSIP: 64115T104<br>LEI: 5493005HZXZ3SPNVCV76 | Long             | EC               | CORP              | US        |    305340 | NS      | $7599912.60   | 0.86%             |  |  |  | No            |                  1 | On Loan: No      |
| NETSTREIT CORP                          | NETSTREIT Corp                                | CUSIP: 64119V303<br>LEI: 254900CJL0EDMEQ25Q84 | Long             | EC               | CORP              | US        |    453030 | NS      | $8285918.70   | 0.93%             |  |  |  | No            |                  1 | On Loan: No      |
| NEWMARK GROUP INC                       | Newmark Group Inc                             | CUSIP: 65158N102<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    769922 | NS      | $14020279.62  | 1.58%             |  |  |  | No            |                  1 | On Loan: No      |
| ORIGIN BANCORP INC                      | Origin Bancorp Inc                            | CUSIP: 68621T102<br>LEI: 54930055DO2Z4KO3NB45 | Long             | EC               | CORP              | US        |    116030 | NS      | $4512406.70   | 0.51%             |  |  |  | No            |                  1 | On Loan: No      |
| OUTFRONT MEDIA INC                      | Outfront Media Inc                            | CUSIP: 69007J304<br>LEI: 549300RZWDYU5722I639 | Long             | EC               | CORP              | US        |    623810 | NS      | $11652770.80  | 1.31%             |  |  |  | No            |                  1 | On Loan: No      |
| PATRICK IND                             | Patrick Industries Inc                        | CUSIP: 703343103<br>LEI: 549300FZ5ZTCCJ3Z5W14 | Long             | EC               | CORP              | US        |    116133 | NS      | $12989476.05  | 1.46%             |  |  |  | No            |                  1 | On Loan: No      |
| PENGUIN SOLUTIONS INC                   | Penguin Solutions Inc                         | CUSIP: 706915105<br>LEI: 529900RDGHQY8XKXPN14 | Long             | EC               | CORP              | KY        |    236840 | NS      | $5714949.20   | 0.64%             |  |  |  | No            |                  1 | On Loan: No      |
| PENNYMAC FINANCIAL SERVICES INC         | PennyMac Financial Services Inc               | CUSIP: 70932M107<br>LEI: 549300W1D516HBDEU871 | Long             | EC               | CORP              | US        |     97606 | NS      | $10746420.60  | 1.21%             |  |  |  | No            |                  1 | On Loan: No      |
| PERDOCEO EDUCATION CORP                 | Perdoceo Education Corp                       | CUSIP: 71363P106<br>LEI: 529900X5PCA184O9LY90 | Long             | EC               | CORP              | US        |    285458 | NS      | $9345894.92   | 1.05%             |  |  |  | No            |                  1 | On Loan: No      |
| POPULAR INC                             | Popular Inc                                   | CUSIP: 733174700<br>LEI: 5493008CARDZMVQ3LO89 | Long             | EC               | CORP              | PR        |     32991 | NS      | $4144989.24   | 0.47%             |  |  |  | No            |                  1 | On Loan: No      |
| PORTLAND GENERAL ELECTRIC COMP          | Portland General Electric Co                  | CUSIP: 736508847<br>LEI: GJOUP9M7C39GLSK9R870 | Long             | EC               | CORP              | US        |    290601 | NS      | $12431910.78  | 1.40%             |  |  |  | No            |                  1 | On Loan: No      |
| POTLATCHDELTIC CORP                     | PotlatchDeltic Corp                           | CUSIP: 737630103<br>LEI: 54930084YA8UJOTXDM59 | Long             | EC               | CORP              | US        |    336890 | NS      | $14159486.70  | 1.60%             |  |  |  | No            |                  1 | On Loan: No      |
| PRICESMART INC.                         | PriceSmart Inc                                | CUSIP: 741511109<br>LEI: 549300JVYOSTGVSDZS46 | Long             | EC               | CORP              | US        |     52520 | NS      | $5633295.20   | 0.64%             |  |  |  | No            |                  1 | On Loan: No      |
| RADIAN GRP INC                          | Radian Group Inc                              | CUSIP: 750236101<br>LEI: 54930005OUP720S5U385 | Long             | EC               | CORP              | US        |    367441 | NS      | $12816342.08  | 1.44%             |  |  |  | No            |                  1 | On Loan: No      |
| RESIDEO TECHNOLOGIES INC                | Resideo Technologies Inc                      | CUSIP: 76118Y104<br>LEI: 5493006TKFPYISY2DS54 | Long             | EC               | CORP              | US        |    390176 | NS      | $13285492.80  | 1.50%             |  |  |  | No            |                  1 | On Loan: No      |
| RUSH ENTERPRISES INC                    | Rush Enterprises Inc                          | CUSIP: 781846209<br>LEI: 529900V3XHTN7A6DFT17 | Long             | EC               | CORP              | US        |    228780 | NS      | $13131972.00  | 1.48%             |  |  |  | No            |                  1 | On Loan: No      |
| AMNEAL PHARMACEUTICALS INC              | Amneal Pharmaceuticals Inc                    | CUSIP: 03168L105<br>LEI: N/A                  | Long             | EC               | CORP              | US        |   1171334 | NS      | $11197953.04  | 1.26%             |  |  |  | No            |                  1 | On Loan: No      |
| ARCBEST CORP                            | ArcBest Corp                                  | CUSIP: 03937C105<br>LEI: 5493006S3KBNE8ZS3J23 | Long             | EC               | CORP              | US        |     99106 | NS      | $7310058.56   | 0.82%             |  |  |  | No            |                  1 | On Loan: No      |
| ARDENT HEALTH INC                       | Ardent Health Inc                             | CUSIP: 03980N107<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    525860 | NS      | $6673163.40   | 0.75%             |  |  |  | No            |                  1 | On Loan: No      |
| BANKUNITED INC                          | BankUnited Inc                                | CUSIP: 06652K103<br>LEI: 96OVUWEW5D4YX6PFXD91 | Long             | EC               | CORP              | US        |    343878 | NS      | $13476578.82  | 1.52%             |  |  |  | No            |                  1 | On Loan: No      |
| GREENBRIER CO INC                       | Greenbrier Cos Inc/The                        | CUSIP: 393657101<br>LEI: 5493001GAGGFR30BP390 | Long             | EC               | CORP              | US        |    107540 | NS      | $5014590.20   | 0.57%             |  |  |  | No            |                  1 | On Loan: No      |
| GREIF INC                               | Greif Inc                                     | CUSIP: 397624107<br>LEI: 54930070FVGKR0Z1FI24 | Long             | EC               | CORP              | US        |    180333 | NS      | $11777548.23  | 1.33%             |  |  |  | No            |                  1 | On Loan: No      |
| GRIFFON CORP                            | Griffon Corp                                  | CUSIP: 398433102<br>LEI: 5493000C2SGNC7ZN3242 | Long             | EC               | CORP              | US        |    156454 | NS      | $11915536.64  | 1.34%             |  |  |  | No            |                  1 | On Loan: No      |
| GULFPORT ENERGY CORP                    | Gulfport Energy Corp                          | CUSIP: 402635502<br>LEI: 254900DJRGZBUZ3F5038 | Long             | EC               | CORP              | US        |     62594 | NS      | $10893233.82  | 1.23%             |  |  |  | No            |                  1 | On Loan: No      |
| HNI CORP                                | HNI Corp                                      | CUSIP: 404251100<br>LEI: 549300SKRADLC4O5HV93 | Long             | EC               | CORP              | US        |    172523 | NS      | $7753183.62   | 0.87%             |  |  |  | No            |                  1 | On Loan: No      |
| HEALTHCARE SERV GRP INC.                | Healthcare Services Group Inc                 | CUSIP: 421906108<br>LEI: 529900GVZ1D2FBYMS719 | Long             | EC               | CORP              | US        |    258500 | NS      | $4035185.00   | 0.45%             |  |  |  | No            |                  1 | On Loan: No      |
| HELIOS TECHNOLOGIES INC                 | Helios Technologies Inc                       | CUSIP: 42328H109<br>LEI: 5493000BCJSMIDRNTI78 | Long             | EC               | CORP              | US        |    130750 | NS      | $7091880.00   | 0.80%             |  |  |  | No            |                  1 | On Loan: No      |
| HORACE MANN EDUCATORS CORP              | Horace Mann Educators Corp                    | CUSIP: 440327104<br>LEI: 254900G5YAV3A2YK8T32 | Long             | EC               | CORP              | US        |    182410 | NS      | $8387211.80   | 0.95%             |  |  |  | No            |                  1 | On Loan: No      |
| ICU MEDICAL INC.                        | ICU Medical Inc                               | CUSIP: 44930G107<br>LEI: 5493003NRW7LS2QZRH60 | Long             | EC               | CORP              | US        |     93720 | NS      | $11964295.20  | 1.35%             |  |  |  | No            |                  1 | On Loan: No      |
| KIRBY CORP                              | Kirby Corp                                    | CUSIP: 497266106<br>LEI: I7NO6FZO5GUCG585T325 | Long             | EC               | CORP              | US        |     94012 | NS      | $9137966.40   | 1.03%             |  |  |  | No            |                  1 | On Loan: No      |
| KONTOOR BRANDS INC                      | Kontoor Brands Inc                            | CUSIP: 50050N103<br>LEI: 549300M0CPXV65RFF271 | Long             | EC               | CORP              | US        |    102835 | NS      | $7944003.75   | 0.90%             |  |  |  | No            |                  1 | On Loan: No      |
| KORN FERRY                              | Korn Ferry                                    | CUSIP: 500643200<br>LEI: 549300FOD20OM0GT5G40 | Long             | EC               | CORP              | US        |    170800 | NS      | $12663112.00  | 1.43%             |  |  |  | No            |                  1 | On Loan: No      |
| LAUREATE EDUCATION INC                  | Laureate Education Inc                        | CUSIP: 518613203<br>LEI: 5JL7TMOH5TY3IUYSUP91 | Long             | EC               | CORP              | US        |    358622 | NS      | $9854932.56   | 1.11%             |  |  |  | No            |                  1 | On Loan: No      |
| LAZARD INC                              | Lazard Inc                                    | CUSIP: 52110M109<br>LEI: 254900RIBCDJSUFG1A11 | Long             | EC               | CORP              | US        |    115387 | NS      | $6595520.92   | 0.74%             |  |  |  | No            |                  1 | On Loan: No      |
| LEGALZOOM.COM INC                       | Legalzoom.com Inc                             | CUSIP: 52466B103<br>LEI: 5493000L8EBF74Y9JZ48 | Long             | EC               | CORP              | US        |    520980 | NS      | $5772458.40   | 0.65%             |  |  |  | No            |                  1 | On Loan: No      |
| LITTELFUSE INC                          | Littelfuse Inc                                | CUSIP: 537008104<br>LEI: 549300MY7HLQFYTPCX75 | Long             | EC               | CORP              | US        |     32840 | NS      | $8532817.20   | 0.96%             |  |  |  | No            |                  1 | On Loan: No      |
| MRC GLOBAL INC                          | MRC Global Inc                                | CUSIP: 55345K103<br>LEI: 529900TSBPOLCLI2L582 | Long             | EC               | CORP              | US        |    338880 | NS      | $5110310.40   | 0.58%             |  |  |  | No            |                  1 | On Loan: No      |
| MACERICH CO (THE)                       | Macerich Co/The                               | CUSIP: 554382101<br>LEI: 529900PSWCB5KI0JYU46 | Long             | EC               | CORP              | US        |    751772 | NS      | $13832604.80  | 1.56%             |  |  |  | No            |                  1 | On Loan: No      |
| MCGRATH                                 | McGrath RentCorp                              | CUSIP: 580589109<br>LEI: 254900EUH6IPWFQ83445 | Long             | EC               | CORP              | US        |     71480 | NS      | $8684105.20   | 0.98%             |  |  |  | No            |                  1 | On Loan: No      |
| PEDIATRIX MEDICAL GROUP INC             | Pediatrix Medical Group Inc                   | CUSIP: 58502B106<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    325710 | NS      | $5605469.10   | 0.63%             |  |  |  | No            |                  1 | On Loan: No      |
| MIMEDX GROUP INC                        | MiMedx Group Inc                              | CUSIP: 602496101<br>LEI: N/A                  | Long             | EC               | CORP              | US        |    473250 | NS      | $3360075.00   | 0.38%             |  |  |  | No            |                  1 | On Loan: No      |

### Signature

**Date Signed:** 2025-09-25

**Name of Applicant:** Columbia Funds Series Trust

**Signature:** Martha Skinner

**Name of Signer:** Martha Skinner

**Title:** Assistant Treasurer