# EDGAR Filing Document

**Accession Number:** 0001026710
**File Stem:** 0001398344-23-003123
**Filing Date:** 2023-2
**Character Count:** 21497
**Document Hash:** 39be1ae76b67d1c002bbf9e3639ddb93
**Contains OCR:** False
**Source Format:** 

## Filing Content

## Filing Summary
**0001398344-23-003123.hdr.sgml**: 20230214

**ACCESSION NUMBER**: 0001398344-23-003123

**CONFORMED SUBMISSION TYPE**: 13F-HR

**PUBLIC DOCUMENT COUNT**: 2

**CONFORMED PERIOD OF REPORT**: 20221231

**FILED AS OF DATE**: 20230214

**DATE AS OF CHANGE**: 20230214

**EFFECTIVENESS DATE**: 20230214

**FILER**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** Granahan Investment Management, LLC
- **CENTRAL INDEX KEY:** 0001026710
- **IRS NUMBER:** 042856200
- **STATE OF INCORPORATION:** MA
- **FISCAL YEAR END:** 1231

**FILING VALUES:**
- **FORM TYPE:** 13F-HR
- **SEC ACT:** 1934 Act
- **SEC FILE NUMBER:** 028-05876
- **FILM NUMBER:** 23623143

**BUSINESS ADDRESS:**
- **STREET 1:** 404 WYMAN STREET
- **STREET 2:** SUITE 460
- **CITY:** WALTHAM
- **STATE:** MA
- **ZIP:** 02451
- **BUSINESS PHONE:** 7818904412

**MAIL ADDRESS:**
- **STREET 1:** 404 WYMAN STREET
- **STREET 2:** SUITE 460
- **CITY:** WALTHAM
- **STATE:** MA
- **ZIP:** 02451

**FORMER COMPANY:**
- **FORMER CONFORMED NAME:** GRANAHAN INVESTMENT MANAGEMENT INC/MA
- **DATE OF NAME CHANGE:** 19990322

### FORM 13F COVER PAGE

**Report for the Calendar Year or Quarter Ended:** 12-31-2022

**Check here if Amendment** [ ] **Amendment Number:** 

**This Amendment (Check only one.):** [ ] is a restatement.

[ ] adds new holdings entries.

**Institutional Investment Manager Filing this Report:**

**Name:** Granahan Investment Management LLC<br>**Address:** 404 WYMAN STREET<br>SUITE 460<br>WALTHAM, MA 02451

**Form 13F File Number:** 028-05876

**CRD Number (if applicable):** 000107745

**SEC File Number (if applicable):** 801-23705

The institutional investment manager filing this report and the person by whom it is signed hereby represent that the person signing the report is authorized to submit it, that all information contained herein is true, correct and complete, and that it is understood that all required items, statements, schedules, lists, and tables, are considered integral parts of this form.

**Person Signing this Report on Behalf of Reporting Manager:**

**Name:** Susan Rawley<br>**Title:** VP/Operations and Account Administration<br>**Phone:** 781-902-1413

**Signature, Place, and Date of Signing:**

Susan Rawley  Waltham, MA  02-14-2023<br>[Signature]  [City, State]  [Date]

**Report Type (Check only one.):**

[x] **13F HOLDINGS REPORT.** (Check here if all holdings of this reporting manager are reported in this report.)

[ ] **13F NOTICE.** (Check here if no holdings are reported in this report, and all holdings are reported by other reporting manager(s).)

[ ] **13F COMBINATION REPORT.** (Check here if a portion of the holdings for this reporting manager are reported in this report and a portion are reported by other reporting manager(s).)

### Form 13F Summary Page

**Report Summary:**

**Number of Other Included Managers:** 1

**Form 13F Information Table Entry Total:** 170

**Form 13F Information Table Value Total:** $3270480003

      (round to nearest dollar)

**List of Other Included Managers:**

Provide a numbered list of the name(s) and Form 13F file number(s) of all institutional investment managers with respect to which this report is filed, other than the manager filing this report.

[If there are no entries in this list, state “NONE” and omit the column headings and list entries.]

| No.   | Name                        | Form 13F File Number   |
|:---|:---|:---|
|  | Destra Capital Advisors LLC | 028-15699              |

### FORM 13F INFORMATION TABLE

---

|  |  |  |  |  |  |  |  |  |  |  |  |  |
| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
| NAME OF ISSUER | TITLE OF CLASS | CUSIP | FIGI | VALUE (x$1000) | SHRS OR PRN AMT | SH/PRN | PUT/CALL | INVESTMENT DISCRETION | OTHER MANAGER | VOTING AUTHORITY | VOTING AUTHORITY | VOTING AUTHORITY |
| NAME OF ISSUER | TITLE OF CLASS | CUSIP | FIGI | VALUE (x$1000) | SHRS OR PRN AMT | SH/PRN | PUT/CALL | INVESTMENT DISCRETION | OTHER MANAGER | SOLE | SHARED | NONE |
| 908 DEVICES INC | COM | 65443P102 |  | 8419368 | 1104904 | SH |  | SOLE |  | 903073 | 0 | 201831 |
| AFFIMED N V | COM | N01045108 |  | 4761040 | 3839548 | SH |  | SOLE |  | 3560510 | 0 | 279038 |
| AIRGAIN INC | COM | 00938A104 |  | 4493684 | 690274 | SH |  | SOLE |  | 525450 | 0 | 164824 |
| AKOUSTIS TECHNOLOGIES INC | COM | 00973N102 |  | 2648406 | 939151 | SH |  | SOLE |  | 701251 | 0 | 237900 |
| AKOYA BIOSCIENCES INC | COM | 00974H104 |  | 5161484 | 539340 | SH |  | SOLE |  | 445621 | 0 | 93719 |
| ALTA EQUIPMENT GROUP INC | COMMON STOCK | 02128L106 |  | 13297248 | 1008131 | SH |  | SOLE |  | 863486 | 0 | 144645 |
| AMBARELLA INC | SHS | G037AX101 |  | 16825327 | 204613 | SH |  | SOLE |  | 185776 | 0 | 18837 |
| AMERESCO INC | CL A | 02361E108 |  | 16986465 | 297278 | SH |  | SOLE |  | 280198 | 0 | 17080 |
| AMERICAN WELL CORP | CL A | 03044L105 |  | 5361390 | 1894484 | SH |  | SOLE |  | 1561737 | 0 | 332747 |
| APELLIS PHARMACEUTICALS INC | COM | 03753U106 |  | 9437489 | 182508 | SH |  | SOLE |  | 141463 | 0 | 41045 |
| APPIAN CORP | CL A | 03782L101 |  | 3335739 | 102449 | SH |  | SOLE |  | 89028 | 0 | 13421 |
| ARCUS BIOSCIENCES INC | COM | 03969F109 |  | 8488726 | 410480 | SH |  | SOLE |  | 327406 | 0 | 83074 |
| ARRAY TECHNOLOGIES INC | COM SHS | 04271T100 |  | 68618929 | 3549867 | SH |  | SOLE |  | 3302729 | 0 | 247138 |
| ARTERIS INC | COM | 04302A104 |  | 1814750 | 422035 | SH |  | SOLE |  | 313718 | 0 | 108317 |
| AXON ENTERPRISE INC | COM | 05464C101 |  | 118101673 | 711756 | SH |  | SOLE |  | 574076 | 0 | 137680 |
| AXT INC | COM | 00246W103 |  | 6278485 | 1433444 | SH |  | SOLE |  | 1160014 | 0 | 273430 |
| AZENTA INC | COM | 114340102 |  | 89502770 | 1537320 | SH |  | SOLE |  | 1237476 | 0 | 299844 |
| BEAM THERAPEUTICS INC | COM | 07373V105 |  | 6405827 | 163790 | SH |  | SOLE |  | 131886 | 0 | 31904 |
| BIOLIFE SOLUTIONS INC | COM NEW | 09062W204 |  | 15393214 | 845781 | SH |  | SOLE |  | 704923 | 0 | 140858 |
| BIOMERICA INC | COM NEW | 09061H307 |  | 3761048 | 1122701 | SH |  | SOLE |  | 1000130 | 0 | 122571 |
| BLUEBIRD BIO INC | COM | 09609G100 |  | 4909941 | 709529 | SH |  | SOLE |  | 600826 | 0 | 108703 |
| BLUEPRINT MEDICINES CORP | COM | 09627Y109 |  | 9849101 | 224814 | SH |  | SOLE |  | 173981 | 0 | 50833 |
| BUMBLE INC | COM CL A | 12047B105 |  | 18193326 | 864291 | SH |  | SOLE |  | 706829 | 0 | 157462 |
| CARIBOU BIOSCIENCES INC | COM | 142038108 |  | 1446541 | 230341 | SH |  | SOLE |  | 208041 | 0 | 22300 |
| CASELLA WASTE SYS INC | CL A | 147448104 |  | 20340874 | 256473 | SH |  | SOLE |  | 201568 | 0 | 54905 |
| CASTLE BIOSCIENCES INC | COM | 14843C105 |  | 24583481 | 1044328 | SH |  | SOLE |  | 896227 | 0 | 148101 |
| CEVA INC | COM | 157210105 |  | 12630381 | 493760 | SH |  | SOLE |  | 401715 | 0 | 92045 |
| CHART INDS INC | COM | 16115Q308 |  | 26311042 | 228335 | SH |  | SOLE |  | 215071 | 0 | 13264 |
| CHEGG INC | COM | 163092109 |  | 16118116 | 637836 | SH |  | SOLE |  | 514925 | 0 | 122911 |
| CHICKEN SOUP FOR THE SOUL EN | CL A | 16842Q100 |  | 7061396 | 1379179 | SH |  | SOLE |  | 1289680 | 0 | 89499 |
| CIENA CORP | COM NEW | 171779309 |  | 10890042 | 213614 | SH |  | SOLE |  | 164047 | 0 | 49567 |
| CLEAN ENERGY FUELS CORP | COM | 184499101 |  | 8532321 | 1640831 | SH |  | SOLE |  | 1502594 | 0 | 138237 |
| COLUMBUS MCKINNON CORP N Y | COM | 199333105 |  | 18039942 | 555588 | SH |  | SOLE |  | 467024 | 0 | 88564 |
| COMPUTER TASK GROUP INC | COM | 205477102 |  | 578899 | 76574 | SH |  | SOLE |  | 76574 | 0 | 0 |
| CONFLUENT INC | CLASS A COM | 20717M103 |  | 2383594 | 107176 | SH |  | SOLE |  | 79840 | 0 | 27336 |
| CORVEL CORP | COM | 221006109 |  | 6172165 | 42470 | SH |  | SOLE |  | 32832 | 0 | 9638 |
| COSTAR GROUP INC | COM | 22160N109 |  | 105631095 | 1366862 | SH |  | SOLE |  | 1114835 | 0 | 252027 |
| CRYOPORT INC | COM PAR $0.001 | 229050307 |  | 10103651 | 582343 | SH |  | SOLE |  | 525080 | 0 | 57263 |
| CYBERARK SOFTWARE LTD | SHS | M2682V108 |  | 4285321 | 33053 | SH |  | SOLE |  | 24690 | 0 | 8363 |
| CYTOSORBENTS CORP | COM NEW | 23283X206 |  | 2470252 | 1593711 | SH |  | SOLE |  | 1358955 | 0 | 234756 |
| DARLING INGREDIENTS INC | COM | 237266101 |  | 3046193 | 48669 | SH |  | SOLE |  | 48669 | 0 | 0 |
| DEFINITIVE HEALTHCARE CORP | CLASS A COM | 24477E103 |  | 11793314 | 1073095 | SH |  | SOLE |  | 904063 | 0 | 169032 |
| DIGITAL TURBINE INC | COM NEW | 25400W102 |  | 52432671 | 3440464 | SH |  | SOLE |  | 3180383 | 0 | 260081 |
| DOMO INC | COM CL B | 257554105 |  | 10464264 | 734850 | SH |  | SOLE |  | 605748 | 0 | 129102 |
| DOUGLAS DYNAMICS INC | COM | 25960R105 |  | 707109 | 19555 | SH |  | SOLE |  | 19555 | 0 | 0 |
| EDGIO INC | COM | 53261M104 |  | 11605335 | 10270208 | SH |  | SOLE |  | 8734990 | 0 | 1535218 |
| ENERGY RECOVERY INC | COM | 29270J100 |  | 464898 | 22689 | SH |  | SOLE |  | 22689 | 0 | 0 |
| ENOVIX CORPORATION | COM | 293594107 |  | 23745696 | 1908818 | SH |  | SOLE |  | 1555110 | 0 | 353708 |
| ENPHASE ENERGY INC | COM | 29355A107 |  | 93742848 | 353800 | SH |  | SOLE |  | 296577 | 0 | 57223 |
| EPAM SYS INC | COM | 29414B104 |  | 6861892 | 20937 | SH |  | SOLE |  | 20937 | 0 | 0 |
| ETSY INC | COM | 29786A106 |  | 96775293 | 807942 | SH |  | SOLE |  | 668012 | 0 | 139930 |
| EURONET WORLDWIDE INC | COM | 298736109 |  | 49319685 | 522565 | SH |  | SOLE |  | 472119 | 0 | 50446 |
| EVERQUOTE INC | COM CL A | 30041R108 |  | 24471363 | 1660201 | SH |  | SOLE |  | 1376828 | 0 | 283373 |
| EVOLENT HEALTH INC | CL A | 30050B101 |  | 89208503 | 3176941 | SH |  | SOLE |  | 2761318 | 0 | 415623 |
| EXPEDIA GROUP INC | COM NEW | 30212P303 |  | 2369580 | 27050 | SH |  | SOLE |  | 27050 | 0 | 0 |
| FATE THERAPEUTICS INC | COM | 31189P102 |  | 6075724 | 602153 | SH |  | SOLE |  | 495558 | 0 | 106595 |
| FIGS INC | CL A | 30260D103 |  | 14065242 | 2089932 | SH |  | SOLE |  | 1946975 | 0 | 142957 |
| FIRST SOLAR INC | COM | 336433107 |  | 20916676 | 139640 | SH |  | SOLE |  | 126669 | 0 | 12971 |
| FIRST WATCH RESTAURANT GROUP | COM | 33748L101 |  | 12568924 | 928967 | SH |  | SOLE |  | 756283 | 0 | 172684 |
| FLEX LTD | ORD | Y2573F102 |  | 3504439 | 163301 | SH |  | SOLE |  | 163301 | 0 | 0 |
| GENIUS SPORTS LIMITED | SHARES CL A | G3934V109 |  | 33397960 | 9355171 | SH |  | SOLE |  | 7750060 | 0 | 1605111 |
| GENTHERM INC | COM | 37253A103 |  | 11924696 | 182642 | SH |  | SOLE |  | 171571 | 0 | 11071 |
| GLOBANT S A | COM | L44385109 |  | 26636544 | 158400 | SH |  | SOLE |  | 129844 | 0 | 28556 |
| GREEN BRICK PARTNERS INC | COM | 392709101 |  | 2696217 | 111276 | SH |  | SOLE |  | 111276 | 0 | 0 |
| GRID DYNAMICS HLDGS INC | CL A | 39813G109 |  | 17889134 | 1594397 | SH |  | SOLE |  | 1488464 | 0 | 105933 |
| GROCERY OUTLET HLDG CORP | COM | 39874R101 |  | 5555353 | 190317 | SH |  | SOLE |  | 147858 | 0 | 42459 |
| GUIDEWIRE SOFTWARE INC | COM | 40171V100 |  | 13998989 | 223769 | SH |  | SOLE |  | 172838 | 0 | 50931 |
| HAGERTY INC | CL A COM | 405166109 |  | 13400561 | 1593408 | SH |  | SOLE |  | 1489905 | 0 | 103503 |
| HARMONIC INC | COM | 413160102 |  | 23220588 | 1772564 | SH |  | SOLE |  | 1649195 | 0 | 123369 |
| HARVARD BIOSCIENCE INC | COM | 416906105 |  | 4051006 | 1462457 | SH |  | SOLE |  | 1385079 | 0 | 77378 |
| HEALTH CATALYST INC | COM | 42225T107 |  | 8604826 | 809485 | SH |  | SOLE |  | 669596 | 0 | 139889 |
| HEALTHEQUITY INC | COM | 42226A107 |  | 37923270 | 615238 | SH |  | SOLE |  | 500824 | 0 | 114414 |
| HESKA CORP | COM RESTRC NEW | 42805E306 |  | 5004439 | 80509 | SH |  | SOLE |  | 62604 | 0 | 17905 |
| HUBSPOT INC | COM | 443573100 |  | 61610712 | 213090 | SH |  | SOLE |  | 173198 | 0 | 39892 |
| HUDSON TECHNOLOGIES INC | COM | 444144109 |  | 10092696 | 997302 | SH |  | SOLE |  | 889028 | 0 | 108274 |
| ICAD INC | COM NEW | 44934S206 |  | 5213897 | 2849124 | SH |  | SOLE |  | 2637653 | 0 | 211471 |
| IMMUNOGEN INC | COM | 45253H101 |  | 10842352 | 2185958 | SH |  | SOLE |  | 1752300 | 0 | 433658 |
| INDIE SEMICONDUCTOR INC | CLASS A COM | 45569U101 |  | 42503318 | 7290449 | SH |  | SOLE |  | 6849675 | 0 | 440774 |
| INMODE LTD | SHS | M5425M103 |  | 4320914 | 121034 | SH |  | SOLE |  | 113547 | 0 | 7487 |
| INSMED INC | COM PAR $.01 | 457669307 |  | 4194941 | 209957 | SH |  | SOLE |  | 162093 | 0 | 47864 |
| INTELLIA THERAPEUTICS INC | COM | 45826J105 |  | 7592866 | 217623 | SH |  | SOLE |  | 173426 | 0 | 44197 |
| ISHARES TR | RUS 2000 GRW ETF | 464287648 |  | 603230 | 2812 | SH |  | SOLE |  | 2812 | 0 | 0 |
| JAMES RIV GROUP LTD | COM | G5005R107 |  | 19499014 | 932521 | SH |  | SOLE |  | 736855 | 0 | 195666 |
| KEZAR LIFE SCIENCES INC | COM | 49372L100 |  | 4860282 | 690381 | SH |  | SOLE |  | 561781 | 0 | 128600 |
| KINIKSA PHARMACEUTICALS LTD | COM CL A | G5269C101 |  | 7552811 | 504193 | SH |  | SOLE |  | 416087 | 0 | 88106 |
| KINSALE CAP GROUP INC | COM | 49714P108 |  | 6183640 | 23645 | SH |  | SOLE |  | 18154 | 0 | 5491 |
| KORNIT DIGITAL LTD | SHS | M6372Q113 |  | 81106611 | 3530980 | SH |  | SOLE |  | 3099201 | 0 | 431779 |
| LENDINGTREE INC NEW | COM | 52603B107 |  | 8722925 | 408951 | SH |  | SOLE |  | 379548 | 0 | 29403 |
| LIFEMD INC | COM | 53216B104 |  | 414520 | 213670 | SH |  | SOLE |  | 213670 | 0 | 0 |
| LIFE TIME GROUP HOLDINGS INC | COMMON STOCK | 53190C102 |  | 9947957 | 831769 | SH |  | SOLE |  | 664736 | 0 | 167033 |
| LIQUIDITY SVCS INC | COM | 53635B107 |  | 30310689 | 2155810 | SH |  | SOLE |  | 1762870 | 0 | 392940 |
| LITHIA MTRS INC | COM | 536797103 |  | 18419229 | 89964 | SH |  | SOLE |  | 85259 | 0 | 4705 |
| LITTELFUSE INC | COM | 537008104 |  | 3371702 | 15312 | SH |  | SOLE |  | 15312 | 0 | 0 |
| LIVEPERSON INC | COM | 538146101 |  | 12211338 | 1204274 | SH |  | SOLE |  | 972337 | 0 | 231937 |
| LOVESAC COMPANY | COM | 54738L109 |  | 28923341 | 1314100 | SH |  | SOLE |  | 1100387 | 0 | 213713 |
| LUXFER HLDGS PLC | SHS | G5698W116 |  | 1662850 | 121199 | SH |  | SOLE |  | 121199 | 0 | 0 |
| MAGNITE INC | COM | 55955D100 |  | 110707330 | 10453950 | SH |  | SOLE |  | 9356941 | 0 | 1097009 |
| MATCH GROUP INC NEW | COM | 57667L107 |  | 2671043 | 64378 | SH |  | SOLE |  | 64378 | 0 | 0 |
| MATERION CORP | COM | 576690101 |  | 20970196 | 239632 | SH |  | SOLE |  | 195788 | 0 | 43844 |
| MAXCYTE INC | COM | 57777K106 |  | 3692543 | 676290 | SH |  | SOLE |  | 553896 | 0 | 122394 |
| MESA LABS INC | COM | 59064R109 |  | 5633522 | 33894 | SH |  | SOLE |  | 26191 | 0 | 7703 |
| MIRATI THERAPEUTICS INC | COM | 60468T105 |  | 4163989 | 91900 | SH |  | SOLE |  | 71245 | 0 | 20655 |
| MONOLITHIC PWR SYS INC | COM | 609839105 |  | 4071819 | 11515 | SH |  | SOLE |  | 8628 | 0 | 2887 |
| NATERA INC | COM | 632307104 |  | 15484209 | 385467 | SH |  | SOLE |  | 339584 | 0 | 45883 |
| NUTANIX INC | CL A | 67059N108 |  | 9882979 | 379385 | SH |  | SOLE |  | 291983 | 0 | 87402 |
| OKTA INC | CL A | 679295105 |  | 857883 | 12555 | SH |  | SOLE |  | 9388 | 0 | 3167 |
| OLINK HLDG AB | SPONSORED ADS | 680710100 |  | 18090915 | 712802 | SH |  | SOLE |  | 586843 | 0 | 125959 |
| ONESPAWORLD HOLDINGS LIMITED | COM | P73684113 |  | 14116784 | 1513053 | SH |  | SOLE |  | 1254284 | 0 | 258769 |
| ONTO INNOVATION INC | COM | 683344105 |  | 7495892 | 110088 | SH |  | SOLE |  | 81975 | 0 | 28113 |
| ORTHOPEDIATRICS CORP | COM | 68752L100 |  | 20160273 | 507432 | SH |  | SOLE |  | 421201 | 0 | 86231 |
| OSI SYSTEMS INC | COM | 671044105 |  | 7747316 | 97426 | SH |  | SOLE |  | 74550 | 0 | 22876 |
| PACIFIC BIOSCIENCES CALIF IN | COM | 69404D108 |  | 5539152 | 677158 | SH |  | SOLE |  | 524762 | 0 | 152396 |
| PAR TECHNOLOGY CORP | COM | 698884103 |  | 7966679 | 305588 | SH |  | SOLE |  | 282190 | 0 | 23398 |
| PAYCOM SOFTWARE INC | COM | 70432V102 |  | 108730142 | 350392 | SH |  | SOLE |  | 287061 | 0 | 63331 |
| PAYLOCITY HLDG CORP | COM | 70438V106 |  | 39073456 | 201140 | SH |  | SOLE |  | 166313 | 0 | 34827 |
| PERFICIENT INC | COM | 71375U101 |  | 14678196 | 210199 | SH |  | SOLE |  | 162097 | 0 | 48102 |
| PHREESIA INC | COM | 71944F106 |  | 23116107 | 714342 | SH |  | SOLE |  | 589385 | 0 | 124957 |
| PLIANT THERAPEUTICS INC | COM | 729139105 |  | 9765207 | 505184 | SH |  | SOLE |  | 407956 | 0 | 97228 |
| PORCH GROUP INC | COM | 733245104 |  | 25152481 | 13378979 | SH |  | SOLE |  | 11691413 | 0 | 1687566 |
| PORTILLOS INC | COM CL A | 73642K106 |  | 8545087 | 523596 | SH |  | SOLE |  | 443670 | 0 | 79926 |
| POWER INTEGRATIONS INC | COM | 739276103 |  | 17922183 | 249891 | SH |  | SOLE |  | 194582 | 0 | 55309 |
| PROS HOLDINGS INC | COM | 74346Y103 |  | 19321537 | 796436 | SH |  | SOLE |  | 644757 | 0 | 151679 |
| PROTHENA CORP PLC | SHS | G72800108 |  | 12045542 | 199926 | SH |  | SOLE |  | 162468 | 0 | 37458 |
| PTC INC | COM | 69370C100 |  | 9288935 | 77382 | SH |  | SOLE |  | 57730 | 0 | 19652 |
| PURE STORAGE INC | CL A | 74624M102 |  | 17787051 | 664688 | SH |  | SOLE |  | 547077 | 0 | 117611 |
| QUALYS INC | COM | 74758T303 |  | 6962637 | 62039 | SH |  | SOLE |  | 46187 | 0 | 15852 |
| QUANTERIX CORP | COM | 74766Q101 |  | 17472495 | 1261552 | SH |  | SOLE |  | 1073554 | 0 | 187998 |
| RADWARE LTD | ORD | M81873107 |  | 10497935 | 531541 | SH |  | SOLE |  | 432417 | 0 | 99124 |
| RAPID7 INC | COM | 753422104 |  | 10354114 | 304712 | SH |  | SOLE |  | 235469 | 0 | 69243 |
| RBC BEARINGS INC | COM | 75524B104 |  | 16111157 | 76958 | SH |  | SOLE |  | 60170 | 0 | 16788 |
| REPLIGEN CORP | COM | 759916109 |  | 18228930 | 107666 | SH |  | SOLE |  | 86554 | 0 | 21112 |
| RESEARCH SOLUTIONS INC | COM | 761025105 |  | 41067 | 21389 | SH |  | SOLE |  | 21389 | 0 | 0 |
| REVANCE THERAPEUTICS INC | COM | 761330109 |  | 6425261 | 348064 | SH |  | SOLE |  | 279538 | 0 | 68526 |
| RITCHIE BROS AUCTIONEERS | COM | 767744105 |  | 30686564 | 530634 | SH |  | SOLE |  | 431997 | 0 | 98637 |
| RUSH ENTERPRISES INC | CL A | 781846209 |  | 2574215 | 49239 | SH |  | SOLE |  | 49239 | 0 | 0 |
| SHOCKWAVE MED INC | COM | 82489T104 |  | 5138811 | 24993 | SH |  | SOLE |  | 19259 | 0 | 5734 |
| SHOTSPOTTER INC | COM | 82536T107 |  | 18892600 | 558457 | SH |  | SOLE |  | 460330 | 0 | 98127 |
| SI-BONE INC | COM | 825704109 |  | 15809782 | 1162484 | SH |  | SOLE |  | 979872 | 0 | 182612 |
| SILICON LABORATORIES INC | COM | 826919102 |  | 5612668 | 41370 | SH |  | SOLE |  | 30847 | 0 | 10523 |
| SILK RD MED INC | COM | 82710M100 |  | 2176574 | 41184 | SH |  | SOLE |  | 41184 | 0 | 0 |
| SILVERGATE CAP CORP | CL A | 82837P408 |  | 8563636 | 492163 | SH |  | SOLE |  | 420762 | 0 | 71401 |
| SITIME CORP | COM | 82982T106 |  | 2882959 | 28370 | SH |  | SOLE |  | 21126 | 0 | 7244 |
| SOLAREDGE TECHNOLOGIES INC | COM | 83417M104 |  | 4374539 | 15443 | SH |  | SOLE |  | 15443 | 0 | 0 |
| SPROUT SOCIAL INC | COM CL A | 85209W109 |  | 35795471 | 633997 | SH |  | SOLE |  | 512434 | 0 | 121563 |
| SPS COMM INC | COM | 78463M107 |  | 105835567 | 824072 | SH |  | SOLE |  | 670807 | 0 | 153265 |
| STOKE THERAPEUTICS INC | COM | 86150R107 |  | 5078521 | 550219 | SH |  | SOLE |  | 453719 | 0 | 96500 |
| STONERIDGE INC | COM | 86183P102 |  | 10445497 | 484485 | SH |  | SOLE |  | 448405 | 0 | 36080 |
| STRATASYS LTD | SHS | M85548101 |  | 20976414 | 1768669 | SH |  | SOLE |  | 1416052 | 0 | 352617 |
| TECHTARGET INC | COM | 87874R100 |  | 6823704 | 154873 | SH |  | SOLE |  | 115560 | 0 | 39313 |
| TEXAS CAP BANCSHARES INC | COM | 88224Q107 |  | 31436708 | 521252 | SH |  | SOLE |  | 408839 | 0 | 112413 |
| THE BEAUTY HEALTH COMPANY | COM CL A | 88331L108 |  | 10598943 | 1164719 | SH |  | SOLE |  | 959829 | 0 | 204890 |
| THREDUP INC | CL A | 88556E102 |  | 2777247 | 2120036 | SH |  | SOLE |  | 1647728 | 0 | 472308 |
| TOAST INC | CL A | 888787108 |  | 92572294 | 5134348 | SH |  | SOLE |  | 4186984 | 0 | 947364 |
| TOPGOLF CALLAWAY BRANDS CORP | COM | 131193104 |  | 16911688 | 856288 | SH |  | SOLE |  | 674428 | 0 | 181860 |
| VARONIS SYS INC | COM | 922280102 |  | 13989339 | 584350 | SH |  | SOLE |  | 518347 | 0 | 66003 |
| VEECO INSTRS INC DEL | COM | 922417100 |  | 19512846 | 1050207 | SH |  | SOLE |  | 845878 | 0 | 204329 |
| VERACYTE INC | COM | 92337F107 |  | 41842302 | 1763266 | SH |  | SOLE |  | 1617645 | 0 | 145621 |
| VERICEL CORP | COM | 92346J108 |  | 8964582 | 340341 | SH |  | SOLE |  | 282050 | 0 | 58291 |
| VERITONE INC | COM | 92347M100 |  | 1576342 | 297423 | SH |  | SOLE |  | 222161 | 0 | 75262 |
| VICOR CORP | COM | 925815102 |  | 8258365 | 153644 | SH |  | SOLE |  | 129282 | 0 | 24362 |
| VICTORY CAP HLDGS INC | COM CL A | 92645B103 |  | 29551421 | 1101432 | SH |  | SOLE |  | 910893 | 0 | 190539 |
| VITA COCO CO INC | COM | 92846Q107 |  | 33730861 | 2440728 | SH |  | SOLE |  | 2278628 | 0 | 162100 |
| VIZIO HLDG CORP | CL A COM | 92858V101 |  | 2852546 | 384959 | SH |  | SOLE |  | 342764 | 0 | 42195 |
| WILLSCOT MOBIL MINI HLDNG CO | COM CL A | 971378104 |  | 43056044 | 953200 | SH |  | SOLE |  | 775004 | 0 | 178196 |
| WNS HLDGS LTD | SPON ADR | 92932M101 |  | 12119845 | 151517 | SH |  | SOLE |  | 115937 | 0 | 35580 |
| WORKIVA INC | COM CL A | 98139A105 |  | 47695632 | 568008 | SH |  | SOLE |  | 462202 | 0 | 105806 |
| XENCOR INC | COM | 98401F105 |  | 5002805 | 192120 | SH |  | SOLE |  | 158516 | 0 | 33604 |
| XENON PHARMACEUTICALS INC | COM | 98420N105 |  | 12785178 | 324250 | SH |  | SOLE |  | 262923 | 0 | 61327 |
| ZETA GLOBAL HOLDINGS CORP | CL A | 98956A105 |  | 58771638 | 7193591 | SH |  | SOLE |  | 5954569 | 0 | 1239022 |
| ZYMEWORKS DEL INC | COM | 98985Y108 |  | 5804201 | 738448 | SH |  | SOLE |  | 622885 | 0 | 115563 |

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