# EDGAR Filing Document

**Accession Number:** 0001656634
**File Stem:** 0001193125-23-030127
**Filing Date:** 2023-2
**Character Count:** 34353
**Document Hash:** d7257e18f43a542d791c6da345b0bb03
**Contains OCR:** False
**Source Format:** 

## Filing Content

## Filing Summary
**0001193125-23-030127.hdr.sgml**: 20230209

**ACCESSION NUMBER**: 0001193125-23-030127

**CONFORMED SUBMISSION TYPE**: SC 13G/A

**PUBLIC DOCUMENT COUNT**: 1

**FILED AS OF DATE**: 20230209

**DATE AS OF CHANGE**: 20230209

**GROUP MEMBERS**: VERSANT AFFILIATES FUND V, L.P.

**GROUP MEMBERS**: VERSANT OPHTHALMIC AFFILIATES FUND I, L.P.

**GROUP MEMBERS**: VERSANT VANTAGE I GP, L.P.

**GROUP MEMBERS**: VERSANT VANTAGE I GP-GP, LLC

**GROUP MEMBERS**: VERSANT VANTAGE I, L.P.

**GROUP MEMBERS**: VERSANT VENTURE CAPITAL V (CANADA) LP

**GROUP MEMBERS**: VERSANT VENTURES V (CANADA), L.P.

**GROUP MEMBERS**: VERSANT VENTURES V GP-GP (CANADA), INC.

**GROUP MEMBERS**: VERSANT VENTURES V, LLC

**SUBJECT COMPANY**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** Gritstone bio, Inc.
- **CENTRAL INDEX KEY:** 0001656634
- **STANDARD INDUSTRIAL CLASSIFICATION:** BIOLOGICAL PRODUCTS (NO DIAGNOSTIC SUBSTANCES) [2836]
- **IRS NUMBER:** 000000000
- **STATE OF INCORPORATION:** DE
- **FISCAL YEAR END:** 1231

**FILING VALUES:**
- **FORM TYPE:** SC 13G/A
- **SEC ACT:** 1934 Act
- **SEC FILE NUMBER:** 005-90655
- **FILM NUMBER:** 23606698

**BUSINESS ADDRESS:**
- **STREET 1:** 5959 HORTON STREET, SUITE 300
- **CITY:** EMERYVILLE
- **STATE:** CA
- **ZIP:** 94608
- **BUSINESS PHONE:** (510) 871-6100

**MAIL ADDRESS:**
- **STREET 1:** 5959 HORTON STREET, SUITE 300
- **CITY:** EMERYVILLE
- **STATE:** CA
- **ZIP:** 94608

**FORMER COMPANY:**
- **FORMER CONFORMED NAME:** Gritstone Oncology, Inc.
- **DATE OF NAME CHANGE:** 20151023
**FILED BY**: 

**COMPANY DATA:**
- **COMPANY CONFORMED NAME:** Versant Venture Capital V, L.P.
- **CENTRAL INDEX KEY:** 0001589097
- **IRS NUMBER:** 000000000
- **STATE OF INCORPORATION:** DE
- **FISCAL YEAR END:** 1231

**FILING VALUES:**
- **FORM TYPE:** SC 13G/A

**BUSINESS ADDRESS:**
- **STREET 1:** ONE SANSOME STREET
- **STREET 2:** SUITE 3630
- **CITY:** SAN FRANCISCO
- **STATE:** CA
- **ZIP:** 94104
- **BUSINESS PHONE:** 650-233-7877

**MAIL ADDRESS:**
- **STREET 1:** ONE SANSOME STREET
- **STREET 2:** SUITE 3630
- **CITY:** SAN FRANCISCO
- **STATE:** CA
- **ZIP:** 94104

------

------

**UNITED STATES** 

**SECURITIES AND EXCHANGE COMMISSION** 

**Washington, D.C. 20549** 

------

**SCHEDULE 13G** 

**(Rule 13d-102)**

**INFORMATION TO BE INCLUDED IN STATEMENTS FILED PURSUANT TO RULES 13d-1 (b), (c) AND (d) AND AMENDMENTS THERETO FILED PURSUANT TO 13d-2 (b)** 

**(Amendment No. 3)** 

------

## Gritstone Oncology, Inc.
**(Name of Issuer)** 

**Common Stock, par value $0.0001 per share** 

**(Title of Class of Securities)** 

**39868T105** 

**(CUSIP Number)** 

**December 31, 2022** 

**(Date of Event Which Requires Filing of this Statement)** 

------

Check the appropriate box to designate the rule pursuant to which this Schedule is filed:

☐ Rule 13d-1(b)

☐ Rule 13d-1(c)

☒ Rule 13d-1(d)

The information required on the remainder of this cover page shall not be deemed to be "filed" for the purpose of Section 18 of the Securities Exchange Act of 1934 ("Act") or otherwise subject to the liabilities of that section of the Act but shall be subject to all other provisions of the Act (however, see the Notes).

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Venture Capital V, L.P.

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;2,723,031 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;2,723,031 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 2,723,031 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 3.3% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; PN

------

(1) This Schedule 13G is filed by Versant Venture Capital V, L.P. ("VVC V"), Versant Affiliates Fund V,
L.P. ("VAF V"), Versant Ophthalmic Affiliates Fund I, L.P. ("VOA"), Versant Venture Capital V (Canada) LP ("VVC CAN") and Versant Vantage I, L.P. ("VV I"). Versant Ventures V, LLC ("VV V") is the
sole general partner of VVC V, VAF V and VOA. Samuel D. Colella ("Colella"), Jerel C. Davis ("Davis"), William J. Link ("Link"), Bradley Bolzon, Ph.D. ("Bolzon"), Kirk G. Nielsen ("Nielsen"), Thomas
Woiwode ("Woiwode") and Robin L. Praeger ("Praeger") are managing directors of VV V. Versant Ventures V GP-GP (Canada), Inc. ("VV V CAN GP") is the sole general partner of Versant
Ventures V (Canada), L.P. ("VV V CAN") and VV V CAN is the sole general partner of VVC CAN. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are directors of VV V CAN GP. Versant Vantage I GP-GP, LLC ("VV I GP-GP") is the sole general partner of Versant Vantage I GP, L.P. ("VV I GP", and, together with VVC V, VV V, VAF V, VOA, VVC CAN,
VV V CAN, VV V CAN GP, VV I and VV I GP-GP, the "Reporting Persons"). VV I GP is the sole general partner of VV I. Bolzon, Davis, Clare Ozawa ("Ozawa"), Woiwode and Praeger are managing
directors of VV I GP-GP. The Reporting Persons expressly disclaim status as a "group" for purposes of this Schedule 13G.

(2) These shares are held by VVC V. VV V is the sole general partner of VVC V and may be deemed to have voting and
dispositive power over the securities held by VVC V. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are managing directors of VV V and share voting and dispositive power over the shares held by VVC V.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's quarterly report on Form 10-Q for the quarter ended September 30, 2022, filed with the United States Securities and Exchange Commission on
November 3, 2022 (the "Form 10-Q").

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Affiliates Fund V, L.P.

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;81,908 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;81,908 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 81,908 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.1% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; PN

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VAF V. VV V is the sole general partner of VAF V and may be deemed to have voting and
dispositive power over the securities held by VAF V. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are managing directors of VV V and share voting and dispositive power over the shares held by VAF V.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Ophthalmic Affiliates Fund I, L.P.

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;90,756 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;90,756 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 90,756 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.1% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; PN

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VOA. VV V is the sole general partner of VOA and may be deemed to have voting and
dispositive power over the securities held by VOA. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are managing directors of VV V and share voting and dispositive power over the shares held by VOA.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Ventures V, LLC

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;2,895,695 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;2,895,695 shares of Common Stock (2) |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 2,895,695 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 3.5% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; OO

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) Includes (i) 2,723,031 shares held by VVC V, (ii) 81,908 shares held by VAF V and (iii) 90,756 shares held by
VOA. VV V is the sole general partner of each of VVC V, VAF V and VOA and may be deemed to have voting and dispositive power over the securities held by each of VVC V, VAF V and VOA. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are
managing directors of VV V and share voting and dispositive power over the shares held by each of VVC V, VAF V and VOA.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Venture Capital V (Canada) LP

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Ontario,
Canada

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;207,234 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;207,234 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 207,234 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.2% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; PN

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VVC CAN. VV V CAN GP is the sole general partner of VV V CAN and VV V CAN is the sole
general partner of VVC CAN. Each of VV CAN GP and VV V CAN share voting and dispositive power over the shares held by VVC CAN. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are directors of VV V CAN GP and share voting and dispositive
power over the shares held by VVC CAN.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Ventures V GP-GP (Canada), Inc.

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;207,234 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;207,234 shares of Common Stock (2) |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 207,234 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.2% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; CO

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VVC CAN. VV V CAN GP is the sole general partner of VV V CAN and VV V CAN is the sole
general partner of VVC CAN. Each of VV CAN GP and VV V CAN share voting and dispositive power over the shares held by VVC CAN. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are directors of VV V CAN GP and share voting and dispositive
power over the shares held by VVC CAN.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Ventures V (Canada), L.P.

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;207,234 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;207,234 shares of Common Stock (2) |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 207,234 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.2% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; PN

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VVC CAN. VV V CAN GP is the sole general partner of VV V CAN and VV V CAN is the sole
general partner of VVC CAN. Each of VV CAN GP and VV V CAN share voting and dispositive power over the shares held by VVC CAN. Colella, Davis, Link, Bolzon, Praeger, Nielsen and Woiwode are directors of VV V CAN GP and share voting and dispositive
power over the shares held by VVC CAN.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Vantage I, L.P.

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;458,221 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;458,221 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 458,221 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.5% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; PN

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VV I. VV I GP-GP is the sole general partner
of VV I GP and VV I GP is the sole general partner of VV I. Each of VV I GP-GP and VV I GP share voting and dispositive power over the shares held by VV I. Bolzon, Davis, Ozawa, Woiwode and Praeger are the
managing directors of VV I GP-GP and share voting and dispositive power over the shares held by VV I.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Vantage I GP, L.P.

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;458,221 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;458,221 shares of Common Stock (2) |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 458,221 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.5% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; PN

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VV I. VV I GP-GP is the sole general partner
of VV I GP and VV I GP is the sole general partner of VV I. Each of VV I GP-GP and VV I GP share voting and dispositive power over the shares held by VV I. Bolzon, Davis, Ozawa, Woiwode and Praeger are the
managing directors of VV I GP-GP and share voting and dispositive power over the shares held by VV I.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

CUSIP No. 39868T105 13G

1. &nbsp;&nbsp;&nbsp;&nbsp; Name of Reporting Persons &nbsp;&nbsp;&nbsp;&nbsp; Versant Vantage I GP-GP, LLC

2. &nbsp;&nbsp;&nbsp;&nbsp; Check the Appropriate Box
if a Member of a Group (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; (a) ☐ (b) ☒(1)

3. &nbsp;&nbsp;&nbsp;&nbsp; SEC Use Only

4. &nbsp;&nbsp;&nbsp;&nbsp; Citizenship or Place of
Organization &nbsp;&nbsp;&nbsp;&nbsp; Delaware

---

| | | |
|:---|:---|:---|
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 5. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 6. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Voting Power<br>&nbsp;&nbsp;&nbsp;&nbsp;458,221 shares of Common Stock (2) |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 7. | &nbsp;&nbsp;&nbsp;&nbsp;Sole Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;0 |
| &nbsp;&nbsp;&nbsp; Number of<br> Shares<br> Beneficially <br> Owned by<br> Each<br> Reporting<br> Person<br> With | 8. | &nbsp;&nbsp;&nbsp;&nbsp;Shared Dispositive Power<br>&nbsp;&nbsp;&nbsp;&nbsp;458,221 shares of Common Stock (2) |

---

9. &nbsp;&nbsp;&nbsp;&nbsp; Aggregate Amount Beneficially Owned by Each Reporting Person &nbsp;&nbsp;&nbsp;&nbsp; 458,221 shares of Common Stock (2)

10. &nbsp;&nbsp;&nbsp;&nbsp; Check if the Aggregate
Amount in Row 9 Excludes Certain Shares (See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; ☐

11. &nbsp;&nbsp;&nbsp;&nbsp; Percent of Class
Represented by Amount in Row 9 &nbsp;&nbsp;&nbsp;&nbsp; 0.5% (3)

12. &nbsp;&nbsp;&nbsp;&nbsp; Type of Reporting Person
(See Instructions) &nbsp;&nbsp;&nbsp;&nbsp; OO

------

(1) This Schedule 13G is filed by the Reporting Persons. The Reporting Persons expressly disclaim status as a
"group" for purposes of this Schedule 13G.

(2) These shares are held by VV I. VV I GP-GP is the sole general partner
of VV I GP and VV I GP is the sole general partner of VV I. Each of VV I GP-GP and VV I GP share voting and dispositive power over the shares held by VV I. Bolzon, Davis, Ozawa, Woiwode and Praeger are the
managing directors of VV I GP-GP and share voting and dispositive power over the shares held by VV I.

(3) This calculation is based upon 83,366,277 shares of the Issuer's Common Stock outstanding as of
November 1, 2022, as reported in the Issuer's Form 10-Q.

------

**Introductory Note**: This Amendment No. 3 (this "Amendment") amends and supplements the Schedule 13G originally filed by the Reporting Persons with the United States Securities and Exchange Commission (the "Commission") on February 14, 2019, as amended by Amendment No. 1 filed with the Commission on February 16, 2021 and Amendment No. 2 filed with the Commission on February 11, 2022 (collectively, the "Original Schedule 13G"). Only those items that are hereby reported are amended; all other items reported in the Original Schedule 13G remain unchanged. Information given in response to each item shall be deemed incorporated by reference in all other items, as applicable. Capitalized terms not defined in this Amendment have the meanings ascribed to them in the Original Schedule 13G.

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| | |
|:---|:---|
| **Item 4** | **Ownership.**  |

---

The following information with respect to the ownership of the Issuer's Common Stock by the Reporting Persons filing this statement on Schedule 13G is provided as of December 31, 2022:

(a) Amount beneficially owned:

See line 9 of each cover sheet.

(b) Percent of class:

See line 11 of each cover sheet.

(c) Number of shares as to which the person has:

i. Sole power to vote or to direct the vote: see line 5 of cover sheets.\*

ii. Shared power to vote or to direct the vote: see line 6 of cover sheets.\*

iii. Sole power to dispose or to direct the disposition of: see line 7 of cover sheets.\*

iv. Shared power to dispose or to direct the disposition of: see line 8 of cover sheets.\*

\* Each Reporting Person disclaims beneficial ownership of such shares of Common Stock, except for the shares, if any, such Reporting Person holds of record and to the extent of its pecuniary interest therein.

**Item 5.** **Ownership of Five Percent or Less of a Class.** <br>

If this statement is being filed to report the fact that as of the date hereof the Reporting Persons have ceased to be the beneficial owners of more than 5 percent of the class of securities, check the following [X].

------

**SIGNATURE** 

After reasonable inquiry and to the best of my knowledge and belief, I certify that the information set forth in this statement is true, complete and correct.

February 9, 2023

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| | |
|:---|:---|
| **Versant Venture Capital V, L.P.** | **Versant Venture Capital V, L.P.** |
| By: | Versant Ventures V, LLC |
| Its: | General Partner |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Affiliates Fund V, L.P.** | **Versant Affiliates Fund V, L.P.** |
| By: | Versant Ventures V, LLC |
| Its: | General Partner |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Ophthalmic Affiliates Fund I, L.P.** | **Versant Ophthalmic Affiliates Fund I, L.P.** |
| By: | Versant Ventures V, LLC |
| Its: | General Partner |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Ventures V, LLC** | **Versant Ventures V, LLC** |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Venture Capital V (Canada) LP** | **Versant Venture Capital V (Canada) LP** |
| By: | Versant Ventures V (Canada), L.P. |
| Its: | General Partner |
| By: | Versant Ventures V GP-GP (Canada), Inc. |
| Its: | General Partner |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Ventures V (Canada), L.P.** | **Versant Ventures V (Canada), L.P.** |
| By: | Versant Ventures V GP-GP (Canada), Inc. |
| Its: | General Partner |

---

------

---

| | |
|:---|:---|
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Ventures V GP-GP (Canada), Inc.** | **Versant Ventures V GP-GP (Canada), Inc.** |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Vantage I, L.P.** | **Versant Vantage I, L.P.** |
| By: | Versant Vantage I GP, L.P. |
| Its: | General Partner |
| By: | Versant Vantage I GP-GP, LLC |
| Its: | General Partner |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Vantage I GP, L.P.** | **Versant Vantage I GP, L.P.** |
| By: | Versant Vantage I GP-GP, LLC |
| Its: | General Partner |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |
| **Versant Vantage I GP-GP, LLC** | **Versant Vantage I GP-GP, LLC** |
| By: | /s/ Max Eisenberg |
|  | Max Eisenberg, Chief Operating Officer |

---