Patent Publication Number: US-2021189472-A1

Title: Sexual maturation in rainbow trout

Description:
CROSS-REFERENCE TO RELATED APPLICATIONS 
     This application claims priority to European Application 19217649.3 filed on Dec. 18, 2020, which is incorporated herein by reference in its entirety. 
     FIELD OF THE DISCLOSURE 
     The present disclosure relates generally to polymorphisms, and in particular single nucleotide polymorphisms (SNP) associated with late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ). In particular, the present disclosure provides methods for predicting late onset of sexual maturation in rainbow trout, methods for selecting a rainbow trout having late onset of sexual maturation and kit suitable for carrying out said methods. The present disclosure further provides rainbow trout cells, sperm and unfertilized eggs carrying at least one allele conferring late onset of sexual maturation in their genome. 
     BACKGROUND 
     Aquaculture has developed rapidly over the last decades in order to meet the growing demand for marine foods, a demand that cannot be met by increased harvest of wild stocks as they already are under stress from commercial fisheries. Many species, ranging from sea urchins to anadromous fish, such as salmon and trout, are now farmed effectively at a large scale. However, there are at present very few breeding programs established for cultured marine species. 
     Atlantic salmon ( Salmo salar ) and Rainbow Trout ( Oncorhynchus mykiss ) are two of the species where breeding programs have been successfully established. Both species have been actively bred for more than four decades, and substantial advances have been made with regard to disease resistance and growth-related traits. Among the growth-related traits, age and size at sexual maturation is one of the most important from an economically point of view. 
     Sexual maturation, or puberty, is the process in which a juvenile individual develops into a sexually functional adult with the ability to produce gametes and gains the somatic and behavioral competence to function as a mating partner and a parent. Gonadal development is initiated already before hatching with the initiation of germ cell formation, gonadal development and continues in stages until final sexual maturation occurs. The heavy investments in gonadal development and gamete production has several negative consequences for growth rate, welfare and product quality of farmed fish. 
     Growth rate is typically high at onset of puberty. However, a few months into the maturation process, feeding will cease, and somatic weight will decrease as resources are routed to gonadal development and gamete production. Early sexual maturation is therefore considered a serious drawback for aquaculture due to its undesirable impacts on growth. 
     Further, maturation also represents a welfare issue since the anadromous fish gradually will lose tolerance to seawater due to compromised hypo-osmoregulation ability during maturation. When the wild fish begins to mature, they typically migrate to their native rivers, an escape that is not possible for farmed fish due to their confinement in cages in seawater facilities. Without the possibility to escape seawater, the consequence of reduced sea water tolerance is dehydration, stress and possible negative effects on the immune system. Thus, early sexual maturation is therefore considered a serious drawback for aquaculture due to its undesirable impacts on fish welfare. 
     It has also been reported that sexual maturation causes reduction in fat- and protein content of fish filets; loss of flesh pigmentation, as the pigment is shifted to skin and gametes; and changes in smell and taste of the fish filets. Thus, early sexual maturation is therefore considered a serious drawback for aquaculture due to its undesirable impacts on product quality. 
     The behavior of farmed fish also changes during sexual maturation. Behavioral changes have been observed in females, but these changes seem to especially affect males that take on a dominant role during mate competition and become more aggressive toward other males. 
     Further, when immature salmon escape, they are likely to run out to sea where survival is low; subsequently they are less likely to survive to mate with wild fish. A mature fish is likely to seek up-river in order to mate. Thus, keeping the fish in an immature stage throughout the production cycle will not only be beneficial to the industry with regards to growth, product quality and animal welfare, but is also an effort that could benefit the wild fish strains whose gene pools are less likely to be negatively impacted by farmed fish with late maturation. 
     Thus, there is a need for anadromous fish that enters sexual maturation at a late stage. In particular, there is a need for anadromous fish that remain sexually immature throughout the production cycle. 
     Maturation seems to be controlled by several intrinsic factors such as size, growth rate and fat deposition, in addition to several extrinsic factors such as photoperiod, temperature, diet, stress exposure and competition. When these intrinsic and extrinsic factors together meet certain conditions, the brain-pituitary-gonad axis is triggered, and maturation is continued to the next step. 
     Based on the knowledge of how the extrinsic factors affect sexual maturation of fish, it has been common practice to expose farmed fish to continuous light to delay sexual maturation and thereby avoid the negative effects of early maturation. 
     However, increased water temperature is known to be associated with early maturation, and the increasing ocean temperatures caused by global warming may therefore to some extent overrun the inhibitory effect of continuous light on maturation. In the future it may therefore be necessary to use other means than continuous light to delay sexual maturation of farmed fish. 
     An alternative to the use of continuous light is to look at the genetic variability in the search for a genetic pattern associated with late onset of sexual maturation. However, as will be discussed further below, there are some challenges that need to be solved if fish are to be selected for late sexual maturation based on genetics. 
     Farmed fish has typically been selected for improved growth and is provided with optimal feeding condition. The result being that farmed anadromous fish often show a phenotypic response to their improved growth conditions where both age and size at puberty is reduced, accompanying the high growth rate and large size achieved through selective breeding. For this reason, it may be challenging for breeding programs to select for late maturity as the phenotypic response can mask the genetic variability in age and size at maturity. 
     There is therefore a need for improved methodologies for assessing late onset of sexual maturation in rainbow trout, particularly methodologies that allow the direct assaying and selection of individual&#39;s having late onset of sexual maturation. 
     Journal of fish biology, vol 78, no. 2, 11 Jan. 2011, pages 602-623 teaches correlations between quantitative trait locus (QTL) involved in developmental rate and age at maturation in rainbow trout  Oncorhynchus mykiss.    
     Molecular Genetics and Genomics, springer, Berlin, De, vol. 277, no. 6, 17 Feb. 2007, pages 647-661, XP019517829 relates to putative growth and maturation timing QTL locations in Arctic charr, several of these regions showing homologies to the identified maturation timing QTL regions in Atlantic salmon and/or rainbow trout. 
     Aquaculture, vol. 410-411, 1 Oct. 2013, pages 164-171, XP055692262, Amsterdam, NL relates to QTL associated with smolting and maturation in rainbow trout. 
     Neogen provides a high-density genotyping array for high-resolution genetics research and aquaculture breeding in rainbow trout. The array includes 57,501 markers and is available in both 96 and 384 formats. 
     SUMMARY OF THE INVENTION 
     The present inventors have solved this need by having identified polymorphisms, and in particular single nucleotide polymorphisms (SNP), within the genome of rainbow trout which are associated with late onset of sexual maturation. 
     In a first aspect, a method for predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ) comprises determining the presence of at least one allele, such as at least two alleles, conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain aspects, provided is a method for predicting late onset of sexual maturation in rainbow trout, the method comprising determining the identity of a nucleotide of at least one allele, optionally of at least two alleles, of at least one single nucleotide polymorphism (SNP) associated with late onset of sexual maturation within the genome of said rainbow trout, said at least one SNP being selected from the group consisting of SNP #1 to SNP #164. 
     According to certain embodiments, provided is a method for predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 164, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 1 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     In a second aspect, a method for selecting a rainbow trout having late onset of sexual maturation comprises determining the presence of at least one allele, such as at least two alleles, conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; and selecting said rainbow trout as having late onset of sexual maturation when the at least one late maturation allele is present; wherein the at least one late maturation allele is an allele of at least one SNP, the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain aspects, a method for selecting a rainbow trout having late onset of sexual maturation comprises determining the identity of a nucleotide of at least one allele, such as at least two alleles, of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being selected from the group consisting of SNP #1 to SNP #164; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain aspects, a method for selecting a rainbow trout having late onset of sexual maturation comprises determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 164, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 1 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     In a further aspect, provided herein is a rainbow trout, such as an isolated rainbow trout, or a progeny thereof having late onset of sexual maturation comprising within its genome at least one allele, such as at least two alleles, conferring late onset of sexual maturation (“late maturation allele”); wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     Late onset of sexual maturation will have positive impacts on growth since the fish does not need to invest energy in gonadal development and gamete production. Further, late onset of sexual maturation will also help the fish to maintain tolerance to seawater for a prolonged period of time which will have a positive effect on fish welfare. Late onset of sexual maturation may also i) improve fat- and protein content of fish filets; ii) improve flesh pigmentation; and iii) improve smell and taste of the fish filet. Late onset of sexual maturation may also reduce aggressive behavior of the fish, in particular for male fish, for a prolonged period of time. Further, gene pools of wild fish strains are less likely to be negatively impacted by farmed fish with late maturation. 
     Thus, the purpose of providing an (isolated) rainbow trout having late onset of sexual maturation may include:
         improving growth of the (isolated) rainbow trout;   maintaining optimal tolerance to seawater for a prolonged period of time;   improving fat- and protein content of fish filets;   improving flesh pigmentation;   improving smell and taste of the fish filet;   reducing undesirable behavior, such as aggressive behavior, commonly associated with mature fish; and/or   reducing negative impact of escaped farmed fish on gene pools of wild fish strains.       

     In view of the above, it is clear that the fish farming industry has a need for an (isolated) rainbow trout having late onset of sexual maturation. Thus, another purpose of providing an (isolated) rainbow trout having late onset of sexual maturation is to fulfill this need by providing a plurality of such fish. A plurality of such fish may be obtained by using the (isolated) rainbow trout for breeding purposes. 
     According to certain aspects, a rainbow trout, such as an isolated rainbow trout, or progeny thereof comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     In another aspect, provided herein is a population of rainbow trout, such as an isolated population of rainbow trout, each individual rainbow trout within the population being a rainbow trout according to the further aspect referred to above. 
     In another aspect, provided is a population of rainbow trout, such as an isolated population of rainbow trout, wherein at least 50%, such as at least 60%, at least 70%, at least 80%, at least 90%, at least 95% or at least 99% of the rainbow trout within the population comprises within its genome at least one allele conferring late onset of sexual maturation (“late maturation allele”); the at least one late maturation allele being an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     The purpose of providing an (isolated) rainbow trout having late onset of sexual maturation is discussed above. However, the fish farming industry is farming a population of fish, not single individuals, and it is common that all fish are treated in a similar manner. It is therefore an advantage if the population of fish at each facility is as homogenous as possible. By having a homogenous population of fish, the response to feeding and other treatments would be expected to be similar within this population. 
     Further provided is a rainbow trout or progeny thereof which comprises in its genome at least one allele, such as at least two alleles, conferring late onset of sexual maturation obtainable by a process comprising the steps of: genotyping the rainbow trout, selecting individuals having at least one allele, preferably two alleles, conferring late onset of sexual maturation (“late maturation allele”); and mating individuals in such a way that at least one individual within each mated pair has two alleles conferring late onset of sexual maturation. 
     In a third aspect, an isolated rainbow trout cell comprises within its genome at least one allele conferring late onset of sexual maturation (“late maturation allele”); wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain embodiments, provided herein is a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     In another aspect, a population of rainbow trout cells is provided, such as an isolated population of rainbow trout cells, each individual cell within the population being a cell according to the third aspect. 
     In another aspect, a population of rainbow trout cells is provided, such as an isolated population of rainbow trout cells, wherein at least 50%, such as at least 60%, at least 70%, at least 80%, at least 90%, at least 95% or at least 99% of the rainbow trout cells within the population comprises within its genome at least one allele conferring late onset of sexual maturation (“late maturation allele”); the at least one late maturation allele being an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     In a fourth aspect, an isolated rainbow trout egg or spermatozoa comprises within its genome at least one allele conferring late onset of sexual maturation (“late maturation allele”); wherein the isolated rainbow trout egg is unfertilized; and the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     Late onset of sexual maturation will have positive impacts on growth since the fish does not need to invest energy in gonadal development and gamete production, Further, late onset of sexual maturation will also help the fish to maintain tolerance to seawater for a prolonged period of time which will have a positive effect on fish welfare. Late onset of sexual maturation may also i) improve fat- and protein content of fish filets; ii) improve flesh pigmentation; and iii) improve smell and taste of the fish filet. Late onset of sexual maturation may also reduce aggressive behavior of the fish, in particular for male fish, for a prolonged period of time. Further, gene pools of wild fish strains are less likely to be negatively impacted by farmed fish with late maturation. 
     Thus, the purpose of providing an (isolated) rainbow trout having late onset of sexual maturation may include:
         improving growth of the (isolated) rainbow trout;   maintaining optimal tolerance to seawater for a prolonged period of time;   improving fat- and protein content of fish filets;   improving flesh pigmentation;   improving smell and taste of the fish filet;   reducing undesirable behavior, such as aggressive behavior, commonly associated with mature fish; and/or   reducing negative impact of escaped farmed fish on gene pools of wild fish strains.       

     In view of the above, it is clear that there is a need in the fish farming industry for an (isolated) rainbow trout having late onset of sexual maturation. This need may be fulfilled by providing a plurality of such fish. A plurality of such fish may be obtained by breeding using the isolated rainbow trout egg or spermatozoa according to the fourth aspect of the present invention. 
     Thus, the (isolated) rainbow trout egg or spermatozoa according to the fourth aspect is intended to be used for breeding purposes for obtaining an (isolated) rainbow trout having late onset of sexual maturation. Said in other words, the (isolated) rainbow trout egg or spermatozoa according to the fourth aspect invention will provide the genetic material necessary to obtain an (isolated) rainbow trout having late onset of sexual maturation and thereby:
         improve growth of the (isolated) rainbow trout;   maintain optimal tolerance to seawater for a prolonged period of time;   improve fat- and protein content of fish filets;   improve flesh pigmentation;   improve smell and taste of the fish filet;   reduce undesirable behavior, such as aggressive behavior, commonly associated with mature fish; and/or   reduce negative impact of escaped farmed fish on gene pools of wild fish strains.       

     According to certain aspects, provided is an isolated rainbow trout egg or spermatozoa which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     In another aspect, provided is a population of rainbow trout eggs or spermatozoa, such as an isolated population of rainbow trout egg or spermatozoa, each individual egg or spermatozoa within the population being an egg or spermatozoa according to the present disclosure. 
     Further provided is a population of rainbow trout eggs or spermatozoa, such as an isolated population of rainbow trout eggs or spermatozoa, wherein at least 50%, such as at least 60%, at least 70%, at least 80%, at least 90%, at least 95% or at least 99% of the rainbow trout eggs or spermatozoa within the population comprises within its genome at least one allele conferring late onset of sexual maturation (“late maturation allele”); the at least one late maturation allele being an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1 and each individual rainbow trout egg within the population being unfertilized. 
     The purpose of providing an (isolated) rainbow trout egg or spermatozoa according to the fourth aspect is discussed above. When these eggs or spermatozoa are used for breeding purposes, they are not used individually but rather as a population of eggs or spermatozoa. By using a population of such eggs or spermatozoa, the resulting fish population will be homogenous with respect to the late maturation allele. Thus, the purpose of providing a population of (isolated) rainbow trout eggs or spermatozoa according is to provide a fish population which will be homogenous with respect to the late maturation allele. 
     In a fifth aspect, provided is oligonucleotide, such as an isolated oligonucleotide, comprising at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 164, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 1 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof, provided that said nucleotide substitutions are not at position 31 of said derived sequence, with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     The purpose of obtaining an oligonucleotide according to the fifth aspect is to provide a tool to be used in the method according to the first and/or second aspect. The oligonucleotide according to the fifth aspect is suitable for determining the presence of at least one allele, such as at least two alleles, conferring late onset of sexual maturation (“late maturation allele”) within a genome of a rainbow trout. 
     In one embodiment according to the fifth aspect, the oligonucleotide has a total length of 8 to 61 nucleotides, such as a total length of 10 to 61 nucleotides, a total length of 12 to 61 nucleotides, a total length of 14 to 61 nucleotides, a total length of 16 to 61 nucleotides, a total length of 18 to 61 nucleotides or a total length of 20 to 61 nucleotides. 
     In another embodiment according to the fifth aspect, the oligonucleotide has a total length of 8 to 50 nucleotides, such as a total length of 10 to 50 nucleotides, a total length of 12 to 50 nucleotides, a total length of 14 to 50 nucleotides, a total length of 16 to 50 nucleotides, a total length of 18 to 50 nucleotides or a total length of 20 to 50 nucleotides. 
     In another embodiment according to the fifth aspect, the oligonucleotide has a total length of 8 to 40 nucleotides, such as a total length of 10 to 40 nucleotides, a total length of 12 to 40 nucleotides, a total length of 14 to 40 nucleotides, a total length of 16 to 40 nucleotides, a total length of 18 to 40 nucleotides or a total length of 20 to 40 nucleotides. 
     Also provided in a sixth aspect is a kit for predicting late onset of sexual maturation in rainbow trout, the kit comprising at least one of the oligonucleotides according to the fifth aspect. 
     Further provided in a seventh aspect is a kit for selecting a rainbow trout having late onset of sexual maturation, the kit comprising at least one of the oligonucleotides according to the fifth aspect. 
     In an eighth aspect, a method for obtaining gametes from rainbow trout ( Oncorhynchus mykiss ) having late onset of sexual maturation comprises:
         providing an initial population of rainbow trout;   obtaining a nucleic acid sample from at least one individual rainbow trout within said initial population;   detecting in each of said nucleic acid samples the presence of at least one allele conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1;   selecting a rainbow trout from said initial population based on the presence of the at least one late maturation allele in the nucleic acid samples; and   isolating gametes from said selected trout.       

     In one embodiment according to the eight aspect, said gametes are eggs. 
     In one embodiment according to the eight aspect, said gametes are spermatozoa. 
     In a ninth aspect, a method for obtaining rainbow trout gametes comprising within their genome at least one late maturation allele comprises
         providing an initial population of rainbow trout gametes;   obtaining a nucleic acid sample from at least one individual gamete within said initial population;   detecting in each of said nucleic acid samples the presence of at least one allele conferring late onset of sexual maturation (“late maturation allele”) within the genome of said gamete; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1; and   selecting and isolating a gamete from said initial population based on the presence of the at least one late maturation allele in the nucleic acid samples; and       

     In one embodiment according to the nineth aspect, said gametes are eggs. 
     In one embodiment according to the nineth aspect, said gametes are spermatozoa. 
     In a tenth aspect, a method for obtaining rainbow trout having late onset of sexual maturation comprises
         providing rainbow trout eggs comprising within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1;   providing rainbow trout spermatozoa; and   utilizing the rainbow trout eggs and the rainbow trout spermatozoa to produce rainbow trout having late onset of sexual maturation.       

     In one embodiment according to the tenth aspect, the rainbow trout eggs are obtained by the foregoing methods. 
     In one embodiment according to the tenth aspect, the rainbow trout spermatozoa comprises within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     In one embodiment according to the tenth aspect, the rainbow trout spermatozoa
         comprise within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1; and   are obtained by the method according to the eight and/or nineth aspect       

     In an eleventh aspect, a method for obtaining rainbow trout having late onset of sexual maturation comprises
         providing rainbow trout spermatozoa comprising within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1;   providing rainbow trout eggs; and   utilizing the rainbow trout eggs and the rainbow trout spermatozoa to produce rainbow trout having late onset of sexual maturation.       

     In one embodiment according to the eleventh aspect, the rainbow trout spermatozoa are obtained by the method according to the eight and/or ninth aspect. 
     In one embodiment according to the eleventh aspect, the rainbow trout eggs comprises within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     In one embodiment according to the tenth aspect, the rainbow trout eggs:
         comprise within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1; and   are obtained by the method according to the eight and/or nineth aspect of the present invention.       

     In a twelfth aspect, a method for obtaining fertilized rainbow trout eggs which will mature into a rainbow trout having late onset of sexual maturation comprises
         providing rainbow trout eggs comprising within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1;   providing rainbow trout spermatozoa; and   fertilizing the rainbow trout eggs using the rainbow trout spermatozoa thereby producing a fertilized rainbow trout egg which will mature into a rainbow trout having late onset of sexual maturation.       

     In one embodiment according to the twelfth aspect, the rainbow trout eggs are obtained by the method according to the eight and/or nineth aspect. 
     In one embodiment according to the twelfth aspect, the rainbow trout spermatozoa comprises within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     In one embodiment according to the twelfth aspect, the rainbow trout spermatozoa:
         comprise within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1; and   are obtained by the method according to the eight and/or nineth aspect of the present invention.       

     In a thirteenth aspect, a method for obtaining fertilized rainbow trout eggs which will mature into a rainbow trout having late onset of sexual maturation comprises
         providing rainbow trout spermatozoa comprising within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1;   providing rainbow trout eggs; and   fertilizing the rainbow trout eggs using the rainbow trout spermatozoa thereby producing a fertilized rainbow trout egg which will mature into a rainbow trout having late onset of sexual maturation.       

     In one embodiment according to the thirteenth aspect, the rainbow trout spermatozoa are obtained by the method according to the eight and/or nineth aspect. 
     In one embodiment according to the thirteenth aspect, the rainbow trout eggs comprises within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     In one embodiment according to the thirteenth aspect, the rainbow trout eggs
         comprise within their genome at least one late maturation allele; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1; and   are obtained by the method according to the eight and/or nineth aspect of the present invention.       

     In one embodiment according to the thirteenth aspect, the method further comprises a step of rinsing the fertilized rainbow trout eggs; preferably using water. 
     In one embodiment according to the thirteenth aspect, the method further comprising subjecting the fertilized rainbow trout eggs to disinfection treatment. 
     In one embodiment according to the thirteenth aspect, the method further comprising the steps of:
         rinsing the fertilized rainbow trout eggs, preferably using water, to obtain rinsed fertilized rainbow trout eggs; and   subjecting the rinsed fertilized rainbow trout eggs to disinfection treatment.       

     In a fourteenth aspect, a method for obtaining rainbow trout having late onset of sexual maturation comprises
         providing an initial population of rainbow trout;   obtaining a nucleic acid sample from at least one individual rainbow trout within said initial population;   detecting in each of said nucleic acid samples the presence of at least one allele conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1;   selecting a rainbow trout from said initial population based on the presence of the at least one late maturation allele in the nucleic acid samples; and   mating said selected rainbow trout with a second rainbow trout to produce progeny rainbow trout having late onset of sexual maturation.       

     In one embodiment according to the fourteenth aspect, at least one rainbow trout within said mating pair is homozygous for the late maturation allele. 
     In one embodiment according to the fourteenth aspect, each rainbow trout within said mating pair is homozygous for the late maturation allele. 
    
    
     
       BRIEF DESCRIPTION OF THE DRAWINGS 
       The present invention is described in detail by reference to the following drawings: 
         FIG. 1 a    shows Genome Wide Association Study on maturity for A17 Nucleus eggs from 2017 (AquaGen). X-axis=Chromosome number. Y-axis −log 10(P). A number of chromosomes have distinct peaks, but the most distinctive and highly significant QTL was detected on chromosome 28 (see  FIG. 1 b   ). 
         FIG. 1 b    is a regional Manhattan plot showing the QTL on chromosome 28 and thus represents an extract of the information related to chromosome 28 presented in  FIG. 1   a.  X-axis=position on chromosome 28. Y-axis −log 10(P). 
       A distinctive and highly significant QTL for maturity is detected on chromosome 28. The most preferred SNPs are marked in light grey color (SNP #120, 127-131, 134-135, 137-138, 140, 142, 143-144, 146-147; 16 SNPs in total). All of these SNPs (16 in total) have a p-value lower than 3.12e-7 and are located within a genomic region of 1.9 megabases. 
         FIG. 2 a    illustrates the effect of SNP #140 on sexual maturation in male and female rainbow trout. X-axis indicates the allelic variants of SNP #140. Y-axis indicates the fraction of fish that are sexually mature. 
       
         
           
             
                 
                 
               
                 
                     
                 
                 
                   Genotype 
                   Allelic variants 
                 
                 
                     
                 
               
              
                 
                   qq 
                   homozygous for normal maturation allele 
                 
                 
                   qQ 
                   heterozygous for late maturation allele 
                 
                 
                   QQ 
                   homozygous for late maturation allele 
                 
                 
                     
                 
              
             
           
         
       
       The figure shows how the genotype correlate with maturation without correcting for polygenic effect (effect of all other genes, i.e. family effect) and other potential co-factors. The effect size of the allele is adjusted for the polygenic effect and sex; hence it is a more accurate measure. The effect is significant in both male and female Trout; but seems to be somewhat larger in female Trout. 
         FIG. 2 b    shows unadjusted effect of SNP #140 on sexual maturation in both sexes combined. X-axis indicates the allelic variants of SNP #140. Y-axis indicates the fraction of fish that are sexually mature. 
       
         
           
             
                 
                 
               
                 
                     
                 
                 
                   Genotype 
                   Allelic variants 
                 
                 
                     
                 
               
              
                 
                   qq 
                   homozygous for normal maturation allele 
                 
                 
                   qQ 
                   heterozygous for late maturation allele 
                 
                 
                   QQ 
                   homozygous for late maturation allele 
                 
                 
                     
                 
              
             
           
         
       
         FIG. 3  shows a squared correlation (R2) matrix between the most preferred SNPs on chromosome 28 (SNP #120, 127-131, 134-135, 137-138, 140, 142, 143-144, 146-147; 16 SNPs in total). The SNPs are clustered together based on R2 and all show varying degree of elevated squared correlations to each other. 
     
    
    
     DETAILED DESCRIPTION 
     Unless specifically defined herein, all technical and scientific terms used have the same meaning as commonly understood by a skilled artisan in the fields of genetics, biochemistry, and molecular biology. 
     All methods and materials similar or equivalent to those described herein can be used in the practice or testing of the present invention, with suitable methods and materials being described herein. All publications, patent applications, patents, and other references mentioned herein are incorporated by reference in their entirety. In case of conflict, the present specification, including definitions, will prevail. 
     Where a numerical limit or range is stated herein, the endpoints are included. Also, all values and sub ranges within a numerical limit or range are specifically included as if explicitly written out. 
     Definitions 
     As used herein, late onset of sexual maturation means that an individual having late onset of sexual maturation has a higher probability of maturing later than a random individual (under the same conditions, i.e. under same intrinsic and extrinsic factors) with whom it is comparable. Two individuals are comparable if they are, with regards to all discriminating factors except the genotype at the SNP which is used for predicting late onset of sexual maturation, random representatives of one and the same population of rainbow trout. 
     As used herein, a “late maturation allele” is an allele conferring late onset of sexual maturation. This means that a rainbow trout having such allele at the position of a polymorphism detailed herein shows late onset of sexual maturation. The “late maturation allele” may identify a single nucleotide polymorphism that can be used to detect or determine the degree of late sexual maturation. 
     As used herein, a “polymorphism” is a variation in a genomic sequence. In particular, a polymorphism is a position on the genome where different allelic variants are generally found between individuals of a population, or between individuals from different populations. The polymorphism may be a single nucleotide difference present at a locus or may be an insertion or deletion of one or a few nucleotides at a position of a gene. 
     As used herein, a “single nucleotide polymorphism” or “SNP” refers to a single base (nucleotide) polymorphism in a DNA sequence among individuals in a population. As such, a single nucleotide polymorphism is characterized by the presence in a population of one or two, three or four different nucleotides (i.e. adenine, cytosine, guanine or thymine), typically less than all four nucleotides, at a particular locus in a genome, such as the genome of rainbow trout. 
     As used herein, “polymorphic sequence” refers to a nucleotide sequence including a polymorphic site at which a SNP or another type of polymorphism occurs. 
     As used herein, a “polymorphic site” is the locus or position within a given sequence at which divergence occurs. Preferred polymorphic sites have at least two alleles, each occurring at frequency greater than 1%, and more preferably greater than 10%. Those skilled in the art will recognize that nucleic acid molecules may be double-stranded molecules and that reference to a particular site on one strand refers, as well, to the corresponding site on a complementary strand. In defining a polymorphic site or allele reference to an adenine, a thymine, a cytosine, or a guanine at a particular site on one strand of a nucleic acid molecule also defines the thymine, adenine, guanine, or cytosine (respectively) at the corresponding site on a complementary strand of the nucleic acid. 
     Herein, when a polymorphism is specified as having a particular allele, then it is understood that that particular allele goes together with the sequence given for the polymorphism. For example, when it is said that thymine is the late maturation allele of Affx-88940276, then it is understood that the late maturation allele of Affx-88940276 harbors a thymine nucleotide in the polymorphic site defined in Table 2 when the DNA is read in the direction defined in Table 2. In other words, as stated in Table 2, the late maturation form of the DNA sequence of Affx-88940276 (with flanking sequences) is CCGCCAGTGTCGATGCCAAACCCTTGAAAATAGGT[C/T]GGGAAAGGAAATCC TTTCCCTCTAAATATCTGTCG (SEQ ID NO: 329; polymorphic site in brackets) when read in the direction defined in Table 2. When read in the opposite direction, the sequence of Affx-88940276 (with flanking sequence) is GCTGTCTATAAATCTCCCTTTCCTAAAGGAAAGGG[T/C]TGGATAAAAGTTCCC AAACCGTAGCTGTGACCGCC (SEQ ID NO: 330; polymorphic site in brackets). Although only one direction is used when late maturation alleles and normal maturation alleles are defined herein, the two read directions are equivalent. 
     As used herein, a “sample”, such as a biological sample that includes nucleic acid molecules, is a sample obtained from a rainbow trout, including, but not limited to, cells, tissue, and bodily fluids. 
     As used herein, an “oligonucleotide” is a plurality of joined nucleotides joined by native phosphodiester bonds, typically from 8 to 300 nucleotides in length. 
     As used herein, “probes” and “primer” are isolated oligonucleotides of at least 8 nucleotides, such as at least 10 nucleotides, capable of hybridizing to a target nucleic acid. 
     As used herein, “isolated” means that an organism or a biological component, such as a cell, population of cells or a nucleic acid molecule, has been separated from its natural environment. 
     As used herein, “genetic linkage” refers to the tendency of polymorphisms that are located close to each other on a chromosome to be inherited together during meiosis. Thus, polymorphisms located close to each other on the same chromosome are said to be genetically linked. Alleles at two such genetically linked loci are co-inherited (from parents to offspring) more often than they are not. Assume, for example, two polymorphisms; polymorphism A having alleles A1 and A2, and polymorphism B having alleles B1 and B2. Assume further that a given rainbow trout carries all of the alleles A1, A2, B1, and B2 (in other words, this rainbow trout is heterozygous at both marker and marker B). If alleles A1 and B1 are, in this particular rainbow trout, located on the same chromosome copy, then alleles A1 and B1 are co-inherited, to the offspring of the rainbow trout, more often than not. 
     As used herein, “genetic linkage analysis” refers to a statistical procedure where genotype data, coming from sets of animals comprising parents and their offspring, are investigated in order to test for the presence of genetic linkage between polymorphisms. Genetic linkage analysis can be used in order to assign polymorphisms to chromosomes, provided that the analysis incorporates polymorphisms that have already been assigned to chromosome using, for example, Fluorescence In Situ Hybridization. 
     As used herein “Fluorescence In Situ Hybridization” or “FISH” refers to a technique that detect the presence or absence of specific DNA sequences on chromosomes. FISH can be used in order to assign known DNA polymorphisms to chromosomes. 
     “Centi-Morgen” is a unit of measurement, used to describe genetic distances, where genetic distance is a measure of the extent to which two polymorphisms are genetically linked. 
     Linkage disequilibrium (LD) or, more precisely, gametic phase linkage disequilibrium, is used in order to describe the co-inheritance of alleles at genetically linked polymorphisms, at the population level. Assume, for example, two polymorphisms located on the same chromosome; polymorphism A having alleles A1 and A2, and polymorphism B having alleles B1 and B2. All copies of the chromosome in question will harbor a combination of alleles at the two loci (i.e. a haplotype), and there are four possible haplotypes: A1-B1, A1-B2, A2-B1, and A2-B2. The two loci are in said to be LD with each other if the number of A1-B1 and A2-B2 haplotypes within the population are significantly larger or significantly smaller than the number of A1-B2 and A2-B1 haplotypes. 
     Polymorphisms and Late Maturation Allele(s) of the Invention 
     The present inventors have identified several quantitative trait locus (QTL) responsible for a significant fraction of the genetic variation related to sexual maturation in rainbow trout. More specifically, the present inventors have identified polymorphisms, and in particular single nucleotide polymorphisms (SNP), within the genome, more particularly on chromosome 1 to 29, of rainbow trout which are associated with late onset of sexual maturation. Specific details of single nucleotide polymorphisms of the invention are provided in Table 1 below. The respective nucleotide sequences including the SNP (at position 31) are shown in Table 2. 
     The polymorphisms of the invention can be present in either of two forms, i.e., the polymorphisms have two alleles. 
     One allele can be characterized as being an allele conferring late onset of sexual maturation. This means that a rainbow trout having such allele at the position of a polymorphism detailed herein shows late onset of sexual maturation. This allele is herein denoted “late maturation allele”. The respective late maturation allele for each of the single nucleotide polymorphism of the invention is specified in Table 1 below. A late maturation allele according to the present invention may therefore be used to predict late onset of sexual maturation in a rainbow trout. A late maturation allele according to the present invention may also be used to select a rainbow trout having late onset of sexual maturation. 
     The other allele can be characterized as being an allele that does not confer late onset of sexual maturation. Such allele is herein denoted “normal maturation allele”. 
     Rainbow trout are diploid, in some case triploid organisms, and thus possess at least two copies of the polymorphisms of the invention (one copy to be found on each copy of the chromosome). 
     As demonstrated herein, if at least one allele of a polymorphism, and more particularly of a SNP, is the respective late maturation allele, then the rainbow trout has late onset of sexual maturation as compared to a rainbow trout wherein both alleles are normal maturation alleles (i.e. such rainbow trout being homozygous for the normal maturation allele). In a great number of cases the onset of sexual maturation is even further delayed if both alleles of a polymorphism, and more particularly of a SNP, are the respective late maturation alleles (such rainbow trout being homozygous for the late maturation allele). Such further increase is, for example, seen for SNP #140 ( FIGS. 2 a  and 2 b   ) which is the most statistically significant SNP associated with late onset of sexual maturation (see table 3). 
     A polymorphism of the invention may be any of several polymorphisms associated with late onset of sexual maturation of a rainbow trout. Particularly, a polymorphism of the invention is a polymorphism located on any one of chromosome 1 to 29 of rainbow trout (following the nomenclature of Palti et al. (2011)), i.e. a polymorphism found to be located on any one of chromosome 1 to 29 on the basis of genetic linkage analysis, Fluorescence In Situ Hybridization (FISH) or any other method that assigns DNA polymorphisms to their respective chromosomes. 
     A polymorphism of the invention may be any polymorphism, including single nucleotide polymorphism, located within any of the rainbow trout genomic sequences listed in the column titled “GenBank contig” in Table 1. 
     A polymorphism of the invention may be any polymorphism, including single nucleotide polymorphism, which is in strong linkage disequilibrium (LD) with any one of the SNPs listed in table 1. Here, two polymorphisms are defined to be in strong LD if the square of the correlation coefficient between the two loci (r2, the most commonly used measure of LD) is equal to or larger than 0.5, more preferably equal to or larger than 0.7, even more preferably equal to or larger than 0.8 and most preferably equal to or larger than 0.9. A person who is skilled in the art will know how to estimate r2, as well as what data is required for this estimation. 
     A polymorphism of the invention may be at least one of the single nucleotide polymorphisms listed in Table 1. Therefore, according to certain embodiments, the at least one SNP of the invention is selected from the SNPs listed in Table 1. Each of the SNPs listed in Table 1 is contemplated as being disclosed individually as part of the present invention. 
     
       
         
           
               
             
               
                 TABLE 1 
               
             
            
               
                   
               
               
                 SNPs associated with sexual maturation in Rainbow trout. 
               
            
           
           
               
               
               
               
               
               
               
            
               
                   
                   
                   
                   
                 Position in 
                 Late 
                 Normal 
               
               
                   
                   
                   
                   
                 Genbank 
                 maturation 
                 maturation 
               
               
                 SNP# 
                 Affymetrix ID 
                 Chromosome 
                 GenBank contig 
                 contig 
                 allele 
                 allele 
               
               
                   
               
            
           
           
               
               
               
               
               
               
               
            
               
                 1 
                 Affx-88904935 
                 chr1  
                 CM007935.1 
                 63783597 
                 T 
                 C 
               
               
                 2 
                 Affx-88955402 
                 chr1  
                 CM007935.1 
                 64700514 
                 A 
                 C 
               
               
                 3 
                 Affx-88936832 
                 chr1  
                 CM007935.1 
                 65578255 
                 G 
                 A 
               
               
                 4 
                 Affx-88950378 
                 chr1  
                 CM007935.1 
                 65615419 
                 A 
                 G 
               
               
                 5 
                 Affx-88925365 
                 chr2  
                 CM007936.1 
                 54285214 
                 A 
                 G 
               
               
                 6 
                 Affx-88946858 
                 chr2  
                 CM007936.1 
                 56918053 
                 C 
                 T 
               
               
                 7 
                 Affx-88954034 
                 chr2  
                 CM007936.1 
                 57044148 
                 C 
                 T 
               
               
                 8 
                 Affx-88936668 
                 chr2  
                 CM007936.1 
                 59081958 
                 A 
                 C 
               
               
                 9 
                 Affx-88953569 
                 chr2  
                 CM007936.1 
                 62341749 
                 A 
                 G 
               
               
                 10 
                 Affx-88961158 
                 chr2  
                 CM007936.1 
                 62572758 
                 C 
                 T 
               
               
                 11 
                 Affx-88934762 
                 chr2  
                 CM007936.1 
                 63306724 
                 G 
                 A 
               
               
                 12 
                 Affx-88923797 
                 chr2  
                 CM007936.1 
                 63685704 
                 T 
                 C 
               
               
                 13 
                 Affx-88905095 
                 chr3  
                 CM007937.1 
                 19524784 
                 A 
                 G 
               
               
                 14 
                 Affx-88915123 
                 chr3  
                 CM007937.1 
                 19535483 
                 A 
                 G 
               
               
                 15 
                 Affx-88954577 
                 chr3  
                 CM007937.1 
                 19603901 
                 G 
                 T 
               
               
                 16 
                 Affx-88938133 
                 chr3  
                 CM007937.1 
                 30879389 
                 A 
                 C 
               
               
                 17 
                 Affx-88915067 
                 chr4  
                 CM007938.1 
                  8909969 
                 C 
                 T 
               
               
                 18 
                 Affx-88941840 
                 chr4  
                 CM007938.1 
                 11240214 
                 G 
                 A 
               
               
                 19 
                 Affx-88904592 
                 chr4  
                 CM007938.1 
                 12590767 
                 T 
                 G 
               
               
                 20 
                 Affx-88958567 
                 chr4  
                 CM007938.1 
                 71235287 
                 C 
                 T 
               
               
                 21 
                 Affx-88920423 
                 chr5  
                 CM007939.1 
                 51055810 
                 A 
                 G 
               
               
                 22 
                 Affx-88951600 
                 chr5  
                 CM007939.1 
                 71720042 
                 A 
                 G 
               
               
                 23 
                 Affx-88958428 
                 chr5  
                 CM007939.1 
                 81401808 
                 A 
                 G 
               
               
                 24 
                 Affx-88921734 
                 chr5  
                 CM007939.1 
                 82479766 
                 G 
                 T 
               
               
                 25 
                 Affx-88952785 
                 chr6  
                 CM007940.1 
                  6340153 
                 G 
                 A 
               
               
                 26 
                 Affx-88954186 
                 chr6  
                 CM007940.1 
                 18411308 
                 G 
                 A 
               
               
                 27 
                 Affx-88940319 
                 chr6  
                 CM007940.1 
                 29053396 
                 A 
                 C 
               
               
                 28 
                 Affx-88936711 
                 chr6  
                 CM007940.1 
                 32381726 
                 A 
                 G 
               
               
                 29 
                 Affx-88909681 
                 chr7  
                 CM007941.1 
                 18849968 
                 C 
                 A 
               
               
                 30 
                 Affx-88920716 
                 chr7  
                 CM007941.1 
                 33271630 
                 T 
                 C 
               
               
                 31 
                 Affx-88936725 
                 chr7  
                 CM007941.1 
                 34065749 
                 C 
                 A 
               
               
                 32 
                 Affx-88930546 
                 chr7  
                 CM007941.1 
                 35811286 
                 G 
                 T 
               
               
                 33 
                 Affx-88932380 
                 chr8  
                 CM007942.1 
                 69313777 
                 T 
                 C 
               
               
                 34 
                 Affx-88919018 
                 chr8  
                 CM007942.1 
                 70769104 
                 A 
                 G 
               
               
                 35 
                 Affx-88948691 
                 chr8  
                 CM007942.1 
                 70793135 
                 T 
                 C 
               
               
                 36 
                 Affx-88916802 
                 chr8  
                 CM007942.1 
                 77412834 
                 G 
                 A 
               
               
                 37 
                 Affx-88916833 
                 chr9  
                 CM007943.1 
                 22487940 
                 A 
                 G 
               
               
                 38 
                 Affx-88929636 
                 chr9  
                 CM007943.1 
                 22627473 
                 A 
                 G 
               
               
                 39 
                 Affx-88926137 
                 chr9  
                 CM007943.1 
                 22675057 
                 A 
                 G 
               
               
                 40 
                 Affx-88926394 
                 chr9  
                 CM007943.1 
                 31913561 
                 A 
                 G 
               
               
                 41 
                 Affx-88934825 
                 chr10 
                 CM007944.1 
                 31683283 
                 G 
                 A 
               
               
                 42 
                 Affx-88904537 
                 chr10 
                 CM007944.1 
                 33539443 
                 T 
                 C 
               
               
                 43 
                 Affx-88959739 
                 chr10 
                 CM007944.1 
                 39056184 
                 T 
                 G 
               
               
                 44 
                 Affx-88956881 
                 chr11 
                 CM007945.1 
                  3987284 
                 G 
                 T 
               
               
                 45 
                 Affx-88951001 
                 chr11 
                 CM007945.1 
                 49414158 
                 G 
                 T 
               
               
                 46 
                 Affx-88954986 
                 chr11 
                 CM007945.1 
                 57541195 
                 T 
                 C 
               
               
                 47 
                 Affx-88916907 
                 chr11 
                 CM007945.1 
                 67172627 
                 C 
                 T 
               
               
                 48 
                 Affx-88947853 
                 chr12 
                 CM007946.1 
                 27799868 
                 C 
                 A 
               
               
                 49 
                 Affx-88939958 
                 chr12 
                 CM007946.1 
                 57549580 
                 G 
                 A 
               
               
                 50 
                 Affx-88911641 
                 chr12 
                 CM007946.1 
                 86816745 
                 T 
                 C 
               
               
                 51 
                 Affx-88957555 
                 chr13 
                 CM007947.1 
                 12693726 
                 G 
                 T 
               
               
                 52 
                 Affx-88942222 
                 chr13 
                 CM007947.1 
                 32421268 
                 G 
                 A 
               
               
                 53 
                 Affx-88904308 
                 chr13 
                 CM007947.1 
                 45827883 
                 C 
                 T 
               
               
                 54 
                 Affx-88931117 
                 chr14 
                 CM007948.1 
                 19467149 
                 C 
                 T 
               
               
                 55 
                 Affx-88959989 
                 chr14 
                 CM007948.1 
                 21894963 
                 C 
                 T 
               
               
                 56 
                 Affx-88918911 
                 chr14 
                 CM007948.1 
                 24115228 
                 T 
                 G 
               
               
                 57 
                 Affx-88922027 
                 chr14 
                 CM007948.1 
                 24920208 
                 G 
                 A 
               
               
                 58 
                 Affx-88921698 
                 chr15 
                 CM007949.1 
                 24569160 
                 T 
                 C 
               
               
                 59 
                 Affx-88925753 
                 chr15 
                 CM007949.1 
                 27726385 
                 T 
                 C 
               
               
                 60 
                 Affx-88956336 
                 chr15 
                 CM007949.1 
                 27801757 
                 C 
                 A 
               
               
                 61 
                 Affx-88961133 
                 chr15 
                 CM007949.1 
                 54519499 
                 G 
                 T 
               
               
                 62 
                 Affx-88921645 
                 chr16 
                 CM007950.1 
                 36765420 
                 T 
                 G 
               
               
                 63 
                 Affx-88949977 
                 chr16 
                 CM007950.1 
                 48753210 
                 A 
                 G 
               
               
                 64 
                 Affx-88957804 
                 chr16 
                 CM007950.1 
                 49749575 
                 A 
                 G 
               
               
                 65 
                 Affx-88913266 
                 chr17 
                 CM007951.1 
                 38785093 
                 A 
                 G 
               
               
                 66 
                 Affx-88939862 
                 chr17 
                 CM007951.1 
                 38831122 
                 G 
                 T 
               
               
                 67 
                 Affx-88915748 
                 chr18 
                 CM007952.1 
                 21596568 
                 T 
                 C 
               
               
                 68 
                 Affx-88922933 
                 chr18 
                 CM007952.1 
                 22420190 
                 G 
                 A 
               
               
                 69 
                 Affx-88915837 
                 chr18 
                 CM007952.1 
                 27494999 
                 C 
                 T 
               
               
                 70 
                 Affx-88929378 
                 chr19 
                 CM007953.1 
                 30636790 
                 C 
                 A 
               
               
                 71 
                 Affx-88961378 
                 chr19 
                 CM007953.1 
                 34580702 
                 A 
                 C 
               
               
                 72 
                 Affx-88953889 
                 chr19 
                 CM007953.1 
                 36847192 
                 C 
                 T 
               
               
                 73 
                 Affx-88922131 
                 chr19 
                 CM007953.1 
                 36901034 
                 G 
                 A 
               
               
                 74 
                 Affx-88958970 
                 chr20 
                 CM007954.1 
                  2000433 
                 G 
                 T 
               
               
                 75 
                 Affx-88910072 
                 chr20 
                 CM007954.1 
                 10825499 
                 T 
                 G 
               
               
                 76 
                 Affx-88933134 
                 chr20 
                 CM007954.1 
                 15246572 
                 A 
                 C 
               
               
                 77 
                 Affx-88939049 
                 chr21 
                 CM007955.1 
                 12015696 
                 T 
                 G 
               
               
                 78 
                 Affx-88950195 
                 chr21 
                 CM007955.1 
                 14532693 
                 T 
                 C 
               
               
                 79 
                 Affx-88933377 
                 chr21 
                 CM007955.1 
                 14717955 
                 G 
                 A 
               
               
                 80 
                 Affx-88909724 
                 chr21 
                 CM007955.1 
                 34227983 
                 A 
                 G 
               
               
                 81 
                 Affx-88917486 
                 chr22 
                 CM007956.1 
                  4433427 
                 A 
                 G 
               
               
                 82 
                 Affx-88941752 
                 chr22 
                 CM007956.1 
                 23757569 
                 G 
                 T 
               
               
                 83 
                 Affx-88942920 
                 chr22 
                 CM007956.1 
                 24124923 
                 A 
                 T 
               
               
                 84 
                 Affx-88945639 
                 chr23 
                 CM007957.1 
                 46442301 
                 A 
                 G 
               
               
                 85 
                 Affx-88905397 
                 chr23 
                 CM007957.1 
                 46444905 
                 T 
                 G 
               
               
                 86 
                 Affx-88949049 
                 chr23 
                 CM007957.1 
                 47383604 
                 A 
                 G 
               
               
                 87 
                 Affx-88923532 
                 chr23 
                 CM007957.1 
                 48284540 
                 C 
                 T 
               
               
                 88 
                 Affx-88924387 
                 chr23 
                 CM007957.1 
                 48657925 
                 G 
                 T 
               
               
                 89 
                 Affx-88905064 
                 chr23 
                 CM007957.1 
                 48711098 
                 A 
                 G 
               
               
                 90 
                 Affx-88957818 
                 chr24 
                 CM007958.1 
                 20952836 
                 A 
                 G 
               
               
                 91 
                 Affx-88945529 
                 chr24 
                 CM007958.1 
                 25019945 
                 T 
                 C 
               
               
                 92 
                 Affx-88940262 
                 chr24 
                 CM007958.1 
                 25020252 
                 T 
                 C 
               
               
                 93 
                 Affx-88928194 
                 chr25 
                 CM007959.1 
                 10849854 
                 C 
                 A 
               
               
                 94 
                 Affx-88944575 
                 chr25 
                 CM007959.1 
                 14555851 
                 T 
                 C 
               
               
                 95 
                 Affx-88911550 
                 chr25 
                 CM007959.1 
                 14574366 
                 A 
                 G 
               
               
                 96 
                 Affx-88940618 
                 chr25 
                 CM007959.1 
                 15498993 
                 C 
                 T 
               
               
                 97 
                 Affx-88918400 
                 chr25 
                 CM007959.1 
                 47014119 
                 T 
                 G 
               
               
                 98 
                 Affx-88926265 
                 chr25 
                 CM007959.1 
                 47014758 
                 G 
                 A 
               
               
                 99 
                 Affx-88945363 
                 chr25 
                 CM007959.1 
                 47015074 
                 A 
                 G 
               
               
                 100 
                 Affx-88914956 
                 chr25 
                 CM007959.1 
                 48229209 
                 G 
                 A 
               
               
                 101 
                 Affx-88959143 
                 chr25 
                 CM007959.1 
                 48397292 
                 C 
                 T 
               
               
                 102 
                 Affx-88907320 
                 chr26 
                 CM007960.1 
                 10225818 
                 T 
                 G 
               
               
                 103 
                 Affx-88950639 
                 chr26 
                 CM007960.1 
                 10495031 
                 A 
                 C 
               
               
                 104 
                 Affx-88951915 
                 chr26 
                 CM007960.1 
                 12901743 
                 G 
                 A 
               
               
                 105 
                 Affx-88939677 
                 chr26 
                 CM007960.1 
                 26154364 
                 C 
                 T 
               
               
                 106 
                 Affx-88921625 
                 chr27 
                 CM007961.1 
                  7278740 
                 G 
                 A 
               
               
                 107 
                 Affx-88918503 
                 chr27 
                 CM007961.1 
                  7764626 
                 C 
                 T 
               
               
                 108 
                 Affx-88958017 
                 chr27 
                 CM007961.1 
                  7820269 
                 C 
                 A 
               
               
                 109 
                 Affx-88911327 
                 chr27 
                 CM007961.1 
                 10784017 
                 C 
                 T 
               
               
                 110 
                 Affx-88950248 
                 chr28 
                 CM007962.1 
                  9037911 
                 C 
                 A 
               
               
                 111 
                 Affx-88932371 
                 chr28 
                 CM007962.1 
                  9072612 
                 C 
                 T 
               
               
                 112 
                 Affx-88923416 
                 chr28 
                 CM007962.1 
                  9082601 
                 C 
                 T 
               
               
                 113 
                 Affx-88955399 
                 chr28 
                 CM007962.1 
                  9102905 
                 T 
                 C 
               
               
                 114 
                 Affx-88908191 
                 chr28 
                 CM007962.1 
                  9446099 
                 T 
                 C 
               
               
                 115 
                 Affx-88923808 
                 chr28 
                 CM007962.1 
                  9632653 
                 A 
                 G 
               
               
                 116 
                 Affx-88908821 
                 chr28 
                 CM007962.1 
                  9909392 
                 G 
                 A 
               
               
                 117 
                 Affx-88955530 
                 chr28 
                 CM007962.1 
                 10202248 
                 A 
                 G 
               
               
                 118 
                 Affx-88941807 
                 chr28 
                 CM007962.1 
                 10237515 
                 A 
                 G 
               
               
                 119 
                 Affx-88904869 
                 chr28 
                 CM007962.1 
                 10252908 
                 T 
                 G 
               
               
                 120 
                 Affx-88912748 
                 chr28 
                 CM007962.1 
                 10378833 
                 C 
                 A 
               
               
                 121 
                 Affx-88924717 
                 chr28 
                 CM007962.1 
                 10406672 
                 A 
                 C 
               
               
                 122 
                 Affx-88946195 
                 chr28 
                 CM007962.1 
                 10407234 
                 C 
                 A 
               
               
                 123 
                 Affx-88931804 
                 chr28 
                 CM007962.1 
                 10500800 
                 G 
                 A 
               
               
                 124 
                 Affx-88929559 
                 chr28 
                 CM007962.1 
                 10502719 
                 T 
                 G 
               
               
                 125 
                 Affx-88905073 
                 chr28 
                 CM007962.1 
                 10564775 
                 G 
                 T 
               
               
                 126 
                 Affx-88947012 
                 chr28 
                 CM007962.1 
                 10593406 
                 G 
                 A 
               
               
                 127 
                 Affx-88939287 
                 chr28 
                 CM007962.1 
                 10605800 
                 A 
                 G 
               
               
                 128 
                 Affx-88947787 
                 chr28 
                 CM007962.1 
                 11093692 
                 C 
                 T 
               
               
                 129 
                 Affx-88932913 
                 chr28 
                 CM007962.1 
                 11096202 
                 C 
                 T 
               
               
                 130 
                 Affx-88952141 
                 chr28 
                 CM007962.1 
                 11139401 
                 G 
                 T 
               
               
                 131 
                 Affx-88913454 
                 chr28 
                 CM007962.1 
                 11148320 
                 T 
                 C 
               
               
                 132 
                 Affx-88959734 
                 chr28 
                 CM007962.1 
                 11200364 
                 T 
                 G 
               
               
                 133 
                 Affx-88915384 
                 chr28 
                 CM007962.1 
                 11251982 
                 T 
                 G 
               
               
                 134 
                 Affx-88927547 
                 chr28 
                 CM007962.1 
                 11382381 
                 A 
                 C 
               
               
                 135 
                 Affx-88946147 
                 chr28 
                 CM007962.1 
                 11414120 
                 G 
                 A 
               
               
                 136 
                 Affx-88938817 
                 chr28 
                 CM007962.1 
                 11414342 
                 C 
                 T 
               
               
                 137 
                 Affx-88953900 
                 chr28 
                 CM007962.1 
                 11414399 
                 A 
                 G 
               
               
                 138 
                 Affx-88947402 
                 chr28 
                 CM007962.1 
                 11487704 
                 C 
                 T 
               
               
                 139 
                 Affx-88918233 
                 chr28 
                 CM007962.1 
                 11523747 
                 C 
                 T 
               
               
                 140 
                 Affx-88940276 
                 chr28 
                 CM007962.1 
                 11537736 
                 T 
                 C 
               
               
                 141 
                 Affx-88913684 
                 chr28 
                 CM007962.1 
                 11575843 
                 G 
                 T 
               
               
                 142 
                 Affx-88921970 
                 chr28 
                 CM007962.1 
                 11667915 
                 A 
                 G 
               
               
                 143 
                 Affx-88945246 
                 chr28 
                 CM007962.1 
                 11704242 
                 A 
                 G 
               
               
                 144 
                 Affx-88958541 
                 chr28 
                 CM007962.1 
                 11745469 
                 T 
                 C 
               
               
                 145 
                 Affx-88949728 
                 chr28 
                 CM007962.1 
                 11771654 
                 A 
                 G 
               
               
                 146 
                 Affx-88961021 
                 chr28 
                 CM007962.1 
                 12063041 
                 T 
                 C 
               
               
                 147 
                 Affx-88907679 
                 chr28 
                 CM007962.1 
                 12252318 
                 A 
                 G 
               
               
                 148 
                 Affx-88951316 
                 chr28 
                 CM007962.1 
                 12259307 
                 G 
                 T 
               
               
                 149 
                 Affx-88930743 
                 chr28 
                 CM007962.1 
                 12593967 
                 G 
                 T 
               
               
                 150 
                 Affx-88941593 
                 chr28 
                 CM007962.1 
                 12700154 
                 T 
                 C 
               
               
                 151 
                 Affx-88948678 
                 chr28 
                 CM007962.1 
                 12735308 
                 C 
                 T 
               
               
                 152 
                 Affx-88912846 
                 chr28 
                 CM007962.1 
                 12847942 
                 A 
                 C 
               
               
                 153 
                 Affx-88915764 
                 chr28 
                 CM007962.1 
                 12982469 
                 C 
                 T 
               
               
                 154 
                 Affx-88905807 
                 chr28 
                 CM007962.1 
                 13010185 
                 C 
                 A 
               
               
                 155 
                 Affx-88948195 
                 chr28 
                 CM007962.1 
                 13365774 
                 A 
                 C 
               
               
                 156 
                 Affx-88937868 
                 chr28 
                 CM007962.1 
                 13877922 
                 A 
                 G 
               
               
                 157 
                 Affx-88929314 
                 chr28 
                 CM007962.1 
                 13927035 
                 A 
                 G 
               
               
                 158 
                 Affx-88919686 
                 chr28 
                 CM007962.1 
                 14120232 
                 G 
                 T 
               
               
                 159 
                 Affx-88948530 
                 chr28 
                 CM007962.1 
                 14551487 
                 C 
                 T 
               
               
                 160 
                 Affx-88911474 
                 chr28 
                 CM007962.1 
                 14570553 
                 C 
                 T 
               
               
                 161 
                 Affx-88944645 
                 chr28 
                 CM007962.1 
                 18046487 
                 C 
                 T 
               
               
                 162 
                 Affx-88952966 
                 chr29 
                 CM007963.1 
                 12570007 
                 C 
                 T 
               
               
                 163 
                 Affx-88904400 
                 chr29 
                 CM007963.1 
                 17738627 
                 T 
                 G 
               
               
                 164 
                 Affx-88914995 
                 chr29 
                 CM007963.1 
                 38728561 
                 T 
                 C 
               
               
                   
               
               
                 A = Adenine, G = Guanine; C = Cytosine, T = Thymine. 
               
               
                 Affymetrix ID is a unique identifier given to each SNP by Affymetrix, the provider of a commercial genotyping assay which incorporates many of the SNPs listed in the table; the Affymetrix ID serves as a link to further details pertaining to the SNPs, provided in a file which can be downloaded from http://www.affymetrix.com/estore/. 
               
               
                 Chromosome indicates on which chromosome the SNPs are situated. 
               
               
                 GenBank contig is the name of a GenBank DNA contig (a genome sequence from rainbow trout) wherein the SNP resides, and the position is the position of the SNP within this contig. 
               
               
                 SNPs in bold are of particular interest. All of these SNPs are located within a genomic region of 1.9 megabases on chromosome 28 (FIG. 1b), have a p-value lower than 3.12e−7 (FIG. 1b) and show varying degree of elevated squared correlations to each other (FIG. 3). 
               
            
           
         
       
     
     
       
         
           
               
             
               
                 TABLE 2 
               
             
            
               
                   
               
               
                 Nucleotide sequence for each SNP including the polymorphic site. The polymorphic site and the 
               
               
                 corresponding allelic variants are indicated in brackets. The column labeled ″Nucleotide sequence 
               
               
                 including SNP site″ provides a reference nucleotide sequence for identification of the SNP within 
               
               
                 the genome of a rainbow trout. The sequences SEQ ID NO: 1 to 164 are each polymorphic sequences 
               
               
                 including a polymorphic site. SNPs in bold are of particular interest. All of these SNPs are 
               
               
                 located within a genomic region of 1.9 megabases on chromosome 28 (FIG. 1b), have a p-value lower 
               
               
                 than 3.12e-7 (FIG. 1b) and show varying degree of elevated squared correlations to each other 
               
               
                 (FIG. 3). 
               
            
           
           
               
               
               
               
               
            
               
                   
                 SEQ 
                   
                 Late 
                 Normal 
               
               
                   
                 ID 
                   
                 maturation 
                 maturation 
               
               
                 SNP# 
                 NO 
                 Nucleotide sequence including SNP site 
                 allele 
                 allele 
               
               
                   
               
            
           
           
               
               
               
               
               
            
               
                 1 
                 1 
                 AGTATATGATGATTCAATGAGATACAGTAC[C/T]TTACACTTTATTGCCCATTTCCATGAAAAT 
                 T 
                 C 
               
               
                   
               
               
                 2 
                 2 
                 TCAGTCCATAAGTAATGCAAAGATCAAAGT[A/C]ATTCTACAGAAACCGATTGGGCAGACTAAC 
                 A 
                 C 
               
               
                   
               
               
                 3 
                 3 
                 AGTACAAACAGAGATGTGTTATGTTAGACA[A/G]CTGAAGTGAACCGCTACACCTGCTTGGTGC 
                 G 
                 A 
               
               
                   
               
               
                 4 
                 4 
                 GAGTGCAGTGGATAGAGACAGCTCCTCAGT[A/G]CATAAAGGCCCACCTGTCCTGGGGGAAGAT 
                 A 
                 G 
               
               
                   
               
               
                 5 
                 5 
                 TACTATAATCGGGGACAGTGACATGCATCG[A/G]CCCACAAAGTTTTTAAGACTGCAGTTATGT 
                 A 
                 G 
               
               
                   
               
               
                 6 
                 6 
                 GGAACCAGTCACTGTCTCACTACATTTTCA[C/T]GTGGCAGTTTTGTCTTCCACCGTGCAAGCC 
                 C 
                 T 
               
               
                   
               
               
                 7 
                 7 
                 TTGGTTCTTGCAACTCTATAGCTCTGGGTC[C/T]TTCCCTTACCTCGGCACGGCAGCCAGTCAG 
                 C 
                 T 
               
               
                   
               
               
                 8 
                 8 
                 CTGTATAAAGTTGTTACTGCAGGTACAGGC[A/C]GTGATGCTGAGACTCTTCTGCCAAGACACC 
                 A 
                 C 
               
               
                   
               
               
                 9 
                 9 
                 ATCGTACTGAGCTTGTTGTCATTGCAGGCA[A/G]TCTCAACTCTGTGCATTACACTGAAGACTT 
                 A 
                 G 
               
               
                   
               
               
                 10 
                 10 
                 AGACAGATCATCAAAAGCTTTTATTCTGAT[C/T]AAGTTCAGTAGTTTGTTTAGGACACTGAAA 
                 C 
                 T 
               
               
                   
               
               
                 11 
                 11 
                 TGACCTGCAGCTATGCACCATAATCTAGCA[A/G]GTTCATTTGAACACCCTTTGAAAAGGTAAT 
                 G 
                 A 
               
               
                   
               
               
                 12 
                 12 
                 TCTATACTACGGTCTATTGCCTATTTTTAA[C/T]GTATCTTTAATTTCGTATCCCAGTTATTAG 
                 T 
                 C 
               
               
                   
               
               
                 13 
                 13 
                 TTACCATACTAACTTGTAGGGCTGAGCAAT[A/G]TATTTTGAATACAGGCACAGAGCCACATAC 
                 A 
                 G 
               
               
                   
               
               
                 14 
                 14 
                 CTACACTAAAATGCAATTTGATCTGGACAG[A/G]TTGTCTGTTATGCTATTGCAGTGTTATGAC 
                 A 
                 G 
               
               
                   
               
               
                 15 
                 15 
                 CACTACCTGATGCAGTCCCAGTTTGTGATT[G/T]TTATCTGCAGAAACTCAAATATAAATTCCA 
                 G 
                 T 
               
               
                   
               
               
                 16 
                 16 
                 GACGCTGCTGTTCCTGCTGCTGCCACCACA[A/C]CCATTTCCTCTCGTCATGAGCAAAAGCTAT 
                 A 
                 C 
               
               
                   
               
               
                 17 
                 17 
                 AGGTACCCTGCACTACATTCCTAGCGCGAC[C/T]AGAGGATGGTATAGAAAATGTAATGGTATA 
                 C 
                 T 
               
               
                   
               
               
                 18 
                 18 
                 TCTCTGAGTGAGATCAAGAACGGTTCGGTT[A/G]TCTACGACTGTTGGGGCCACTTCATAGAAC 
                 G 
                 A 
               
               
                   
               
               
                 19 
                 19 
                 TGCAGATATGAACAGCTTGAGTAAAAAGAT[G/T]ATGTTACCCACTGAACATCAATGAACAAAT 
                 T 
                 G 
               
               
                   
               
               
                 20 
                 20 
                 CAAAGGGACTTATCTTCTCCCAAAAGACAA[C/T]GGGCCGATCATTTAACGAATCTTCTCTTGA 
                 C 
                 T 
               
               
                   
               
               
                 21 
                 21 
                 CTGCGACATGTTTTGAGTTAGCGTAATTTC[A/G]TACTAAAGATTGGAGAAGTGTGCCTAATTA 
                 A 
                 G 
               
               
                   
               
               
                 22 
                 22 
                 TTCGCCTTCACCCTGAGTTAGGACGGCTCC[A/G]AGCCCCATATTTGAGGCGTCTACCTGCACA 
                 A 
                 G 
               
               
                   
               
               
                 23 
                 23 
                 AATTGACTAACAATATTGTCTAACAAGTGC[A/G]GTATAAATAAATCCATCCTTCTCATCCTCC 
                 A 
                 G 
               
               
                   
               
               
                 24 
                 24 
                 GTATGGTTTTGAAGAGTACAACTGTGTGAG[G/T]GTGGATTGAACAAAATAGTATCTTAAACAC 
                 G 
                 T 
               
               
                   
               
               
                 25 
                 25 
                 TAAACAGCAGTTGAACGAGCTGGGCAGCGA[A/G]TCGGCCAAGATCAAGGCCATGGGCATCACC 
                 G 
                 A 
               
               
                   
               
               
                 26 
                 26 
                 GATGCGTCCCTCCAACACAGTGCATCTGCT[A/G]GTTTTGTGTGAGGACCACAGAGCCGGCATG 
                 G 
                 A 
               
               
                   
               
               
                 27 
                 27 
                 GTGCGGCAGGCGGCAGTGGTGGACAACTTC[A/C]TGTCCCAGCAAGAGAAGAAGCAGAAACACC 
                 A 
                 C 
               
               
                   
               
               
                 28 
                 28 
                 GCCTGCAGATGTTCCTCACGTGATGTGATG[A/G]CCTTTTAACTGGGCGTCCTTTGAATATAAG 
                 A 
                 G 
               
               
                   
               
               
                 29 
                 29 
                 ACGTGCCTTTTGATGGTTATTACTAGACCG[A/C]TTATTGTACCTGCCCTATTGATCAACCGGA 
                 C 
                 A 
               
               
                   
               
               
                 30 
                 30 
                 GACATAGGGACGTTTCCACATGAAGTGAAT[C/T]GGAAAAGCATACACTTACATGACTTTCAAT 
                 T 
                 C 
               
               
                   
               
               
                 31 
                 31 
                 ATCAGATTCTCCAAAAAGGTCCAGGGGAAA[A/C]AGTTTGCTGCTTTTGTTGGATATTTTTACA 
                 C 
                 A 
               
               
                   
               
               
                 32 
                 32 
                 ATGAGTGATATTACAGTTTGTCCTTCAGAT[G/T]AAACAATTGAGGAGCCAACTATGTGTAATC 
                 G 
                 T 
               
               
                   
               
               
                 33 
                 33 
                 GCATAAAGTTGATACAATTCACACAAAGTC[C/T]TTGTCGGGGGACTCCAATCCTCTGTGTTTC 
                 T 
                 C 
               
               
                   
               
               
                 34 
                 34 
                 ACACACTGCAGAGTAAACAGCAAACACTGA[A/G]AAAGCTGCACCCAGACTGGCATTCACACAC 
                 A 
                 G 
               
               
                   
               
               
                 35 
                 35 
                 TGCTCTGAGAAGGGGGTTCTGATTTCTGTC[C/T]ACAGGGGGCTCCCTCCTGCCTATCCAACGA 
                 T 
                 C 
               
               
                   
               
               
                 36 
                 36 
                 CTGTAGTATCTGGAAGCCTAGGCCCAGTAG[A/G]ATAGTGTTCTTATTCCCTATGGAGCGCATC 
                 G 
                 A 
               
               
                   
               
               
                 37 
                 37 
                 ATGAGATATGAATTAGACCTAAAGGCCTCA[A/G]CATGCTTCAGTTTTGCTGGTGCCTAGCTTG 
                 A 
                 G 
               
               
                   
               
               
                 38 
                 38 
                 CTTGAGTATGTGTGTTCCATTAGAGTGTAT[A/G]CAGAGTGTGTCGCTCGCACAGACAGATTGG 
                 A 
                 G 
               
               
                   
               
               
                 39 
                 39 
                 TGTGATTCGATTTTGTAGCTACAACAAGCG[A/G]TGGCTCAAAACCAACCTACACATTTTCAGT 
                 A 
                 G 
               
               
                   
               
               
                 40 
                 40 
                 ACAAATACAACGGGATGGAGTGCTCGTCTC[A/G]TCACATAATGTTCCCTGGCAGTGCTTCGTT 
                 A 
                 G 
               
               
                   
               
               
                 41 
                 41 
                 GTTGACCTTCTTGTTGGTGGTGAAATCGCA[A/G]TCTGTGCACTGGTACTTCTTCTTGAAGATG 
                 G 
                 A 
               
               
                   
               
               
                 42 
                 42 
                 GTCTTAACTGCAAATGTGAGCTTAAATCGG[C/T]TTTCAGCTCCCATGTATTACGGTATTCAAA 
                 T 
                 C 
               
               
                   
               
               
                 43 
                 43 
                 GGGAATGGATTGTAGCAGTGAAGGAGGAGA[G/T]TTATTGGTTTAGAGCTACACCAAGGAGCAC 
                 T 
                 G 
               
               
                   
               
               
                 44 
                 44 
                 AACAGTAAATAGAGCCTGAGATGACTCAAA[G/T]GTCTGATACATGGAGGCAGACTCCTTTCCA 
                 G 
                 T 
               
               
                   
               
               
                 45 
                 45 
                 AAGAACTGTCTGACCGGAACCACTCACACA[G/T]AAAACAGAGAGGCCCGAAGAGATACTGTTG 
                 G 
                 T 
               
               
                   
               
               
                 46 
                 46 
                 TCGTAGCATGTCTGTGTTCCTGGTGAAGAA[C/T]CTTCCAGCGGGAGTGACGGTGGAGAACCTG 
                 T 
                 C 
               
               
                   
               
               
                 47 
                 47 
                 TCACAAAATATGTATACAAGTGTAACCATG[C/T]TGGTTTTGTGGTAATATGTCATGTGTATCT 
                 C 
                 T 
               
               
                   
               
               
                 48 
                 48 
                 CGGTAAAGCCAATGGGAGCGTCCTTAACCT[A/C]AATCAGCCGGGCTACAATCACCACAACCTC 
                 C 
                 A 
               
               
                   
               
               
                 49 
                 49 
                 ATATTAGTAATCTCAAGCCATATTCATACA[A/G]TTTTGTTGAATAGGAAATACGTCATATAAT 
                 G 
                 A 
               
               
                   
               
               
                 50 
                 50 
                 GAATGGACAATGAAGTGGTATTCATTTCTA[C/T]GGGGATATTCACGGTTCTAAAAATAATGAC 
                 T 
                 C 
               
               
                   
               
               
                 51 
                 51 
                 TCTGTCAACGGTCTTCCAAAACAGTATGAC[G/T]GAATCAACAGCAGGGCTGAAACTGACTAAC 
                 G 
                 T 
               
               
                   
               
               
                 52 
                 52 
                 AAAGAAACACGTCACATTTCTAATGATCCA[A/G]CAATCACACTTGGAGGTGTCCATATCCCAC 
                 G 
                 A 
               
               
                   
               
               
                 53 
                 53 
                 TTTCATGGCAATAAATAGAAGTTGGAGAAC[C/T]CTGAGAATGTTGGAGAACCCTGAGAATGCT 
                 C 
                 T 
               
               
                   
               
               
                 54 
                 54 
                 AAAGCCATAGATAGGATATAGGCCTACCTC[C/T]GACACAGAACTGACTGAGTGTCAGTCAACT 
                 C 
                 T 
               
               
                   
               
               
                 55 
                 55 
                 CTTTAAAGTCCCTTGTCCTACATATAGTCT[C/T]TAGCTTGTAAAAAGCCCACAATACAGAGTT 
                 C 
                 T 
               
               
                   
               
               
                 56 
                 56 
                 ATTGCAGTCTCCATTCTGGCTCTGTATCCT[G/T]TTCTCACGGAAGCCACGGTGATTTTTATAG 
                 T 
                 G 
               
               
                   
               
               
                 57 
                 57 
                 GCATGGCTTTCTGTGGCTATATTTGGGGTT[A/G]TCTCAGTCTCCAAAAATCTCTCTGAAATAA 
                 G 
                 A 
               
               
                   
               
               
                 58 
                 58 
                 CATAGTTTATTGTTGCGGACAATAACTTGC[C/T]CTCACATTAAAACTATAGTCTCTCTTACGT 
                 T 
                 C 
               
               
                   
               
               
                 59 
                 59 
                 CATTTTGGATGCATCTGAAGTCATCAGTCG[C/T]CGTATCAACAAAGGGCTTTTTATTGACTGT 
                 T 
                 C 
               
               
                   
               
               
                 60 
                 60 
                 AACTCGGTATTAGGGAAGCAAGCAGGACAA[A/C]ATGGGTTTGGATGCGGGTGAACTAACGTGA 
                 C 
                 A 
               
               
                   
               
               
                 61 
                 61 
                 ACAATGATTACCTTCAAAAAATAATGAATA[G/T]ATTATGTTTTTGCATTTGTAACTGAGCTCA 
                 G 
                 T 
               
               
                   
               
               
                 62 
                 62 
                 TCTCATTGACACCCACTGTGCTCTGTAAAT[G/T]ACTTGATTGTTTGAAGTAAGGCATACTAGG 
                 T 
                 G 
               
               
                   
               
               
                 63 
                 63 
                 GAGAGTGGTAATATCAGCAGCAGAAGCAAC[A/G]CGGTTTCCTGCTGTCCTTACTGTCGAGAGA 
                 A 
                 G 
               
               
                   
               
               
                 64 
                 64 
                 CACCACAAACCCTAGACGTATTATGAATAG[A/G]ATAAAATTTAGAGTAGGTCTCAAATTTAGG 
                 A 
                 G 
               
               
                   
               
               
                 65 
                 65 
                 ACAGGTGGTGGTAACAGAATATTCATTAAG[A/G]TTGCTACTGGCTTTTCAACTAATTCTTAGT 
                 A 
                 G 
               
               
                   
               
               
                 66 
                 66 
                 GGAGCATAGAGCAGATGATGAGAAAGGAGA[G/T]GATGTTCTGATTGAACCAACCCCTTTTGAC 
                 G 
                 T 
               
               
                   
               
               
                 67 
                 67 
                 TTTGCTGCCAATGTCATGATTGATTCAATG[C/T]ATTCGGTGCCAGTTTAGAGCTTCGTTACTC 
                 T 
                 C 
               
               
                   
               
               
                 68 
                 68 
                 ATCTCAGGGTTATAAAGGTAGAAAGGCACT[A/G]TGGGTTCAATTTTGGAAAACAAAAGGAGAG 
                 G 
                 A 
               
               
                   
               
               
                 69 
                 69 
                 TTGTTCCACATGCCTGGATGCCACACAGCA[C/T]TAGATTTTCACTGCTGCAGATACCGGTATG 
                 C 
                 T 
               
               
                   
               
               
                 70 
                 70 
                 GACTGAATTGAAATGGTATTGACCCCAAAC[A/C]TGAGAGGGAAGCATCAATACAAAAGGCACA 
                 C 
                 A 
               
               
                   
               
               
                 71 
                 71 
                 CGACACAGCTGCCGTTTGGACTGCAGCCTT[A/C]AACAGTTTGACAGTTTTTGGTGTTTCTCGT 
                 A 
                 C 
               
               
                   
               
               
                 72 
                 72 
                 TGCGGTGGGTGTCTTTGTGGTGTTTCTGGC[C/T]GTCCTCTTCCTCTTCATCAATAAGAAGCTG 
                 C 
                 T 
               
               
                   
               
               
                 73 
                 73 
                 TTGTGATTAAGTAGAGGGTCACTCTGAGTA[A/G]AACAGTGCTTCATTAGACTCAAGCCTAAAA 
                 G 
                 A 
               
               
                   
               
               
                 74 
                 74 
                 TGGATCAACCATTTGTTCCAAATTCCACTT[G/T]AATGATGTAAAAGGTGTCCGCCTGCCTGCG 
                 G 
                 T 
               
               
                   
               
               
                 75 
                 75 
                 ATGTAGCATTATGTCAGACTTTCGTCAAAA[G/T]CAGTTCAAAAGGAATGGGTGCAGTGACCCT 
                 T 
                 G 
               
               
                   
               
               
                 76 
                 76 
                 GTTCCAATGGTACATTTGAGTAAATAACTT[A/C]GTTTTTACAATATTGTTTATCATTTAGTCT 
                 A 
                 C 
               
               
                   
               
               
                 77 
                 77 
                 AGAAAACACTGCCAGTTGGTTACCAACCCG[G/T]CCCATGTCCAGCTCAAAAATATGTTTTGTC 
                 T 
                 G 
               
               
                   
               
               
                 78 
                 78 
                 TGTTGTCATCCTCGTCAGAGCTGTGGACTT[C/T]CTCCTCAGAGTGAGCATCATCCAATCCATG 
                 T 
                 C 
               
               
                   
               
               
                 79 
                 79 
                 CATATACTGTTTTTATAAACTCACTGTGAT[A/G]TCCAGCTTCTGCAGAGCAAAAAGCTGGTGA 
                 G 
                 A 
               
               
                   
               
               
                 80 
                 80 
                 AAACCGTCTTTTAAGTGTTGATGACATCAG[A/G]GAGAGAATTCATTAATGATCAGAGGTACTT 
                 A 
                 G 
               
               
                   
               
               
                 81 
                 81 
                 AAGTCATTTTGGCTGTCCTGACAAGCTTTG[A/G]GTTTTAATTTTGAGGATGCCTTGGAGGACT 
                 A 
                 G 
               
               
                   
               
               
                 82 
                 82 
                 GGAATGATATTAAAGGAATTGTTGTGAGTT[G/T]TAAATGTGTTTCTTAATATGTGTATGTGTC 
                 G 
                 T 
               
               
                   
               
               
                 83 
                 83 
                 CACCTACGGATGGTACACACAATTATGACA[A/T]ACTTCTCAACTATCATTACCTCAGCCGCTA 
                 A 
                 T 
               
               
                   
               
               
                 84 
                 84 
                 CTTTCTAGCATAGCGTCGCCCACCTGCAGC[A/G]AAGCCGTCAAATATCCTGTTCTTCCTCAGG 
                 A 
                 G 
               
               
                   
               
               
                 85 
                 85 
                 AAAAGATTAATTGAGAAATAGCCATGGTAT[G/T]CTGATTTATTACTGACGTAACGGTCTCCTC 
                 T 
                 G 
               
               
                   
               
               
                 86 
                 86 
                 TAAGGCGTAGTGGTGCACTAATGGGGGAAC[A/G]CGCACTACATCTTGGACCAGAGATAGAGGG 
                 A 
                 G 
               
               
                   
               
               
                 87 
                 87 
                 AGGTGTGACTGACTGACTGAAGTGTAGGTG[C/T]GAGTGAGTAACTGAATAAGAAGGTACTGTA 
                 C 
                 T 
               
               
                   
               
               
                 88 
                 88 
                 CTGTTGTGAAGGTAACTCACACGTATGTAT[G/T]TATGTAGACAAAATACTGTCATGAGTTACA 
                 G 
                 T 
               
               
                   
               
               
                 89 
                 89 
                 ACTAAAAAATACAACAGAACATCTTCCTTT[A/G]GATTCTTAAAACCAACAAACTGGGCTTTAG 
                 A 
                 G 
               
               
                   
               
               
                 90 
                 90 
                 CACATGCTACAATGAGTTACTCTATGAACC[A/G]AACACATTACTGTCAAGTAAATTGTCTCCC 
                 A 
                 G 
               
               
                   
               
               
                 91 
                 91 
                 ACTGGACTATTTGACATGAACCTATTACCT[C/T]ACTCTCTCTCTCCCCTGACCTCCCTACCTC 
                 T 
                 C 
               
               
                   
               
               
                 92 
                 92 
                 GTGTGTGCTATACGACGTCCTCAGCGTAGT[C/T]AAGGAGAAGAAGTACATGGCCCTGGACCCA 
                 T 
                 C 
               
               
                   
               
               
                 93 
                 93 
                 ACAAATCCACGTTGATTTGACTCGCACGGC[A/C]AGGAGTATGACGCTGATAGTAAGATTATTT 
                 C 
                 A 
               
               
                   
               
               
                 94 
                 94 
                 AAGACGGTAAATCACATCAAGTTTAGATGT[C/T]GAGTGTCTGAATACTGAGGAAGTATGTCTT 
                 T 
                 C 
               
               
                   
               
               
                 95 
                 95 
                 GCATCGCTGCAGGTAAAATGAGAACCTTCA[A/G]GTTGTTTTTCATTGGGCTCATCTTCATCAT 
                 A 
                 G 
               
               
                   
               
               
                 96 
                 96 
                 CCTTCCTCACTTTCTACTTCGACTATCGCT[C/T]TCTGTCACACGCTCGTGCACACAGAATGCA 
                 C 
                 T 
               
               
                   
               
               
                 97 
                 97 
                 CATTCAAATAGGCCAAAAGTTTACATACAC[G/T]TTTTTCTCTTTTTGCAAAGCTAATCGCTAA 
                 T 
                 G 
               
               
                   
               
               
                 98 
                 98 
                 GCCTAGAATAAGCATAGATGAGATATCTTC[A/G]TCGTCTCATCAAAACATTTTGGTATTGTGC 
                 G 
                 A 
               
               
                   
               
               
                 99 
                 99 
                 CAGGTTGTCTTTGTTGAGCTTGATGAGGGC[A/G]GAGTAGAGCTCTGAGGCTCGGCCCATGCCC 
                 A 
                 G 
               
               
                   
               
               
                 100 
                 100 
                 TTGATGTAGTCTATTCCTGAAATACAAGGG[A/G]GATGACAATATAAATCATGCAGTTATGTTT 
                 G 
                 A 
               
               
                   
               
               
                 101 
                 101 
                 CAGTACAGACAAGATTTATCGATAGCATTT[C/T]ATGTATGTAACAGGAGACTGTTTATATTTC 
                 C 
                 T 
               
               
                   
               
               
                 102 
                 102 
                 CTGTGCTCGAAGCTAGCAATCACTGATAAA[G/T]CAAATAACAAGCTTTTTCTGAAAACCTTTA 
                 T 
                 G 
               
               
                   
               
               
                 103 
                 103 
                 CATTAAACCACTGATGACAACAAATATGCT[A/C]ATTTCCCCATGTCTCTGTAATGTCATCCTC 
                 A 
                 C 
               
               
                   
               
               
                 104 
                 104 
                 AACATGCAAGGCTATAACTCCCCAGCAGGA[A/G]AGAGAGGGGCTTTTACCACTCCATCAAGTC 
                 G 
                 A 
               
               
                   
               
               
                 105 
                 105 
                 AAGTATGATTCCAAATTGAACTGCAGGTGA[C/T]GCAATGCAACTCATTTCAAGCCTGCACTCA 
                 C 
                 T 
               
               
                   
               
               
                 106 
                 106 
                 GACAGTTGAATTTAACAGGTGAACATTACA[A/G]TTGATATTTTACATTGCATCATGTATGCAA 
                 G 
                 A 
               
               
                   
               
               
                 107 
                 107 
                 ATATCTATACTCCTATTAGTGAAATTGGAA[C/T]ATTATCTAGGCCTACCTACAATCATGGATT 
                 C 
                 T 
               
               
                   
               
               
                 108 
                 108 
                 AGGCCCAATACATTCTGAGATACAGGGTCC[A/C]AAGGTCTAAAACTAATAGTTCAAAACTTTC 
                 C 
                 A 
               
               
                   
               
               
                 109 
                 109 
                 CTCATTATTTCACTTGAAAGGTTTTTCCCC[C/T]GTATCACCTTTCAGTGCACAATTTAAAGGA 
                 C 
                 T 
               
               
                   
               
               
                 110 
                 110 
                 TGTATGTGTGTCTTGAAAGGTCAGGAGCTT[A/C]ACAGAGGGACTGACATTCACTGAGACCAAC 
                 C 
                 A 
               
               
                   
               
               
                 111 
                 111 
                 TTCATTTCAACACCAAGCAGCAGATCAGCT[C/T]AGAACATAAACATAAAACACCACAGTGACC 
                 C 
                 T 
               
               
                   
               
               
                 112 
                 112 
                 CATTTGCAAATGCACACTAGAAAATTTGAG[C/T]CTGGTCTGACCTAGCCCGAGCCCTACCCGA 
                 C 
                 T 
               
               
                   
               
               
                 113 
                 113 
                 ATAGCTACCATTTTGGCCCCATTACGGAAA[C/T]GTTAAGGTTTATGATATCTCCTCTAGTAAT 
                 T 
                 C 
               
               
                   
               
               
                 114 
                 114 
                 GGAGAAAGAACTTAAAAAGATCAAGAACAG[C/T]ACCAATAATGCTGGGACTCCCACCAAAACA 
                 T 
                 C 
               
               
                   
               
               
                 115 
                 115 
                 CTGAGACTGTTAAGGAAGAGGAGGTGAAAA[A/G]GGAAGAAAAGCCCCTACAGTTGATTAGGGG 
                 A 
                 G 
               
               
                   
               
               
                 116 
                 116 
                 TACTCGGTATTCTCTTTAAACTACTTCCCT[A/G]TATCCCCTTCTGGTGAGCTTGGGACAGAAA 
                 G 
                 A 
               
               
                   
               
               
                 117 
                 117 
                 AAATCATCTAAAAAGCAAGATATGAAATTG[A/G]AAATGCATTCACATTTCACAGATCGTCGTC 
                 A 
                 G 
               
               
                   
               
               
                 118 
                 118 
                 GTTCATCAAAGCTCGAATTTTCCTAGTAGC[A/G]AGCCGGGACTTGGAGGTAAAACTGCAGTCT 
                 A 
                 G 
               
               
                   
               
               
                 119 
                 119 
                 CAATTAACAGAAACTGAGTCAATTCAAAAC[G/T]TTAGTTAGAGAAATTGCACATAAAAACACA 
                 T 
                 G 
               
               
                   
               
               
                 120 
                 120 
                 CCTCAACTCTCTCCCAACACAATCTTTTCC[A/C]CACTTTTTCAAATTGGATTTATTCCCAAGT 
                 C 
                 A 
               
               
                   
               
               
                 121 
                 121 
                 CACCAATACTTGGTCACCTGGCTGTAGTGG[A/C]CAAGAAAAAGCCTGTTTATCATTGTGTCTT 
                 A 
                 C 
               
               
                   
               
               
                 122 
                 122 
                 GTTGTCCAGCCCAAATTACAGTTTTGTATT[A/C]TCTGTGTGACGAAAAGTATACAATTTCGAT 
                 C 
                 A 
               
               
                   
               
               
                 123 
                 123 
                 TTCTCAGGGTCCATCATGGCGATCTCTGCA[A/G]TGGGTACTCAGTGGGTCCAGCTGCATGGTG 
                 G 
                 A 
               
               
                   
               
               
                 124 
                 124 
                 CATCAATTCCCCTGCAAGACCACAGAAAGT[G/T]ATTTGCCTTGTATTTTGCAAAGACCAAACT 
                 T 
                 G 
               
               
                   
               
               
                 125 
                 125 
                 CTGTTTGTACTGGAGAGCCATGTAATCCTA[G/T]ATCCTGTACTATCTATCCTCTTATTGTTTA 
                 G 
                 T 
               
               
                   
               
               
                 126 
                 126 
                 GGAGGTGTTCTTTACAGGGGACGAGGAGGA[A/G]GAGCTGTCTGAGAGAAAGTCCCAGAGAAAT 
                 G 
                 A 
               
               
                   
               
               
                 127 
                 127 
                 GCCCGAACGTCCTGTCCTCTATCCGCGTCA[A/G]GGTGTTGAAGGACAAATCCAGGTAAGCAAG 
                 A 
                 G 
               
               
                   
               
               
                 128 
                 128 
                 AGGCGAATGGCAGCGTCAGTAAAGTTATGT[C/T]TCTCCCTCTCAAAGTTCCTCCTCTGCTCGT 
                 C 
                 T 
               
               
                   
               
               
                 129 
                 129 
                 GTCTAATGAACACTGCCGTGGATTGTCATC[C/T]GAGTTGTCTGTTGAGTCCACAGTAACACAA 
                 C 
                 T 
               
               
                   
               
               
                 130 
                 130 
                 ATACTCTGCTAAAATTAGAACTGACCAGTT[G/T]ATTTTTTTTACATGATACATGATAATAATG 
                 G 
                 T 
               
               
                   
               
               
                 131 
                 131 
                 GTTGAATTTGGTCATATAAGGAAATGTGCC[C/T]TACTGCCTTTCGAATTTTGTGAAACTTAAA 
                 T 
                 C 
               
               
                   
               
               
                 132 
                 132 
                 GATACAGTCTGCATTTCAACAGGCTGCATT[G/T]TATTGCACCACACCCTGCTGTTATCAGATA 
                 T 
                 G 
               
               
                   
               
               
                 133 
                 133 
                 GGAAAATGTAACAACCAAACAGGACAGCAC[G/T]CGGTCTGTTGATGTTACAGGCAGCTCAGCT 
                 T 
                 G 
               
               
                   
               
               
                 134 
                 134 
                 CCATATTTCTTTGTTGTTGTTTGGAATAGT[A/C]TCCTACTCGAAAAAATATGCATTTGTATTT 
                 A 
                 C 
               
               
                   
               
               
                 135 
                 135 
                 GCCAGCGGGTCGGCACAGCCTAGTCATGCG[A/G]TCACAAGCCATTCAGTCTGGTCGCAGTCTA 
                 G 
                 A 
               
               
                   
               
               
                 136 
                 136 
                 AACAAACTAATCATCTCTAGTGGAGCGCAC[C/T]CGCACCACAAGGGGGGCCTTATCGTCGACC 
                 C 
                 T 
               
               
                   
               
               
                 137 
                 137 
                 AACACTGCCCTGCCCACCACAGAGGACATC[A/G]CGCTGTTCACGGATTTAGACCAGGGCAACA 
                 A 
                 G 
               
               
                   
               
               
                 138 
                 138 
                 AACTTTGTGTGCGTTCTATCCTTGTTTCCC[C/T]CCTACAGCCAGCGGTAACATCCCGGCCATG 
                 C 
                 T 
               
               
                   
               
               
                 139 
                 139 
                 TCAACTGAACAGACAGCAACTTTCCCTCCT[C/T]GTTCATTTCATCTTCACCTGCTTCTTCTTT 
                 C 
                 T 
               
               
                   
               
               
                 140 
                 140 
                 AGTGTCGATGCCAAACCCTTGAAAATAGGT[C/T]GGGAAAGGAAATCCTTTCCCTCTAAATATC 
                 T 
                 C 
               
               
                   
               
               
                 141 
                 141 
                 ACTAGTGGAGATGAAGGGGACTGGTAATGA[G/T]AATACAGTGACAGTAGGCAGTGTTGTAAAT 
                 G 
                 T 
               
               
                   
               
               
                 142 
                 142 
                 TGTACTTGTTTTATTAAAACACAATGCAAG[A/G]CTTAAAACGAACTGTACTGTAGATATAACC 
                 A 
                 G 
               
               
                   
               
               
                 143 
                 143 
                 CCGGGTTCTGAACGAATAGGCCTACATAAC[A/G]GGACATGGATTACGATGGGACCGTTACGAA 
                 A 
                 G 
               
               
                   
               
               
                 144 
                 144 
                 CATCTCTTTCCTCCACAATGCATACTTGGG[C/T]TTATGCCTTGGGTTGGTGTATCTGAAGACA 
                 T 
                 C 
               
               
                   
               
               
                 145 
                 145 
                 AATGGCCTTTGCAACGTGTTCCCAGCACAC[A/G]CTCAGCTCGGGGAGCTTTCTGTGGTCCTCC 
                 A 
                 G 
               
               
                   
               
               
                 146 
                 146 
                 TCATTAATGTGTAGTAGCAGTTATTTAGTT[C/T]ATAATAGTGATACAACAATAATACTTCATT 
                 T 
                 C 
               
               
                   
               
               
                 147 
                 147 
                 CATTTCCTCGTATCACTAGGAGTTCCTAAC[A/G]TAACAGATGTAAGCTAGTGAGCTGTATGCT 
                 A 
                 G 
               
               
                   
               
               
                 148 
                 148 
                 AAATAGGACACCATACCTGGTTGATTTCGT[G/T]TCGGTTGTGTGAAGTCAATGTTGTAAATGT 
                 G 
                 T 
               
               
                   
               
               
                 149 
                 149 
                 GAGATTCTAATTATATATTTTTTTAGAGAT[G/T]AATTATTTTGGAGGTTTATATACTCACATA 
                 G 
                 T 
               
               
                   
               
               
                 150 
                 150 
                 CATGAATAGTGGCAAAATGTCCTTCAATTT[C/T]GGTAGCTAGATCATAAAATACATTTGTACT 
                 T 
                 C 
               
               
                   
               
               
                 151 
                 151 
                 TGGTCACCAGCTGCATTAAGTACTGCAGTG[C/T]ATCTCCTCAGACTGCACCAGATATGTTCTT 
                 C 
                 T 
               
               
                   
               
               
                 152 
                 152 
                 AGTTTATTGGCGTAAACATAATCTAGAAGT[A/C]ATTTTCATAATATGCAACAATTGGCATGTA 
                 A 
                 C 
               
               
                   
               
               
                 153 
                 153 
                 ACTCTTGAGATTCGGATCAGCCTAGATGGA[C/T]GATTGAAGCTCCAGTCGATGATGAATCAAT 
                 C 
                 T 
               
               
                   
               
               
                 154 
                 154 
                 ATTTTCCCTATAATTTTGAAGAAATTGTTT[A/C]TTTAGCTCCTAATTATACACCCAGGTGTAG 
                 C 
                 A 
               
               
                   
               
               
                 155 
                 155 
                 GCCGATGGAAACAAGGTTATGGTTAGTGAC[A/C]ATACACAGCGTACCCAGTCTAATCCAAAGA 
                 A 
                 C 
               
               
                   
               
               
                 156 
                 156 
                 GAACGGCCTGAACACCATGGACCTCAACAC[A/G]GAGTTCACTGTGGCTCGCCTGGAAGCAATA 
                 A 
                 G 
               
               
                   
               
               
                 157 
                 157 
                 CTAAGGAGGAAATGGAGCTTGTCTTGCTAA[A/G]GTCAATGGTGATGTCAATGTCTCCCTCTCA 
                 A 
                 G 
               
               
                   
               
               
                 158 
                 158 
                 GATCTTGTACGCTGTCTGTGGGCTCCCTGA[G/T]AAATAATTGTAAGAAGCCTACTGATGCCAT 
                 G 
                 T 
               
               
                   
               
               
                 159 
                 159 
                 TCTGCGTCATTCCATCCCTGCCATCACACT[C/T]CTGCTACACTGCATTCTGCGTGTGGTGTAT 
                 C 
                 T 
               
               
                   
               
               
                 160 
                 160 
                 AAAAACTGCGTGTTTATCAAATTAGAAATT[C/T]ACAACCATAAAGTTTTGCGTTGACAAAAAA 
                 C 
                 T 
               
               
                   
               
               
                 161 
                 161 
                 CTTTGAGCTGGATCCAGTCAAGAAGAAGGC[C/T]AAAGAGAGGATGGTGAAGTCCACCAGCAGC 
                 C 
                 T 
               
               
                   
               
               
                 162 
                 162 
                 AAAGCTGCTTCCAGCTGATGGTGTAGTCTA[C/T]GGACAGGTTCGTTACCCATCCACATCTCTC 
                 C 
                 T 
               
               
                   
               
               
                 163 
                 163 
                 AGAATAAATGCAATCTTAATGTGAATTTAT[G/T]TATGTTTAGAAACTGCTTACTAATAGGTAA 
                 T 
                 G 
               
               
                   
               
               
                 164 
                 164 
                 CTGAATCAAATGCTCTCACTTGCCAGACTT[C/T]GAGGTTGACAGTGCAGCAATGACAAATACA 
                 T 
                 C 
               
               
                   
               
            
           
         
       
     
     SEQ ID NOs: 1 to 164 has an “n” at the polymorphic site (position 31; indicated in brackets in table 2), wherein “n” is either the late maturation allele or the normal maturation allele. SEQ ID NOs: 165 to 328 are identical to SEQ ID NOs: 1 to 164 except that the “n” at the polymorphic site is the late maturation allele. 
     According to particular embodiments, the at least one SNP of the invention is selected from the group consisting of SNP #1 to SNP #164 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 1 and is selected from the group consisting of SNP #1 to SNP #4 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 2 and is selected from the group consisting of SNP #5 to SNP #12 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 3 and is selected from the group consisting of SNP #13 to SNP #16 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 4 and is selected from the group consisting of SNP #17 to SNP #20 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 5 and is selected from the group consisting of SNP #21 to SNP #24 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 6 and is selected from the group consisting of SNP #25 to SNP #28 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 7 and is selected from the group consisting of SNP #29 to SNP #32 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 8 and is selected from the group consisting of SNP #33 to SNP #36 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 9 and is selected from the group consisting of SNP #37 to SNP #40 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 10 and is selected from the group consisting of SNP #41 to SNP #43 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 11 and is selected from the group consisting of SNP #44 to SNP #47 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 12 and is selected from the group consisting of SNP #48 to SNP #50 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 13 and is selected from the group consisting of SNP #51 to SNP #53 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 14 and is selected from the group consisting of SNP #54 to SNP #57 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 16 and is selected from the group consisting of SNP #62 to SNP #64 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 17 and is selected from the group consisting of SNP #65 to SNP #66 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 18 and is selected from the group consisting of SNP #67 to SNP #69 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 19 and is selected from the group consisting of SNP #70 to SNP #73 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 20 and is selected from the group consisting of SNP #74 to SNP #76 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 21 and is selected from the group consisting of SNP #77 to SNP #80 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 22 and is selected from the group consisting of SNP #81 to SNP #83 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 23 and is selected from the group consisting of SNP #84 to SNP #89 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 24 and is selected from the group consisting of SNP #90 to SNP #92 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 25 and is selected from the group consisting of SNP #93 to SNP #101 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 26 and is selected from the group consisting of SNP #102 to SNP #105 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 27 and is selected from the group consisting of SNP #106 to SNP #109 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #110 to SNP #161 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #120, SNP #127 to SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 to SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137 and SNP #128 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128 and SNP #144 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144 and SNP #142 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, and SNP #127 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127 and SNP #146 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146 and SNP #147 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147 and SNP #131 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131 and SNP #129 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129 and SNP #120 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120 and SNP #130 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120, SNP #130 and SNP #138 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 and SNP #143 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143 and SNP #144 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144 and SNP #146 listed in table 1. 
     According to particular preferred embodiments, the at least one SNP of the invention is located on chromosome 28 and is selected from the group consisting of SNP #130, SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is located on chromosome 29 and is selected from the group consisting of SNP #162 to SNP #164 listed in table 1. 
     According to particular embodiments, the at least one SNP of the invention is selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147. 
     According to particular embodiments, the at least one SNP of the invention is selected from the group consisting of SNP #93, SNP #114, SNP #120, SNP #129, SNP #130, SNP #131, SNP #132, SNP #134, SNP #138 and SNP #148. 
     According to particular embodiments, the at least one SNP of the invention is selected from the group consisting of SNP #5, SNP #24, SNP #86, SNP #121, SNP #125, SNP #126, SNP #133, SNP #141, SNP #150 and SNP #154. 
     According to particular embodiments, the at least one SNP of the invention is selected from the group consisting of SNP #7, SNP #9, SNP #19, SNP #85, SNP #111, SNP #116, SNP #117, SNP #119, SNP #153 and SNP #155. 
     According to particular embodiments, the at least one SNP of the invention is selected from the group consisting of SNP #10, SNP #11, SNP #23, SNP #88, SNP #89, SNP #91, SNP #101, SNP #124, SNP #145 and SNP #151. 
     It is understood that the foregoing disclosure regarding the polymorphisms of the invention, and in particular regarding SNPs and late maturation allele(s), is applicable to the following aspects. 
     Methods for Predicting 
     The present invention provides in a first aspect, a method for predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising determining the presence of at least one allele conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 1 and being selected from SNP #1 to SNP #4 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 2 and being selected from SNP #5 to SNP #12 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 3 and being selected from SNP #13 to SNP #16 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 4 and being selected from SNP #17 to SNP #20 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 5 and being selected from SNP #21 to SNP #24 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 6 and being selected from SNP #25 to SNP #28 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 7 and being selected from SNP #29 to SNP #32 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 8 and being selected from SNP #33 to SNP #36 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 9 and being selected from SNP #37 to SNP #40 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 10 and being selected from SNP #41 to SNP #43 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 11 and being selected from SNP #44 to SNP #47 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 12 and being selected from SNP #48 to SNP #50 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 13 and being selected from SNP #51 to SNP #53 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 14 and being selected from SNP #54 to SNP #57 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 15 and being selected from SNP #58 to SNP #61 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 16 and being selected from SNP #62 to SNP #64 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 17 and being selected from SNP #65 to SNP #66 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 18 and being selected from SNP #67 to SNP #69 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 19 and being selected from SNP #70 to SNP #73 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 20 and being selected from SNP #74 to SNP #76 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 21 and being selected from SNP #77 to SNP #80 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 22 and being selected from SNP #81 to SNP #83 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 23 and being selected from SNP #84 to SNP #89 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 24 and being selected from SNP #90 to SNP #92 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 25 and being selected from SNP #93 to SNP #101 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 26 and being selected from SNP #102 to SNP #105 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 27 and being selected from SNP #106 to SNP #109 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from SNP #110 to SNP #161 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #120, SNP #127 to SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 to SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137 and SNP #128 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144 and SNP #142 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, and SNP #127 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147 and SNP #131 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131 and SNP #129 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129 and SNP #120 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120 and SNP #130 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120, SNP #130 and SNP #138 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 and SNP #143 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #130, SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 29 and being selected from SNP #162 to SNP #164 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #148 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #5, SNP #24, SNP #86, SNP #121, SNP #125, SNP #126, SNP #133, SNP #141, SNP #150 and SNP #154 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #7, SNP #9, SNP #19, SNP #85, SNP #111, SNP #116, SNP #117, SNP #119, SNP #153 and SNP #155 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #10, SNP #11, SNP #23, SNP #88, SNP #89, SNP #91, SNP #101, SNP #124, SNP #145 and SNP #151 listed in table 1. 
     A preferred embodiment according to the first aspect of the present invention relates to a method for predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the presence of at least one allele conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being SNP #140 listed in table 1 wherein the presence of a thymine at the position corresponding to position 31 of SEQ ID NO. 140 indicates that the rainbow trout has late onset of sexual maturation. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 164, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 1 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 4, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 1 to 4 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 5 to 12, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 5 to 12 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 13 to 16, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 13 to 16 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 17 to 20, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 17 to 20 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 21 to 24, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 21 to 24 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 25 to 28, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 25 to 28 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 29 to 32, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 29 to 32 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 33 to 36, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 33 to 36 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 37 to 40, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 37 to 40 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 41 to 43, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 41 to 43 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 44 to 47, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 44 to 47 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 48 to 50, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 48 to 50 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 51 to 53, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 51 to 53 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 54 to 57, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 54 to 57 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 58 to 61, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 58 to 61 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 62 to 64, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 62 to 64 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 65 to 66, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 65 to 66 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 67 to 69, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 67 to 69 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 70 to 73, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 70 to 73 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 74 to 76, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 74 to 76 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 77 to 80, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 77 to 80 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 81 to 83, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 81 to 83 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 84 to 89, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 84 to 89 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 90 to 92, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 90 to 92 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 93 to 101, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 93 to 101 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 102 to 105, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 102 to 105 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 106 to 109, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 106 to 109 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain particularly preferred embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 110 to 161, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 110 to 161 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 162 to 164, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 162 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs:127, 128, 135, 137, 140, 142, 143, 144, 146 and 147, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 127, 128, 135, 137, 140, 142, 143, 144, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 93, 114, 120, 129, 130, 131, 132, 134, 138 and 148, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 93, 114, 120, 129, 130, 131, 132, 134, 138 and 148 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 5, 24, 86, 121, 125, 126, 133, 141, 150 and 154, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 5, 24, 86, 121, 125, 126, 133, 141, 150 and 154 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 7, 9, 19, 85, 111, 116, 117, 119, 153 and 155 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 7, 9, 19, 85, 111, 116, 117, 119, 153 and 155 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 10, 11, 23, 88, 89, 91, 101, 124, 145 and 151, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 10, 11, 23, 88, 89, 91, 101, 124, 145 and 151 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 120, 127 to 131, 134, 135, 137, 138, 140, 142 to 144, 146 and 147, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 120, 127 to 131, 134, 135, 137, 138, 140, 142 to 144, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143 and 137, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143 and 137 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137 and 128 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143 and 128 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128 and 144 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 128 and 144 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144 and 142 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144 and 142 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142 and 127 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142 and 127 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127 and 146 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146 and 147 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147 and 131 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147 and 131 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131 and 129 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131 and 129 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129 and 120 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129 and 120 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120 and 130 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120 and 130 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120, 130 and 138 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120, 130 and 138 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143 and 137 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143 and 137 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 135, 137, 138, 140, 142 and 143 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 135, 137, 138, 140, 142 and 143 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 134, 135, 137, 138, 140, 142, 143 and 144 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 134, 135, 137, 138, 140, 142, 143 and 144 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     According to certain embodiments, the present invention provides a method of predicting late onset of sexual maturation in rainbow trout ( Oncorhynchus mykiss ), the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 130, 131, 134, 135, 137, 138, 140, 142, 143, 144, 146 and 147 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     Methods for Selecting 
     The present invention provides in a second aspect, a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising determining the presence of at least one allele conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; and selecting said rainbow trout as having late onset of sexual maturation when the at least one late maturation allele is present; wherein the at least one late maturation allele is an allele of at least one SNP, the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 1 and being selected from SNP #1 to SNP #4 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 2 and being selected from SNP #5 to SNP #12 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 3 and being selected from SNP #13 to SNP #16 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 4 and being selected from SNP #17 to SNP #20 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 5 and being selected from SNP #21 to SNP #24 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 6 and being selected from SNP #25 to SNP #28 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 7 and being selected from SNP #29 to SNP #32 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 8 and being selected from SNP #33 to SNP #36 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 9 and being selected from SNP #37 to SNP #40 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 10 and being selected from SNP #41 to SNP #43 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 11 and being selected from SNP #44 to SNP #47 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 12 and being selected from SNP #48 to SNP #50 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 13 and being selected from SNP #51 to SNP #53 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 14 and being selected from SNP #54 to SNP #57 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 15 and being selected from SNP #58 to SNP #61 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 16 and being selected from SNP #62 to SNP #64 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 17 and being selected from SNP #65 to SNP #66 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 18 and being selected from SNP #67 to SNP #69 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 19 and being selected from SNP #70 to SNP #73 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 20 and being selected from SNP #74 to SNP #76 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 21 and being selected from SNP #77 to SNP #80 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 22 and being selected from SNP #81 to SNP #83 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 23 and being selected from SNP #84 to SNP #89 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 24 and being selected from SNP #90 to SNP #92 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 25 and being selected from SNP #93 to SNP #101 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 26 and being selected from SNP #102 to SNP #105 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 27 and being selected from SNP #106 to SNP #109 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from SNP #110 to SNP #161 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #120, SNP #127 to SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 to SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137 and SNP #128 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144 and SNP #142 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, and SNP #127 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147 and SNP #131 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131 and SNP #129 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129 and SNP #120 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120 and SNP #130 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120, SNP #130 and SNP #138 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 and SNP #143 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #130, SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 29 and being selected from SNP #162 to SNP #164 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #148 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #5, SNP #24, SNP #86, SNP #121, SNP #125, SNP #126, SNP #133, SNP #141, SNP #150 and SNP #154 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #7, SNP #9, SNP #19, SNP #85, SNP #111, SNP #116, SNP #117, SNP #119, SNP #153 and SNP #155 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #10, SNP #11, SNP #23, SNP #88, SNP #89, SNP #91, SNP #101, SNP #124, SNP #145 and SNP #151 listed in table 1. 
     A preferred embodiment according to the first aspect of the present invention relates to a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the presence of at least one allele conferring late onset of sexual maturation (“late maturation allele”) within the genome of said rainbow trout; and selecting said rainbow trout as having late onset of sexual maturation when the at least one late maturation allele is present; wherein the at least one late maturation allele is an allele of at least one SNP, the at least one SNP being SNP #140 listed in table 1 wherein the presence of a thymine at the position corresponding to position 31 of SEQ ID NO. 140 indicates that the rainbow trout has late onset of sexual maturation. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 164, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 1 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 4, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 1 to 4 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 5 to 12, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 5 to 12 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 13 to 16, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 13 to 16 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 17 to 20, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 17 to 20 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 21 to 24, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 21 to 24 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 25 to 28, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 25 to 28 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 29 to 32, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 29 to 32 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 33 to 36, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 33 to 36 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 37 to 40, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 37 to 40 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 41 to 43, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 41 to 43 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 44 to 47, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 44 to 47 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 48 to 50, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 48 to 50 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 51 to 53, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 51 to 53 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 54 to 57, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 54 to 57 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 58 to 61, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 58 to 61 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 62 to 64, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 62 to 64 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 65 to 66, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 65 to 66 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 67 to 69, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 67 to 69 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 70 to 73, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 70 to 73 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 74 to 76, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 74 to 76 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 77 to 80, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 77 to 80 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 81 to 83, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 81 to 83 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 84 to 89, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 84 to 89 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 90 to 92, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 90 to 92 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 93 to 101, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 93 to 101 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 102 to 105, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 102 to 105 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 106 to 109, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 106 to 109 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 110 to 161, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 110 to 161 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 162 to 164, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 162 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 127, 128, 135, 137, 140, 142, 143, 144, 146 and 147, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 127, 128, 135, 137, 140, 142, 143, 144, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 93, 114, 120, 129, 130, 131, 132, 134, 138 and 148, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 93, 114, 120, 129, 130, 131, 132, 134, 138 and 148 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 5, 24, 86, 121, 125, 126, 133, 141, 150 and 154, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 5, 24, 86, 121, 125, 126, 133, 141, 150 and 154 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 7, 9, 19, 85, 111, 116, 117, 119, 153 and 155, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 7, 9, 19, 85, 111, 116, 117, 119, 153 and 155 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 10, 11, 23, 88, 89, 91, 101, 124, 145 and 151, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 10, 11, 23, 88, 89, 91, 101, 124, 145 and 151 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 120, 127 to 131, 134, 135, 137, 138, 140, 142 to 144, 146 and 147, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 120, 127 to 131, 134, 135, 137, 138, 140, 142 to 144, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143 and 137, or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143 and 137 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137 and 128 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143 and 128 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128 and 144 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 128 and 144 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144 and 142 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144 and 142 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142 and 127 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142 and 127 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127 and 146 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146 and 147 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147 and 131 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147 and 131 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131 and 129 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131 and 129 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129 and 120 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129 and 120 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120 and 130 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120 and 130 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120, 130 and 138 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120, 130 and 138 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143 and 137 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 140, 135, 143 and 137 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 135, 137, 138, 140, 142 and 143 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 135, 137, 138, 140, 142 and 143by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 134, 135, 137, 138, 140, 142, 143 and 144 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 134, 135, 137, 138, 140, 142, 143 and 144 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     According to certain particularly preferred embodiments, the present invention provides a method for selecting a rainbow trout having late onset of sexual maturation, the method comprising: determining the identity of a nucleotide of at least one allele of at least one SNP associated with late onset of sexual maturation within the genome of said rainbow trout; the at least one SNP being located within said genome at a position corresponding to position 31 of the nucleotide sequence set forth in any one of SEQ ID NOs: 130, 131, 134, 135, 137, 138, 140, 142, 143, 144, 146 and 147 or at a position corresponding to position 31 of a nucleotide sequence which is derived from any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions; with the proviso that said nucleotide substitution(s) are not at position 31 of said derived sequences; and selecting said rainbow trout as having late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the SNP. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     Numerous techniques are known in the art for determining the identity of a nucleotide of an allele present at a polymorphic site. For example, the determination may involve sequence analysis of the rainbow trout to be tested using, e.g., traditional sequence methodologies (e.g., the “dideoxy-mediated chain termination method,” also known as the “Sanger Method” (Sanger, F., et al., J. Molec. Biol. 94: 441 (1975); Prober et al. Science 238: 336-340 (1987)) and the “chemical degradation method” also known as the “Maxam-Gilbert method” (Maxam, A. M., et al., Proc. Natl. Acad. Sci. (U.S.A.) 74: 560 (1977). Alternatively, the determination may involve single base extension of DNA oligonucleotides terminating at the polymorphic site (e.g. iPLEX assays from Sequenom (San Diego, USA) and Infinium assays from Illumina (San Diego, USA), allele-specific ligation assays (e.g. Axiom technology from Affymetrix (San Diego, USA), allele-specific PCR (e.g. SNPtype assays from Fluidigm (San Francisco) or KASP assays from LGC Genomics (Teddington, UK)), or competitive hybridization of probes complementary to the different alleles (e.g. the TaqMan assay from Applied Biosystems (Foster City, USA)). 
     Methods for the detection of allelic variation are also reviewed by Nollau et al., Clin. Chem. 43, 1114-1120, 1997; and in standard textbooks, for example “Laboratory Protocols for Mutation Detection”, Ed. by U. Landegren, Oxford University Press, 1996 and “PCR”, 2nd Edition by Newton &amp; Graham, BIOS Scientific Publishers Limited, 1997. 
     For analyzing SNPs, it may for example be appropriate to use oligonucleotides specific for alternative SNP alleles. Such oligonucleotides which detect single nucleotide variations in target sequences may be referred to by such terms as “allele-specific oligonucleotides”, “allele-specific probes”, or “allele-specific primers”. The design and use of allele-specific probes for analyzing polymorphisms is described in, e.g., Mutation Detection A Practical Approach, ed. Cotton et al. Oxford University Press, 1998; Saiki et al., Nature 324, 163-166 (1986); Dattagupta, EP235726; and Saiki, WO 89/11548. 
     Rainbow Trout 
     The present invention provides in a further aspect, a rainbow trout, such as an isolated rainbow trout, or a progeny thereof having late onset of sexual maturation comprising within its genome at least one allele, such as at least two alleles, conferring late onset of sexual maturation (“late maturation allele”); wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 1 and being selected from SNP #1 to SNP #4 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 2 and being selected from SNP #5 to SNP #12 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 3 and being selected from SNP #13 to SNP #16 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 4 and being selected from SNP #17 to SNP #20 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 5 and being selected from SNP #21 to SNP #24 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 6 and being selected from SNP #25 to SNP #28 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 7 and being selected from SNP #29 to SNP #32 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 8 and being selected from SNP #33 to SNP #36 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 9 and being selected from SNP #37 to SNP #40 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 10 and being selected from SNP #41 to SNP #43 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 11 and being selected from SNP #44 to SNP #47 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 12 and being selected from SNP #48 to SNP #50 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 13 and being selected from SNP #51 to SNP #53 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 14 and being selected from SNP #54 to SNP #57 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 15 and being selected from SNP #58 to SNP #61 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 16 and being selected from SNP #62 to SNP #64 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 17 and being selected from SNP #65 to SNP #66 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 18 and being selected from SNP #67 to SNP #69 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 19 and being selected from SNP #70 to SNP #73 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 20 and being selected from SNP #74 to SNP #76 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 21 and being selected from SNP #77 to SNP #80 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 22 and being selected from SNP #81 to SNP #83 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 23 and being selected from SNP #84 to SNP #89 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 24 and being selected from SNP #90 to SNP #92 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 25 and being selected from SNP #93 to SNP #101 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 26 and being selected from SNP #102 to SNP #105 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 27 and being selected from SNP #106 to SNP #109 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from SNP #110 to SNP #161 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #120, SNP #127 to SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 to SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137 and SNP #128 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144 and SNP #142 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, and SNP #127 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147 and SNP #131 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131 and SNP #129 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129 and SNP #120 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120 and SNP #130 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120, SNP #130 and SNP #138 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 and SNP #143 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #130, SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 29 and being selected from SNP #162 to SNP #164 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #93, SNP #114, SNP #120, SNP #129, SNP #130, SNP #131, SNP #132, SNP #134, SNP #138 and SNP #148 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #5, SNP #24, SNP #86, SNP #121, SNP #125, SNP #126, SNP #133, SNP #141, SNP #150 and SNP #154 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #7, SNP #9, SNP #19, SNP #85, SNP #111, SNP #116, SNP #117, SNP #119, SNP #153 and SNP #155 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #10, SNP #11, SNP #23, SNP #88, SNP #89, SNP #91, SNP #101, SNP #124, SNP #145 and SNP #151 listed in table 1. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 168, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 168 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 169 to 176, and b) nucleotide sequences derived from any one of SEQ ID NOs: 169 to 176 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 177 to 180, and b) nucleotide sequences derived from any one of SEQ ID NOs: 177 to 180 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 181 to 184, and b) nucleotide sequences derived from any one of SEQ ID NOs: 181 to 184 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 185 to 188, and b) nucleotide sequences derived from any one of SEQ ID NOs: 185 to 188 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 189 to 192, and b) nucleotide sequences derived from any one of SEQ ID NOs: 189 to 192 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 193 to 196, and b) nucleotide sequences derived from any one of SEQ ID NOs: 193 to 196 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 197 to 200, and b) nucleotide sequences derived from any one of SEQ ID NOs: 197 to 200 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 201 to 204, and b) nucleotide sequences derived from any one of SEQ ID NOs: 201 to 204 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 205 to 207, and b) nucleotide sequences derived from any one of SEQ ID NOs: 205 to 207 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 208 to 211, and b) nucleotide sequences derived from any one of SEQ ID NOs: 208 to 211 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 212 to 214, and b) nucleotide sequences derived from any one of SEQ ID NOs: 212 to 214 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 215 to 217, and b) nucleotide sequences derived from any one of SEQ ID NOs: 215 to 217 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 218 to 221, and b) nucleotide sequences derived from any one of SEQ ID NOs: 218 to 221 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 222 to 225, and b) nucleotide sequences derived from any one of SEQ ID NOs: 222 to 225 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 226 to 228, and b) nucleotide sequences derived from any one of SEQ ID NOs: 226 to 228 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 229 to 230, and b) nucleotide sequences derived from any one of SEQ ID NOs: 229 to 230 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 231 to 233, and b) nucleotide sequences derived from any one of SEQ ID NOs: 231 to 233 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 234 to 237, and b) nucleotide sequences derived from any one of SEQ ID NOs: 234 to 237 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 238 to 240, and b) nucleotide sequences derived from any one of SEQ ID NOs: 238 to 240 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 241 to 244, and b) nucleotide sequences derived from any one of SEQ ID NOs: 241 to 244 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 245 to 247, and b) nucleotide sequences derived from any one of SEQ ID NOs: 245 to 247 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 248 to 253, and b) nucleotide sequences derived from any one of SEQ ID NOs: 248 to 253 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 254 to 256, and b) nucleotide sequences derived from any one of SEQ ID NOs: 254 to 256 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 257 to 265, and b) nucleotide sequences derived from any one of SEQ ID NOs: 257 to 265 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 266 to 269, and b) nucleotide sequences derived from any one of SEQ ID NOs: 266 to 269 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 270 to 273, and b) nucleotide sequences derived from any one of SEQ ID NOs: 270 to 273 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain particularly preferred embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 274 to 325, and b) nucleotide sequences derived from any one of SEQ ID NOs: 274 to 325 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 326 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 326 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 291, 292, 299, 301, 304, 306, 307, 308, 310 and 311, and b) nucleotide sequences derived from any one of SEQ ID NOs: 291, 292, 299, 301, 304, 306, 307, 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 257, 278, 284, 293, 294, 295, 296, 298, 302 and 312, and b) nucleotide sequences derived from any one of SEQ ID NOs: 257, 278, 284, 293, 294, 295, 296, 298, 302 and 312by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 169, 188, 250, 285, 289, 290, 297, 305, 314 and 318, and b) nucleotide sequences derived from any one of SEQ ID NOs: 169, 188, 250, 285, 289, 290, 297, 305, 314 and 318 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 171, 173, 183, 249, 275, 280, 281, 283, 317 and 319, and b) nucleotide sequences derived from any one of SEQ ID NOs: 171, 173, 183, 249, 275, 280, 281, 283, 317 and 319 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 174, 175, 187, 252, 253, 255, 265, 288, 309 and 315, and b) nucleotide sequences derived from any one of SEQ ID NOs: 174, 175, 187, 252, 253, 255, 265, 288, 309 and 315 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 284, 291 to 295, 298, 299, 301, 302, 304, 306 to 308, 310 and 311, and b) nucleotide sequences derived from any one of SEQ ID NOs: 284, 291 to 295, 298, 299, 301, 302, 304, 306 to 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307 and 301, and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307 and 301 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301 and 292 and b) nucleotide sequences derived from any one of SEQ ID NOs: SEQ ID NOs: 304, 299, 307, 301 and 292 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292 and 308 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292 and 308 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308 and 306 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308 and 306 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306 and 291 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306 and 291 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291 and 310 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291 and 310 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310 and 311 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311 and 295 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311 and 295 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295 and 293 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295 and 293 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293 and 284 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293 and 284 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284 and 294 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284 and 294 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284, 294 and 302 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284, 294 and 302 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307 and 301 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307 and 301 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 299, 301, 302, 304, 306 and 307 and b) nucleotide sequences derived from any one of SEQ ID NOs: 299, 301, 302, 304, 306 and 307 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 298, 299, 301, 302, 304, 306, 307 and 308 and b) nucleotide sequences derived from any one of SEQ ID NOs: 298, 299, 301, 302, 304, 306, 307 and 308 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 295, 298, 299, 301, 302, 304, 306, 307, 308 and 310 and b) nucleotide sequences derived from any one of SEQ ID NOs: 295, 298, 299, 301, 302, 304, 306, 307, 308 and 310 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout, such as an isolated rainbow trout, or a progeny thereof which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 294, 295, 298, 299, 301, 302, 304, 306, 307, 308, 310 and 311 and b) nucleotide sequences derived from any one of SEQ ID NOs: 294, 295, 298, 299, 301, 302, 304, 306, 307, 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     The present invention provides in a further aspect a population of rainbow trout, such as an isolated population of rainbow trout, each individual rainbow trout within the population being a rainbow trout according to the further aspect as discussed above. 
     According to certain embodiments, the rainbow trout is a female. 
     According to certain other embodiments, the rainbow trout is a male. 
     According to certain embodiments, the rainbow trout or progeny thereof is obtained by a method according to the present invention. 
     In one further aspect of the present invention, a rainbow trout or progeny thereof comprises in its genome at least one allele conferring late sexual maturation obtainable by a process comprising the steps of:
         a) genotyping the trout,   b) selecting individuals having at least one allele preferably two alleles conferring late onset of sexual maturation (“late maturation allele”); and   c) mating individuals in such a way that at least one individual within each mated pair has two alleles conferring late onset of sexual maturation.       

     According to certain embodiments the mating in c) may also be conducted in such a way that the mated pair each has two alleles conferring late onset of sexual maturation, or that each mated pair has one allele conferring late onset of sexual maturation. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     Rainbow Trout Cell 
     The present invention provides in a third aspect, a rainbow trout cell, such as an isolated rainbow trout cell, comprising within its genome at least one allele conferring late onset of sexual maturation (“late maturation allele”); wherein the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 1 and being selected from SNP #1 to SNP #4 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 2 and being selected from SNP #5 to SNP #12 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 3 and being selected from SNP #13 to SNP #16 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 4 and being selected from SNP #17 to SNP #20 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 5 and being selected from SNP #21 to SNP #24 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 6 and being selected from SNP #25 to SNP #28 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 7 and being selected from SNP #29 to SNP #32 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 8 and being selected from SNP #33 to SNP #36 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 9 and being selected from SNP #37 to SNP #40 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 10 and being selected from SNP #41 to SNP #43 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 11 and being selected from SNP #44 to SNP #47 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 12 and being selected from SNP #48 to SNP #50 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 13 and being selected from SNP #51 to SNP #53 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 14 and being selected from SNP #54 to SNP #57 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 15 and being selected from SNP #58 to SNP #61 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 16 and being selected from SNP #62 to SNP #64 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 17 and being selected from SNP #65 to SNP #66 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 18 and being selected from SNP #67 to SNP #69 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 19 and being selected from SNP #70 to SNP #73 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 20 and being selected from SNP #74 to SNP #76 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 21 and being selected from SNP #77 to SNP #80 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 22 and being selected from SNP #81 to SNP #83 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 23 and being selected from SNP #84 to SNP #89 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 24 and being selected from SNP #90 to SNP #92 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 25 and being selected from SNP #93 to SNP #101 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 26 and being selected from SNP #102 to SNP #105 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 27 and being selected from SNP #106 to SNP #109 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from SNP #110 to SNP #161 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #120, SNP #127 to SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 to SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137 and SNP #128 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144 and SNP #142 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, and SNP #127 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147 and SNP #131 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131 and SNP #129 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129 and SNP #120 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120 and SNP #130 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120, SNP #130 and SNP #138 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 and SNP #143 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #130, SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 29 and being selected from SNP #162 to SNP #164 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #93, SNP #114, SNP #120, SNP #129, SNP #130, SNP #131, SNP #132, SNP #134, SNP #138 and SNP #148 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #5, SNP #24, SNP #86, SNP #121, SNP #125, SNP #126, SNP #133, SNP #141, SNP #150 and SNP #154 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #7, SNP #9, SNP #19, SNP #85, SNP #111, SNP #116, SNP #117, SNP #119, SNP #153 and SNP #155 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #10, SNP #11, SNP #23, SNP #88, SNP #89, SNP #91, SNP #101, SNP #124, SNP #145 and SNP #151 listed in table 1. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 168, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 168 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 169 to 176, and b) nucleotide sequences derived from any one of SEQ ID NOs: 169 to 176 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 177 to 180, and b) nucleotide sequences derived from any one of SEQ ID NOs: 177 to 180 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 181 to 184, and b) nucleotide sequences derived from any one of SEQ ID NOs: 181 to 184 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 185 to 188, and b) nucleotide sequences derived from any one of SEQ ID NOs: 185 to 188 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 189 to 192, and b) nucleotide sequences derived from any one of SEQ ID NOs: 189 to 192 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 193 to 196, and b) nucleotide sequences derived from any one of SEQ ID NOs: 193 to 196 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 197 to 200, and b) nucleotide sequences derived from any one of SEQ ID NOs: 197 to 200 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 201 to 204, and b) nucleotide sequences derived from any one of SEQ ID NOs: 201 to 204 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 205 to 207, and b) nucleotide sequences derived from any one of SEQ ID NOs: 205 to 207 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 208 to 211, and b) nucleotide sequences derived from any one of SEQ ID NOs: 208 to 211 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 212 to 214, and b) nucleotide sequences derived from any one of SEQ ID NOs: 212 to 214 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 215 to 217, and b) nucleotide sequences derived from any one of SEQ ID NOs: 215 to 217 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 218 to 221, and b) nucleotide sequences derived from any one of SEQ ID NOs: 218 to 221 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 222 to 225, and b) nucleotide sequences derived from any one of SEQ ID NOs: 222 to 225 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 226 to 228, and b) nucleotide sequences derived from any one of SEQ ID NOs: 226 to 228 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 229 to 230, and b) nucleotide sequences derived from any one of SEQ ID NOs: 229 to 230 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 231 to 233, and b) nucleotide sequences derived from any one of SEQ ID NOs: 231 to 233 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 234 to 237, and b) nucleotide sequences derived from any one of SEQ ID NOs: 234 to 237 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 238 to 240, and b) nucleotide sequences derived from any one of SEQ ID NOs: 238 to 240 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 241 to 244, and b) nucleotide sequences derived from any one of SEQ ID NOs: 241 to 244 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 245 to 247, and b) nucleotide sequences derived from any one of SEQ ID NOs: 245 to 247 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 248 to 253, and b) nucleotide sequences derived from any one of SEQ ID NOs: 248 to 253 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 254 to 256, and b) nucleotide sequences derived from any one of SEQ ID NOs: 254 to 256 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 257 to 265, and b) nucleotide sequences derived from any one of SEQ ID NOs: 257 to 265 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 266 to 269, and b) nucleotide sequences derived from any one of SEQ ID NOs: 266 to 269 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 270 to 273, and b) nucleotide sequences derived from any one of SEQ ID NOs: 270 to 273 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain particularly preferred embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 274 to 325, and b) nucleotide sequences derived from any one of SEQ ID NOs: 274 to 325 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 326 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 326 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 291, 292, 299, 301, 304, 306, 307, 308, 310 and 311, and b) nucleotide sequences derived from any one of SEQ ID NOs: 291, 292, 299, 301, 304, 306, 307, 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 257, 278, 284, 293, 294, 295, 296, 298, 302 and 312, and b) nucleotide sequences derived from any one of SEQ ID NOs: 257, 278, 284, 293, 294, 295, 296, 298, 302 and 312 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 169, 188, 250, 285, 289, 290, 297, 305, 314 and 318, and b) nucleotide sequences derived from any one of SEQ ID NOs: 169, 188, 250, 285, 289, 290, 297, 305, 314 and 318 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 171, 173, 183, 249, 275, 280, 281, 283, 317 and 319, and b) nucleotide sequences derived from any one of SEQ ID NOs: 171, 173, 183, 249, 275, 280, 281, 283, 317 and 319 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 174, 175, 187, 252, 253, 255, 265, 288, 309 and 315, and b) nucleotide sequences derived from any one of SEQ ID NOs: 174, 175, 187, 252, 253, 255, 265, 288, 309 and 315 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 284, 291 to 295, 298, 299, 301, 302, 304, 306 to 308, 310 and 311, and b) nucleotide sequences derived from any one of SEQ ID NOs: 284, 291 to 295, 298, 299, 301, 302, 304, 306 to 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307 and 301, and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307 and 301 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301 and 292 and b) nucleotide sequences derived from any one of SEQ ID NOs: SEQ ID NOs: 304, 299, 307, 301 and 292 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292 and 308 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292 and 308 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308 and 306 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308 and 306 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306 and 291 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306 and 291 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291 and 310 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291 and 310 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310 and 311 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311 and 295 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311 and 295 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295 and 293 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295 and 293 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293 and 284 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293 and 284 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284 and 294 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284 and 294 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284, 294 and 302 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284, 294 and 302 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307 and 301 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307 and 301 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 299, 301, 302, 304, 306 and 307 and b) nucleotide sequences derived from any one of SEQ ID NOs: 299, 301, 302, 304, 306 and 307 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 298, 299, 301, 302, 304, 306, 307 and 308 and b) nucleotide sequences derived from any one of SEQ ID NOs: 298, 299, 301, 302, 304, 306, 307 and 308 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 295, 298, 299, 301, 302, 304, 306, 307, 308 and 310 and b) nucleotide sequences derived from any one of SEQ ID NOs: 295, 298, 299, 301, 302, 304, 306, 307, 308 and 310 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout cell, such as an isolated rainbow trout cell, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 294, 295, 298, 299, 301, 302, 304, 306, 307, 308, 310 and 311 and b) nucleotide sequences derived from any one of SEQ ID NOs: 294, 295, 298, 299, 301, 302, 304, 306, 307, 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     The present invention provides in a further aspect a population of rainbow trout cells, such as an isolated population of rainbow trout cells, each individual cell within the population being a cell according to the third aspect of the present invention. 
     According to certain embodiments, the rainbow trout cell is a gamete. 
     According to certain embodiments, the rainbow trout cell is not a gamete. 
     According to particular embodiments, the rainbow trout cell is an egg, such as an eyed egg. According to more particular embodiments, the egg is unfertilized. According to other more particular embodiments, the egg is fertilized. 
     According to particular embodiments, the rainbow trout cell is a sperm cell. 
     According to certain other embodiments, the rainbow trout cell is a somatic cell. 
     According to certain embodiments, the rainbow trout cell has been isolated from a rainbow trout of the invention. 
     According to particular embodiments, the rainbow trout cell has been isolated from a female rainbow trout of the invention. 
     According to particular embodiments, the rainbow trout cell has been isolated from a male rainbow trout of the invention. 
     Rainbow Trout Egg or Spermatozoa 
     The present invention provides in a fourth aspect, a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, comprising within its genome at least one allele conferring late onset of sexual maturation (“late maturation allele”); wherein the isolated rainbow trout egg is unfertilized; and the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the SNPs listed in Table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 1 and being selected from SNP #1 to SNP #4 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 2 and being selected from SNP #5 to SNP #12 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 3 and being selected from SNP #13 to SNP #16 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 4 and being selected from SNP #17 to SNP #20 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 5 and being selected from SNP #21 to SNP #24 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 6 and being selected from SNP #25 to SNP #28 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 7 and being selected from SNP #29 to SNP #32 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 8 and being selected from SNP #33 to SNP #36 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 9 and being selected from SNP #37 to SNP #40 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 10 and being selected from SNP #41 to SNP #43 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 11 and being selected from SNP #44 to SNP #47 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 12 and being selected from SNP #48 to SNP #50 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 13 and being selected from SNP #51 to SNP #53 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 14 and being selected from SNP #54 to SNP #57 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 15 and being selected from SNP #58 to SNP #61 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 16 and being selected from SNP #62 to SNP #64 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 17 and being selected from SNP #65 to SNP #66 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 18 and being selected from SNP #67 to SNP #69 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 19 and being selected from SNP #70 to SNP #73 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 20 and being selected from SNP #74 to SNP #76 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 21 and being selected from SNP #77 to SNP #80 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 22 and being selected from SNP #81 to SNP #83 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 23 and being selected from SNP #84 to SNP #89 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 24 and being selected from SNP #90 to SNP #92 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 25 and being selected from SNP #93 to SNP #101 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 26 and being selected from SNP #102 to SNP #105 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 27 and being selected from SNP #106 to SNP #109 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from SNP #110 to SNP #161 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #120, SNP #127 to SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 to SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137 and SNP #128 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144 and SNP #142 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, and SNP #127 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146 and SNP #147 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147 and SNP #131 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131 and SNP #129 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129 and SNP #120 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120 and SNP #130 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143, SNP #137, SNP #128, SNP #144, SNP #142, SNP #127, SNP #146, SNP #147, SNP #131, SNP #129, SNP #120, SNP #130 and SNP #138 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #140, SNP #135, SNP #143 and SNP #137 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #135, SNP #137, SNP #138, SNP #140, SNP #142 and SNP #143 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143 and SNP #144 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144 and SNP #146 listed in table 1. 
     According to certain particularly preferred embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 28 and being selected from the group consisting of SNP #130, SNP #131, SNP #134, SNP #135, SNP #137, SNP #138, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being located on chromosome 29 and being selected from SNP #162 to SNP #164 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #127, SNP #128, SNP #135, SNP #137, SNP #140, SNP #142, SNP #143, SNP #144, SNP #146 and SNP #147 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #93, SNP #114, SNP #120, SNP #129, SNP #130, SNP #131, SNP #132, SNP #134, SNP #138 and SNP #148 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #5, SNP #24, SNP #86, SNP #121, SNP #125, SNP #126, SNP #133, SNP #141, SNP #150 and SNP #154 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #7, SNP #9, SNP #19, SNP #85, SNP #111, SNP #116, SNP #117, SNP #119, SNP #153 and SNP #155 listed in table 1. 
     According to certain embodiments, the at least one late maturation allele is an allele of at least one single nucleotide polymorphism (SNP), the at least one SNP being selected from the group consisting of SNP #10, SNP #11, SNP #23, SNP #88, SNP #89, SNP #91, SNP #101, SNP #124, SNP #145 and SNP #151 listed in table 1. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 165 to 168, and b) nucleotide sequences derived from any one of SEQ ID NOs: 165 to 168 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 169 to 176, and b) nucleotide sequences derived from any one of SEQ ID NOs: 169 to 176 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 177 to 180, and b) nucleotide sequences derived from any one of SEQ ID NOs: 177 to 180 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 181 to 184, and b) nucleotide sequences derived from any one of SEQ ID NOs: 181 to 184 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 185 to 188, and b) nucleotide sequences derived from any one of SEQ ID NOs: 185 to 188 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 189 to 192, and b) nucleotide sequences derived from any one of SEQ ID NOs: 189 to 192 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 193 to 196, and b) nucleotide sequences derived from any one of SEQ ID NOs: 193 to 196 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 197 to 200, and b) nucleotide sequences derived from any one of SEQ ID NOs: 197 to 200 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 201 to 204, and b) nucleotide sequences derived from any one of SEQ ID NOs: 201 to 204 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 205 to 207, and b) nucleotide sequences derived from any one of SEQ ID NOs: 205 to 207 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 208 to 211, and b) nucleotide sequences derived from any one of SEQ ID NOs: 208 to 211 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 212 to 214, and b) nucleotide sequences derived from any one of SEQ ID NOs: 212 to 214 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 215 to 217, and b) nucleotide sequences derived from any one of SEQ ID NOs: 215 to 217 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 218 to 221, and b) nucleotide sequences derived from any one of SEQ ID NOs: 218 to 221 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 222 to 225, and b) nucleotide sequences derived from any one of SEQ ID NOs: 222 to 225 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 226 to 228, and b) nucleotide sequences derived from any one of SEQ ID NOs: 226 to 228 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 229 to 230, and b) nucleotide sequences derived from any one of SEQ ID NOs: 229 to 230 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 231 to 233, and b) nucleotide sequences derived from any one of SEQ ID NOs: 231 to 233 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 234 to 237, and b) nucleotide sequences derived from any one of SEQ ID NOs: 234 to 237 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 238 to 240, and b) nucleotide sequences derived from any one of SEQ ID NOs: 238 to 240 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 241 to 244, and b) nucleotide sequences derived from any one of SEQ ID NOs: 241 to 244 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 245 to 247, and b) nucleotide sequences derived from any one of SEQ ID NOs: 245 to 247 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 248 to 253, and b) nucleotide sequences derived from any one of SEQ ID NOs: 248 to 253 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 254 to 256, and b) nucleotide sequences derived from any one of SEQ ID NOs: 254 to 256 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 257 to 265, and b) nucleotide sequences derived from any one of SEQ ID NOs: 257 to 265 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 266 to 269, and b) nucleotide sequences derived from any one of SEQ ID NOs: 266 to 269 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 270 to 273, and b) nucleotide sequences derived from any one of SEQ ID NOs: 270 to 273 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 274 to 325, and b) nucleotide sequences derived from any one of SEQ ID NOs: 274 to 325 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 326 to 328, and b) nucleotide sequences derived from any one of SEQ ID NOs: 326 to 328 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 291, 292, 299, 301, 304, 306, 307, 308, 310 and 311, and b) nucleotide sequences derived from any one of SEQ ID NOs: 291, 292, 299, 301, 304, 306, 307, 308, 310 and 311by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 257, 278, 284, 293, 294, 295, 296, 298, 302 and 312, and b) nucleotide sequences derived from any one of SEQ ID NOs: 257, 278, 284, 293, 294, 295, 296, 298, 302 and 312 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 169, 188, 250, 285, 289, 290, 297, 305, 314 and 318, and b) nucleotide sequences derived from any one of SEQ ID NOs: 169, 188, 250, 285, 289, 290, 297, 305, 314 and 318 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 171, 173, 183, 249, 275, 280, 281, 283, 317 and 319, and b) nucleotide sequences derived from any one of SEQ ID NOs: 171, 173, 183, 249, 275, 280, 281, 283, 317 and 319 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 174, 175, 187, 252, 253, 255, 265, 288, 309 and 315, and b) nucleotide sequences derived from any one of SEQ ID NOs: 174, 175, 187, 252, 253, 255, 265, 288, 309 and 315 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 284, 291 to 295, 298, 299, 301, 302, 304, 306 to 308, 310 and 311, and b) nucleotide sequences derived from any one of SEQ ID NOs: 284, 291 to 295, 298, 299, 301, 302, 304, 306 to 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307 and 301, and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307 and 301 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301 and 292 and b) nucleotide sequences derived from any one of SEQ ID NOs: SEQ ID NOs: 304, 299, 307, 301 and 292 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292 and 308 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292 and 308 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308 and 306 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308 and 306 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306 and 291 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306 and 291by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291 and 310 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291 and 310 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310 and 311 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311 and 295 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311 and 295 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295 and 293 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295 and 293 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293 and 284 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293 and 284 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284 and 294 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284 and 294 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284, 294 and 302 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307, 301, 292, 308, 306, 291, 310, 311, 295, 293, 284, 294 and 302 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 304, 299, 307 and 301 and b) nucleotide sequences derived from any one of SEQ ID NOs: 304, 299, 307 and 301 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 299, 301, 302, 304, 306 and 307 and b) nucleotide sequences derived from any one of SEQ ID NOs: 299, 301, 302, 304, 306 and 307 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 298, 299, 301, 302, 304, 306, 307 and 308 and b) nucleotide sequences derived from any one of SEQ ID NOs: 298, 299, 301, 302, 304, 306, 307 and 308 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 295, 298, 299, 301, 302, 304, 306, 307, 308 and 310 and b) nucleotide sequences derived from any one of SEQ ID NOs: 295, 298, 299, 301, 302, 304, 306, 307, 308 and 310 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     According to certain embodiments, the present invention provides a rainbow trout egg or spermatozoa, such as an isolated rainbow trout egg or spermatozoa, which comprises within its genome at least one nucleotide sequence selected from the group consisting of a) the nucleotide sequences set forth in SEQ ID NOs: 294, 295, 298, 299, 301, 302, 304, 306, 307, 308, 310 and 311 and b) nucleotide sequences derived from any one of SEQ ID NOs: 294, 295, 298, 299, 301, 302, 304, 306, 307, 308, 310 and 311 by 1 to 5, such as 1 to 2, nucleotide substitutions, provide that said nucleotide substitutions are not at position 31 of said derived sequence. 
     The rainbow trout has late onset of sexual maturation when the nucleotide of the at least one allele is a nucleotide corresponding to the late maturation allele of the respective SNP. The late maturation allele of each SNP is specified in Table 1. 
     The present invention provides in a further aspect a population of rainbow trout egg or spermatozoa, such as an isolated population of rainbow trout egg or spermatozoa, each individual egg or spermatozoa within the population being an egg or spermatozoa according to the fourth aspect of the present invention. 
     According to certain embodiments, the rainbow trout egg is unfertilized. 
     According to certain other embodiments, the rainbow trout egg is fertilized. 
     According to particular embodiments, the rainbow trout egg is an eyed egg. 
     According to certain embodiments, the rainbow trout egg has been isolated from a female rainbow trout of the invention. 
     Oligonucleotide 
     The present invention provides in a fifth aspect, an oligonucleotide, such as an isolated oligonucleotide, comprising at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 164, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 1 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In one embodiment according to the fifth aspect of the present invention, the oligonucleotide has a total length of 8 to 61 nucleotides, such as a total length of 10 to 61 nucleotides, a total length of 12 to 61 nucleotides, a total length of 14 to 61 nucleotides, a total length of 16 to 61 nucleotides, a total length of 18 to 61 nucleotides or a total length of 20 to 61 nucleotides. 
     In another embodiment according to the fifth aspect of the present invention, the oligonucleotide has a total length of 8 to 50 nucleotides, such as a total length of 10 to 50 nucleotides, a total length of 12 to 50 nucleotides, a total length of 14 to 50 nucleotides, a total length of 16 to 50 nucleotides, a total length of 18 to 50 nucleotides or a total length of 20 to 50 nucleotides. 
     In another embodiment according to the fifth aspect of the present invention, the oligonucleotide has a total length of 8 to 40 nucleotides, such as a total length of 10 to 40 nucleotides, a total length of 12 to 40 nucleotides, a total length of 14 to 40 nucleotides, a total length of 16 to 40 nucleotides, a total length of 18 to 40 nucleotides or a total length of 20 to 40 nucleotides. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 1 to 4, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 1 to 4 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 5 to 12, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 5 to 12 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 13 to 16, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 13 to 16 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 17 to 20, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 17 to 20 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 21 to 24, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 21 to 24 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 25 to 28, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 25 to 28 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 29 to 32, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 29 to 32 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 33 to 36, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 33 to 36 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 37 to 40, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 37 to 40 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 41 to 43, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 41 to 43 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 44 to 47, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 44 to 47 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 48 to 50, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 48 to 50 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 51 to 53, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 51 to 53 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 54 to 57, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 54 to 57 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 58 to 61, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 58 to 61 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 62 to 64, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 62 to 64 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 65 to 66, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 65 to 66 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 67 to 69, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 67 to 69 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 70 to 73, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 70 to 73 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 74 to 76, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 74 to 76 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 77 to 80, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 77 to 80 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 81 to 83, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 81 to 83 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 84 to 89, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 84 to 89 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 90 to 92, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 90 to 92 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 93 to 101, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 93 to 101 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 102 to 105, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 102 to 105 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 106 to 109, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 106 to 109 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain particularly preferred embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 110 to 161, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 110 to 161 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 162 to 164, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 162 to 164 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 127, 128, 135, 137, 140, 142, 143, 144, 146 and 147, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 127, 128, 135, 137, 140, 142, 143, 144, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 93, 114, 120, 129, 130, 131, 132, 134, 138 and 148, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 93, 114, 120, 129, 130, 131, 132, 134, 138 and 148 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 5, 24, 86, 121, 125, 126, 133, 141, 150 and 154, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 5, 24, 86, 121, 125, 126, 133, 141, 150 and 154 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 7, 9, 19, 85, 111, 116, 117, 119, 153 and 155, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 7, 9, 19, 85, 111, 116, 117, 119, 153 and 155 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 10, 11, 23, 88, 89, 91, 101, 124, 145 and 151, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 10, 11, 23, 88, 89, 91, 101, 124, 145 and 151 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 120, 127 to 131, 134, 135, 137, 138, 140, 142 to 144, 146 and 147, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 120, 127 to 131, 134, 135, 137, 138, 140, 142 to 144, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143 and 137, or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143 and 137 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137 and 128 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143 and 128 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128 and 144 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 128 and 144 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144 and 142 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144 and 142 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142 and 127 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142 and 127 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127 and 146 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146 and 147 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146 and 147 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147 and 131 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147 and 131 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131 and 129 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131 and 129 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129 and 120 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129 and 120 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120 and 130 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120 and 130 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120, 130 and 138 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143, 137, 128, 144, 142, 127, 146, 147, 131, 129, 120, 130 and 138 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 140, 135, 143 and 137 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 140, 135, 143 and 137 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 135, 137, 138, 140, 142 and 143 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 135, 137, 138, 140, 142 and 143 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 134, 135, 137, 138, 140, 142, 143 and 144 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 134, 135, 137, 138, 140, 142, 143 and 144 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     In certain embodiments, the (isolated) oligonucleotide comprises at least 8 contiguous nucleotides, such as at least 10 contiguous nucleotides, at least 12 contiguous nucleotides, at least 14 contiguous nucleotides, at least 16 contiguous nucleotides, at least 18 contiguous nucleotides or at least 20 contiguous nucleotides, of a nucleotide sequence set forth in any one of SEQ ID NOs: 130, 131, 134, 135, 137, 138, 140, 142, 143, 144, 146 and 147 or a complementary sequence thereof; or of a nucleotide sequence derived from any one of SEQ ID NOs: 131, 134, 135, 137, 138, 140, 142, 143, 144 and 146 by 1 to 5, such as 1 to 2, nucleotide substitutions, or a complementary sequence thereof; with the proviso that that the oligonucleotide includes a late maturation allele or a normal maturation allele, the late maturation allele and the normal maturation allele being alleles of single nucleotide polymorphisms (SNP), the SNP being selected from the SNPs listed in Table 1. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of at least 10 nucleotides, such as at least 16 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of at least 16 nucleotides, such as at least 20 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of at least 20 nucleotides, such as at least 25 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of 10 to 200 nucleotides, such as 10 to 150 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of 10 to 100 nucleotides, such as 10 to 70 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of 16 to 100 nucleotides, such as 16 to 70 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of 10 to 50 nucleotides, such as 10 to 40 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of 16 to 50 nucleotides, such as 16 to 40 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of 10 to 30 nucleotides, such as 8 to 25 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof has a length of 16 to 30 nucleotides, such as 16 to 25 nucleotides. 
     According to certain embodiments, the oligonucleotide or complement thereof is a primer, such as a PCR primer. 
     According to certain embodiments, the oligonucleotide or complement thereof is a probe, such as a hybridization probe. 
     According to certain embodiments, the present invention provides a complement to the oligonucleotide specified above. Such complement may be used as a probe, such as a hybridization probe. 
     A probe or primer according to the present invention may have attached to it a detectable label or reporter molecule. Typical labels include radioactive isotopes, enzyme substrates, co-factors, ligands, chemiluminescent or fluorescent agents, haptens, and enzymes. Methods for labeling and guidance in the choice of labels appropriate for various purposes are discussed, for example, in Sambrook et al. (In Molecular Cloning, A Laboratory Manual, CSHL, New York, 1989) and Ausubel et al. (In Current Protocols in Molecular Biology, John Wiley &amp; Sons, New York, 1998). As a particular example, a probe or primer may include one fluorophor, such as an acceptor fluorophore or donor fluorophor. Such fluorophore may be attached at the 5′- or 3′ end of the probe/primer. 
     Probes are generally at least 15 nucleotides in length, such as at least 16, at least 17, at least 18, at least 19, at least 20, at least 25, at least 30, at least 35, at least 40, at least 45, at least 50, at least 55, at least 60, at least 65, at least 70, or more contiguous nucleotides complementary to the target nucleic acid molecule, such as 20 to 70 nucleotides, 20 to 60 nucleotides, 20 to 50 nucleotides, 20 to 40 nucleotides, or 20 to 30 nucleotides. 
     Primers are typically shorter in length. An oligonucleotide used as primer may be at least 10 nucleotides in length. The specificity of a primer increases with its length. Thus, for example, a primer that includes 30 consecutive nucleotides will anneal to a target sequence with a higher specificity that a corresponding primer of only 15 nucleotides. Thus, to obtain greater specificity, primers of the invention are at least 15 nucleotides in length, such as at least 16, at least 17, at least 18, at least 19, at least 20, at least 25, at least 30, at least 35, at least 40, at least 45, at least 50, at least 55, at least 60, at least 65, at least 70, or more contiguous nucleotides complementary to the target nucleic acid molecule, such as 15 to 70 nucleotides, 15 to 60 nucleotides, 15 to 50 nucleotides, 15 to 40 nucleotides, or 15 to 30 nucleotides. Primer pairs can be used for amplification of nucleic acid sequences, for example, by PCT, real-time-PCR, or other nucleic-acid amplification methods known in the art. 
     Kit 
     The present invention provides in a sixth aspect, a kit for predicting late onset of sexual maturation in rainbow trout, the kit comprising at least one of the oligonucleotides according to the fifth aspect of the present invention. 
     The present invention provides in a seventh aspect, a kit for selecting a rainbow trout having late onset of sexual maturation, the kit comprising at least one of the oligonucleotides according to the fifth aspect of the present invention. 
     Having generally described this invention, a further understanding can be obtained by reference to certain specific examples, which are provided herein for purposes of illustration only, and are not intended to be limiting unless otherwise specified. 
     EXAMPLES 
     Example 1 
     Genome Wide Association Study (GWAS) 
     Dataset: The dataset was acquired from rainbow trout ( Oncorhynchus mykiss ) from AquaGen&#39;s breeding nucleus population of the 2017 year-class. 
     Egg incubation and first feeding: Eggs were fertilized in freshwater in February 2017. The eggs and yolk sac larvae were kept in covered trays with 8 h background light and 16 h darkness, and the mean water temperature was 8° C., until first feeding in April 2017. During egg incubation and first feeding each family group was kept separate, in 250 single tanks. Each first-feeding tank had 80 L, and 1000 fish per family. The fish were fed by a Storvik robot feeder, every 10 minutes 24 h per day. When the mean weight was 2.5 g the number of fish per tank was reduced to 250. All deformed and small fish were culled at this step. 
     Smoltification &amp; vaccination: In June 2017, when the mean fish weight was more than 5 g, 140 fish per family were PIT-tagged (Passive Integrated Transponder, Smartrac Unique 12×2 mm, supplied by RFID-solution). After tagging, fish from different families were mixed in three 6 m 3  tanks, and kept at 24 h light and 13° C. In late July 2017, when the ambient water temperature was 12° C., all fish were transferred to two 50 m 3  outdoor tanks, with ambient light and water temperature. Feeding in these tanks was done by Betten automatic feeders. All fish were manually vaccinated in September 2017, by injection of AlphaJect® 5-3. At this stage all fish were individually weighed, and PIT tag registered. Fish weighing less than 20 g and fish with deviations were culled. 
     Freshwater to Seawater: 25 798 fish with a mean weight of 76 g were transferred to seawater in October 2017. From first feeding until transfer to outdoor tanks the fish were kept at 24 h light and 13° C. The fish were fed commercially available feed for rainbow trout, with nutritional composition and pellet size adjusted to the fish developmental stage and body size. 
     The 25 798 smolts that were transferred to seawater were reared in a 90-meter polar circle cage under ambient light and sea water temperature, similar to national statistics for the Trondheim area. The fish were fed according to feeding tables adjusted for weight, with automatic feeding systems from Aqua. 
     Selection &amp; Identification of control fish: In October 2018, the remaining 25.548 fish were anaesthetised and inspected for deformities, skin colour and body shape. Weight and length were measured, and a photo stored for each fish. The purpose of the measurements was to calculate breeding values and select the best fish to be parents for a new generation trout. 
     At this stage, the 30% biggest trout, with the silveriest skin colour, were kept as breeding candidates. 7 000 fish were pre-selected, at a mean weight of 3.93 kg. The culled ones, 18 547 fish, had a mean weight of 3.00 kg. These fish were transported to a processing plant for harvest during October 2018. 
     At the processing plant, a group of 1 526 non-maturing fish (average weight of 3.1 kg), with maximum 7 fish per 250 family, were selected for registration of yield and analysis of pigment and DNA. The gender of these fish was determined when the fish were opened, and the gonads could be seen. This group of fish was considered the control group, i.e. non-maturing fish with known gender. 
     Seawater to freshwater for spawning: 6900 fish of the pre-selected fish, with a mean weight of 5.11 kg, were sorted again in December 2018. All fish with a tendency for maturation were selected. Maturation is more difficult to determine in fish less than 2 years of age, as they have a silverier skin colour and less clear external sexual maturation characteristics. 1 748 fish were selected as broodstock candidates, and tissue samples were taken from all these fish for DNA analysis. 
     The selected fish were transferred to freshwater in AquaGen&#39;s broodstock facilities in Kyrksæterøra, Norway, for spawning in February 2019. Included in this group were also fish of the type neo-males, which are sex-reversed so that genetic females develop masculine reproductive organs and produce sperm. This is done by administering testosterone to the fish at the time when the reproductive organs are first developed (at first feeding, around 0.15 g) (Biotechnology &amp; Biotechnological Equipment, 2009, 23:4, 1509-1514, DOI: 10.2478/V10133-009-0002-X). The change in hormone levels stimulates females to develop male gonads, and they develop male external characteristics when they mature 2 years later. All neo-males were genotyped in advance to ensure that only genetic females were present. 
     During the last period in freshwater (1 month) around 60% of the fish went through final maturation and became ready to spawn. Fish that did not go through final maturation, were culled. The gender of these fish was not checked. For the fish that went through final maturation, gender was determined by whether the spawning product was eggs (from females) or sperm (from males). The fish were kept at a 24 h light, and the mean water temperature was 10° C. during the month in freshwater. 
     Genotyping: As already described above, tissue samples were taken from control fish and broodstock candidates. All tissue samples were taken from the tail fin of the fish for DNA-extraction and genotyping. DNA was extracted from the tissue samples, using a standard method (DNAdvance™ kit from Beckman Coulter, Indianapolis, United States). The DNA was genotyped using the Axiom® Trout genotyping Array, a SNP-chip harbouring 57,501 single nucleotide polymorphisms (SNPs) in 96-well format. This SNP-chip was developed by AquaGen in collaboration with the United Stated Department of Agriculture 20 (USDA) and Affymetrix, and is commercially available from Affymetrix (San Diego, USA). Genotyping was performed using the Affymetrix&#39; proprietary Axiom® platform, following the Axiom® 2.0 Assay Automated Workflow User Guide. 
     Based on the raw data provided by the Axiom® machinery, genotypes were called using the Affymetrix PowerTools software. The analysis and interpretation of the raw data was done according to the Best Practices 30 Workflow provided by Affymetrix. SNPs and animals having quality parameters below the default thresholds, provided in the Best Practices Workflow, were not considered for further analyses. 
     Only markers in the PolyHighResolution and NoMinorHom categories were analysed, as recommended by the genotyping platform producer. SNP markers with missingness above 10% and animals with missing data above 15% was removed, resulting in a dataset of 42,314 SNP markers and 1,647 fish with recorded maturation status and sex. 
     GWAS: To search for DNA markers associated with late maturation in the sampled rainbow trout population, a single marker genome-wide association study was conducted using genotypes for 1 647 fish with recorded maturation status. Sex was included as a covariate in the analysis. The association study was conducted with the GCTA software (Yang et al., 2011, Am J Hum Genet. 2011 88(1): 76-82) which implements a mixed linear model single-marker association analysis using the -mlma-loco option of GCTA (Yang et al., 2014, Nat Genet. February; 46(2):100-6). Sex was included as a covariate in the analysis using the covar parameter. The model fitted to the performance information for each trait and each SNP was: 
       LATEMAT= a+bx+g−+e    
     where LATEMAT is the maturation status of the fish coded as 0 or 1 (0=immature; 1=mature), a is the mean term, b is the fixed additive effect of the candidate SNP to be tested for association, x is the SNP genotype indicator variable coded as 0, 1 or 2, g− is the random polygenic effect, i.e. the accumulated effect of all SNPs except those on the chromosome where the candidate SNP is located, and e is the residual. The var(g−) is re-estimated each time when a chromosome is excluded from calculating the genomic relationship matrix. 
     The results of the genome-wide association study are presented in table 1 and table 3. 
     
       
         
           
               
               
               
               
               
             
               
                   
                 TABLE 3 
               
               
                   
                   
               
               
                   
                 SNP # 
                 b 
                 p 
                 se 
               
               
                   
                   
               
             
            
               
                   
               
            
           
           
               
               
               
               
               
            
               
                   
                 1 
                 0.072621 
                 0.000116 
                 0.018841 
               
               
                   
                 2 
                 0.066487 
                 0.000363 
                 0.018646 
               
               
                   
                 3 
                 −0.0733 
                 0.000143 
                 0.019278 
               
               
                   
                 4 
                 0.069419 
                 0.000374 
                 0.01951 
               
               
                   
                 5 
                 0.232912 
                 1.39E−06 
                 0.048258 
               
               
                   
                 6 
                 −0.20478 
                 1.21E−05 
                 0.046799 
               
               
                   
                 7 
                 −0.0878 
                 3.16E−06 
                 0.018842 
               
               
                   
                 8 
                 0.155719 
                 2.26E−05 
                 0.036752 
               
               
                   
                 9 
                 0.20537 
                 3.29E−06 
                 0.044148 
               
               
                   
                 10 
                 −0.19734 
                 9.78E−06 
                 0.044628 
               
               
                   
                 11 
                 −0.19984 
                 7.04E−06 
                 0.044482 
               
               
                   
                 12 
                 0.19337 
                 1.58E−05 
                 0.044785 
               
               
                   
                 13 
                 0.053904 
                 0.006195 
                 0.019693 
               
               
                   
                 14 
                 0.066758 
                 0.006195 
                 0.024389 
               
               
                   
                 15 
                 −0.07529 
                 0.001926 
                 0.024278 
               
               
                   
                 16 
                 0.062764 
                 0.003561 
                 0.021534 
               
               
                   
                 17 
                 −0.09764 
                 3.73E−05 
                 0.023679 
               
               
                   
                 18 
                 −0.09032 
                 2.55E−05 
                 0.021452 
               
               
                   
                 19 
                 0.125808 
                 4.33E−06 
                 0.02738 
               
               
                   
                 20 
                 −0.07647 
                 0.000213 
                 0.020651 
               
               
                   
                 21 
                 0.115704 
                 1.72E−05 
                 0.026915 
               
               
                   
                 22 
                 0.111758 
                 1.22E−05 
                 0.025546 
               
               
                   
                 23 
                 0.091821 
                 6.78E−06 
                 0.020402 
               
               
                   
                 24 
                 −0.0919 
                 1.31E−06 
                 0.018996 
               
               
                   
                 25 
                 −0.09971 
                 0.005334 
                 0.035786 
               
               
                   
                 26 
                 −0.05523 
                 0.006552 
                 0.020316 
               
               
                   
                 27 
                 0.126665 
                 0.00635 
                 0.046413 
               
               
                   
                 28 
                 0.05894 
                 0.001665 
                 0.018745 
               
               
                   
                 29 
                 −0.1057 
                 0.000305 
                 0.029273 
               
               
                   
                 30 
                 0.147067 
                 0.00117 
                 0.045304 
               
               
                   
                 31 
                 −0.16004 
                 0.000784 
                 0.047657 
               
               
                   
                 32 
                 −0.10704 
                 0.000775 
                 0.031843 
               
               
                   
                 33 
                 0.115041 
                 0.000899 
                 0.034647 
               
               
                   
                 34 
                 0.12055 
                 0.000502 
                 0.034646 
               
               
                   
                 35 
                 0.119566 
                 0.000526 
                 0.034488 
               
               
                   
                 36 
                 −0.08267 
                 0.000725 
                 0.024459 
               
               
                   
                 37 
                 0.108508 
                 0.002291 
                 0.03558 
               
               
                   
                 38 
                 0.110396 
                 0.001872 
                 0.035499 
               
               
                   
                 39 
                 0.108853 
                 0.002157 
                 0.035483 
               
               
                   
                 40 
                 0.110637 
                 0.002715 
                 0.0369 
               
               
                   
                 41 
                 −0.14723 
                 0.000184 
                 0.039366 
               
               
                   
                 42 
                 0.070596 
                 0.000442 
                 0.020092 
               
               
                   
                 43 
                 0.069851 
                 0.00041 
                 0.019768 
               
               
                   
                 44 
                 −0.06207 
                 0.000744 
                 0.018403 
               
               
                   
                 45 
                 −0.07639 
                 0.000417 
                 0.021643 
               
               
                   
                 46 
                 0.073182 
                 0.000715 
                 0.021626 
               
               
                   
                 47 
                 −0.06458 
                 0.000603 
                 0.018827 
               
               
                   
                 48 
                 −0.13404 
                 0.003149 
                 0.045395 
               
               
                   
                 49 
                 −0.10799 
                 0.003762 
                 0.037271 
               
               
                   
                 50 
                 0.056679 
                 0.006409 
                 0.020791 
               
               
                   
                 51 
                 −0.06892 
                 0.000714 
                 0.020366 
               
               
                   
                 52 
                 −0.14191 
                 0.000392 
                 0.040027 
               
               
                   
                 53 
                 −0.05961 
                 0.001254 
                 0.018476 
               
               
                   
                 54 
                 −0.06682 
                 0.000265 
                 0.018321 
               
               
                   
                 55 
                 −0.06498 
                 0.000317 
                 0.018043 
               
               
                   
                 56 
                 0.127081 
                 0.00027 
                 0.034891 
               
               
                   
                 57 
                 −0.11243 
                 0.000252 
                 0.030712 
               
               
                   
                 58 
                 0.159429 
                 0.000285 
                 0.043937 
               
               
                   
                 59 
                 0.069496 
                 0.000553 
                 0.020121 
               
               
                   
                 60 
                 −0.09543 
                 0.000307 
                 0.026438 
               
               
                   
                 61 
                 −0.08153 
                 0.000847 
                 0.02443 
               
               
                   
                 62 
                 0.05459 
                 0.002727 
                 0.018216 
               
               
                   
                 63 
                 0.072264 
                 0.003723 
                 0.024912 
               
               
                   
                 64 
                 0.09553 
                 0.003666 
                 0.032879 
               
               
                   
                 65 
                 0.061393 
                 0.001841 
                 0.019711 
               
               
                   
                 66 
                 −0.06519 
                 0.001032 
                 0.019865 
               
               
                   
                 67 
                 0.063174 
                 0.000705 
                 0.018649 
               
               
                   
                 68 
                 −0.06968 
                 0.000576 
                 0.020241 
               
               
                   
                 69 
                 −0.07971 
                 0.00024 
                 0.021706 
               
               
                   
                 70 
                 −0.08226 
                 0.000214 
                 0.02222 
               
               
                   
                 71 
                 0.099409 
                 1.23E−05 
                 0.022736 
               
               
                   
                 72 
                 −0.08576 
                 0.000127 
                 0.022383 
               
               
                   
                 73 
                 −0.08458 
                 0.000187 
                 0.022635 
               
               
                   
                 74 
                 −0.08047 
                 0.000126 
                 0.020985 
               
               
                   
                 75 
                 0.064418 
                 0.000812 
                 0.019236 
               
               
                   
                 76 
                 0.063253 
                 0.000951 
                 0.019141 
               
               
                   
                 77 
                 0.044405 
                 0.021796 
                 0.019358 
               
               
                   
                 78 
                 0.066171 
                 0.011509 
                 0.026187 
               
               
                   
                 79 
                 −0.04933 
                 0.015787 
                 0.020436 
               
               
                   
                 80 
                 0.042566 
                 0.024577 
                 0.018935 
               
               
                   
                 81 
                 0.060481 
                 0.014357 
                 0.024704 
               
               
                   
                 82 
                 −0.09819 
                 0.011979 
                 0.039075 
               
               
                   
                 83 
                 0.110566 
                 0.009111 
                 0.042397 
               
               
                   
                 84 
                 0.111622 
                 1.93E−05 
                 0.026122 
               
               
                   
                 85 
                 0.105807 
                 3.43E−06 
                 0.022787 
               
               
                   
                 86 
                 0.104419 
                 2.28E−06 
                 0.022091 
               
               
                   
                 87 
                 −0.09058 
                 1.21E−05 
                 0.020704 
               
               
                   
                 88 
                 −0.11374 
                 1.08E−05 
                 0.025847 
               
               
                   
                 89 
                 0.096867 
                  9.5E−06 
                 0.021875 
               
               
                   
                 90 
                 0.104326 
                 5.27E−05 
                 0.025802 
               
               
                   
                 91 
                 0.108363 
                   1E−05 
                 0.024537 
               
               
                   
                 92 
                 0.105876 
                  1.7E−05 
                 0.024617 
               
               
                   
                 93 
                 −0.1047 
                 1.33E−07 
                 0.019847 
               
               
                   
                 94 
                 0.079676 
                 2.82E−05 
                 0.019027 
               
               
                   
                 95 
                 0.077646 
                 4.61E−05 
                 0.019056 
               
               
                   
                 96 
                 −0.0792 
                 3.17E−05 
                 0.019034 
               
               
                   
                 97 
                 0.120882 
                 4.93E−05 
                 0.029782 
               
               
                   
                 98 
                 −0.12623 
                 2.36E−05 
                 0.029858 
               
               
                   
                 99 
                 0.123603 
                 3.31E−05 
                 0.029775 
               
               
                   
                 100 
                 −0.12114 
                 4.72E−05 
                 0.02977 
               
               
                   
                 101 
                 −0.13242 
                 9.56E−06 
                 0.029912 
               
               
                   
                 102 
                 0.075256 
                 0.010179 
                 0.029286 
               
               
                   
                 103 
                 0.043721 
                 0.014083 
                 0.017808 
               
               
                   
                 104 
                 −0.065 
                 0.004754 
                 0.023025 
               
               
                   
                 105 
                 −0.04499 
                 0.016455 
                 0.018755 
               
               
                   
                 106 
                 −0.07967 
                 0.000102 
                 0.020501 
               
               
                   
                 107 
                 −0.07302 
                 9.19E−05 
                 0.018671 
               
               
                   
                 108 
                 −0.0709 
                 0.000138 
                 0.018599 
               
               
                   
                 109 
                 −0.07402 
                 0.00024 
                 0.020152 
               
               
                   
                 110 
                 −0.09654 
                 4.21E−05 
                 0.023571 
               
               
                   
                 111 
                 −0.08881 
                 4.36E−06 
                 0.019335 
               
               
                   
                 112 
                 −0.09654 
                 4.21E−05 
                 0.023571 
               
               
                   
                 113 
                 0.09124 
                 2.02E−05 
                 0.021402 
               
               
                   
                 114 
                 0.094936 
                  3.3E−07 
                 0.018595 
               
               
                   
                 115 
                 0.081369 
                 1.22E−05 
                 0.018605 
               
               
                   
                 116 
                 −0.08526 
                  2.5E−06 
                 0.018109 
               
               
                   
                 117 
                 0.081366 
                 4.64E−06 
                 0.017763 
               
               
                   
                 118 
                 0.07669 
                 1.78E−05 
                 0.017876 
               
               
                   
                 119 
                 0.104425 
                 2.77E−06 
                 0.02228 
               
               
                   
                 120 
                 −0.1101 
                 1.97E−07 
                 0.021165 
               
               
                   
                 121 
                 0.090614 
                 5.93E−07 
                 0.018147 
               
               
                   
                 122 
                 −0.07324 
                 4.21E−05 
                 0.017881 
               
               
                   
                 123 
                 −0.07727 
                 1.56E−05 
                 0.017885 
               
               
                   
                 124 
                 0.162104 
                  6.3E−06 
                 0.035896 
               
               
                   
                 125 
                 −0.08414 
                 1.63E−06 
                 0.017546 
               
               
                   
                 126 
                 −0.09229 
                 8.96E−07 
                 0.018783 
               
               
                   
                 127 
                 0.112832 
                  9.8E−08 
                 0.021168 
               
               
                   
                 128 
                 −0.1145 
                  6.4E−08 
                 0.021176 
               
               
                   
                 129 
                 −0.11025 
                 1.84E−07 
                 0.02114 
               
               
                   
                 130 
                 −0.10958 
                 2.43E−07 
                 0.021225 
               
               
                   
                 131 
                 0.111905 
                 1.32E−07 
                 0.021211 
               
               
                   
                 132 
                 0.093353 
                 3.85E−07 
                 0.018391 
               
               
                   
                 133 
                 0.088411 
                 1.81E−06 
                 0.018519 
               
               
                   
                 134 
                 0.09229 
                 3.12E−07 
                 0.018039 
               
               
                   
                 135 
                 −0.1046 
                 1.68E−08 
                 0.018538 
               
               
                   
                 136 
                 −0.08991 
                 1.52E−05 
                 0.020784 
               
               
                   
                 137 
                 0.10297 
                 2.56E−08 
                 0.018489 
               
               
                   
                 138 
                 −0.09344 
                 3.12E−07 
                 0.018263 
               
               
                   
                 139 
                 −0.08991 
                 1.52E−05 
                 0.020784 
               
               
                   
                 140 
                 0.112587 
                 6.95E−10 
                 0.018256 
               
               
                   
                 141 
                 −0.09459 
                 5.24E−07 
                 0.018852 
               
               
                   
                 142 
                 0.096872 
                 8.73E−08 
                 0.018103 
               
               
                   
                 143 
                 0.101851 
                 1.87E−08 
                 0.018112 
               
               
                   
                 144 
                 0.096511 
                 8.44E−08 
                 0.018014 
               
               
                   
                 145 
                 0.080036 
                 5.37E−06 
                 0.017591 
               
               
                   
                 146 
                 0.117443 
                 1.18E−07 
                 0.022174 
               
               
                   
                 147 
                 0.084425 
                  1.2E−07 
                 0.015949 
               
               
                   
                 148 
                 −0.11203 
                 4.33E−07 
                 0.022169 
               
               
                   
                 149 
                 −0.09458 
                 2.84E−05 
                 0.022593 
               
               
                   
                 150 
                 0.106663 
                   1E−06 
                 0.021805 
               
               
                   
                 151 
                 −0.08603 
                 1.17E−05 
                 0.019631 
               
               
                   
                 152 
                 0.08138 
                 2.35E−05 
                 0.019247 
               
               
                   
                 153 
                 −0.09825 
                 2.32E−06 
                 0.0208 
               
               
                   
                 154 
                 −0.10695 
                 6.85E−07 
                 0.021539 
               
               
                   
                 155 
                 0.10169 
                 2.53E−06 
                 0.021609 
               
               
                   
                 156 
                 0.096217 
                 2.48E−05 
                 0.022817 
               
               
                   
                 157 
                 0.097621 
                 1.77E−05 
                 0.022743 
               
               
                   
                 158 
                 −0.09713 
                 1.93E−05 
                 0.022735 
               
               
                   
                 159 
                 −0.09638 
                 2.24E−05 
                 0.022737 
               
               
                   
                 160 
                 −0.09391 
                 3.63E−05 
                 0.02274 
               
               
                   
                 161 
                 −0.09331 
                 2.82E−05 
                 0.022282 
               
               
                   
                 162 
                 −0.10438 
                 0.008011 
                 0.039364 
               
               
                   
                 163 
                 0.132686 
                 0.008268 
                 0.050242 
               
               
                   
                 164 
                 0.051735 
                 0.01064 
                 0.020254 
               
               
                   
                   
               
               
                   
                 b (also referred to as the beta value) is also called the allele substitution effect and refers to the estimated effect of receiving one copy of the tested allele (the “normal maturation allele” in table 1). Reference is e.g. made to SNP #140, wherein the mean effect on the phenotype of going from genotype TT to TC is 0.112587 while going from TT to CC is 0.112587 X2 = 0.225174. The phenotype in the present case is maturation status coded as 0 when immature and 1 when mature, the effect of 0.112587 translates to an 11 percentage point increase in percentage of mature fish in the data set. 
               
               
                   
                 p measure of statistical evidence against a null hypothesis. A smaller p-value indicates stronger evidence substantiating the alternative hypothesis. In practice, the p-value indicates how likely a putative characteristic associated variant is due to random chance. High beta value and low se value will provide a low p-value, while low beta value and high se value will provide a high p-value. 
               
               
                   
                 se represents the standard error which depends on the sample size and how well the observed data fits the linear model. 
               
            
           
         
       
     
     Linkage Disequilibrium (LD) 
     LD or, more precisely, gametic phase linkage disequilibrium, is used in order to describe the co-inheritance of alleles at genetically linked polymorphisms, at the population level. Two polymorphisms are defined to be in strong LD if the square of the correlation coefficient between the two loci (r2, the most commonly used measure of LD) is equal to or larger than 0.5, more preferably equal to or larger than 0.7, even more preferably equal to or larger than 0.8 and most preferably equal to or larger than 0.9. 
     A distinctive and highly significant QTL for maturity has been detected on chromosome 28 (see  FIGS. 1 a  and 1 b   ), and among these are 16 SNPs of particular interest (SNP #120, 127-131, 134-135, 137-138, 140, 142-144 and 146-147). All 16 of the SNPs are located within a genomic region of 1.9 megabases ( FIG. 1 b   ), and all show varying degree of elevated squared correlations to each other ( FIG. 3 ). 
     Table 4 provides a list of the 16 SNPs on chromosome 28 that are of particular interest and their position on the chromosome. Further, the list is sorted according to the square of the correlation coefficient between two loci. 
                                     TABLE 4               A_SNP #   A_BP #   B_SNP #   B_BP #   R2                                                    135   11414120   137   11414399   0.996177       128   11093692   129   11096202   0.995723       120   10378833   129   11096202   0.990043       120   10378833   128   11093692   0.988618       142   11667915   143   11704242   0.980364       143   11704242   144   11745469   0.979454       142   11667915   144   11745469   0.976509       129   11096202   131   11148320   0.97439       128   11093692   131   11148320   0.974313       120   10378833   131   11148320   0.967308       134   11382381   138   11487704   0.961402       128   11093692   130   11139401   0.934455       129   11096202   130   11139401   0.934442       120   10378833   130   11139401   0.927666       130   11139401   131   11148320   0.923362       138   11487704   140   11537736   0.870889       127   10605800   130   11139401   0.854619       134   11382381   140   11537736   0.841967       127   10605800   128   11093692   0.82477       127   10605800   129   11096202   0.824571       120   10378833   127   10605800   0.818416       127   10605800   131   11148320   0.813752       131   11148320   146   12063041   0.801846       134   11382381   137   11414399   0.776771       134   11382381   135   11414120   0.775718       137   11414399   138   11487704   0.755322       135   11414120   138   11487704   0.754192       137   11414399   140   11537736   0.640884       135   11414120   140   11537736   0.638918       134   11382381   147   12252318   0.63764       138   11487704   147   12252318   0.629467       137   11414399   142   11667915   0.571017       135   11414120   142   11667915   0.569716       137   11414399   143   11704242   0.568289       137   11414399   144   11745469   0.567879       135   11414120   143   11704242   0.566992       135   11414120   144   11745469   0.566587       137   11414399   147   12252318   0.555781       135   11414120   147   12252318   0.553991       140   11537736   147   12252318   0.547934       140   11537736   143   11704242   0.458642       140   11537736   144   11745469   0.456815       140   11537736   142   11667915   0.456404       134   11382381   143   11704242   0.439942       134   11382381   142   11667915   0.439518       134   11382381   144   11745469   0.43747       138   11487704   142   11667915   0.428019       138   11487704   144   11745469   0.425464       138   11487704   143   11704242   0.425421       129   11096202   143   11704242   0.399344       129   11096202   142   11667915   0.397379       128   11093692   142   11667915   0.395977       131   11148320   143   11704242   0.390864       131   11148320   142   11667915   0.388907       131   11148320   144   11745469   0.387393       130   11139401   143   11704242   0.362932       130   11139401   142   11667915   0.360975       130   11139401   144   11745469   0.359626       143   11704242   146   12063041   0.316434       144   11745469   147   12252318   0.3161       144   11745469   146   12063041   0.315437       142   11667915   146   12063041   0.313873       143   11704242   147   12252318   0.313531       142   11667915   147   12252318   0.312988       130   11139401   137   11414399   0.227957       130   11139401   135   11414120   0.227507       127   10605800   137   11414399   0.226163       127   10605800   135   11414120   0.225741       129   11096202   137   11414399   0.219178       128   11093692   137   11414399   0.219139       129   11096202   135   11414120   0.218713       128   11093692   135   11414120   0.218672       131   11148320   137   11414399   0.216527       131   11148320   135   11414120   0.216046       127   10605800   140   11537736   0.211376       127   10605800   134   11382381   0.192605       146   12063041   147   12252318   0.191637       130   11139401   140   11537736   0.188975       127   10605800   138   11487704   0.185399       128   11093692   140   11537736   0.182801       129   11096202   140   11537736   0.182682       131   11148320   140   11537736   0.179886       130   11139401   134   11382381   0.17262       130   11139401   138   11487704   0.166937       128   11093692   134   11382381   0.16544       129   11096202   134   11382381   0.16483       131   11148320   134   11382381   0.163749       137   11414399   146   12063041   0.160392       135   11414120   146   12063041   0.160383       128   11093692   138   11487704   0.159903       129   11096202   138   11487704   0.159742       131   11148320   138   11487704   0.158182       140   11537736   146   12063041   0.133768       138   11487704   146   12063041   0.115124       134   11382381   146   12063041   0.114512               A_SNP # a first SNP, wherein the number refers to the SNP # according to table 1.       A_BP # position of the A_SNP # (first SNP) on chromosome 28.       B_SNP # a second SNP, wherein the number refers to the SNP # according to table 1.       B_BP # position of the B_SNP # (second SNP) on chromosome 28.       R2 correlation coefficient between A_SNP # and B_SNP #.            
TABLE 5A AND TABLE 5B PROVIDES A LIST OF THE 16 SNPS ON CHROMOSOME 28 THAT ARE OF PARTICULAR INTEREST, THEIR POSITION ALONG THE CHROMOSOME AND THEIR P-VALUE. TABLE 5A IS SORTED ACCORDING TO THE POSITION OF THE SNPS ALONG CHROMOSOME 28 WHILE TABLE 5B IS SORTED ACCORDING TO THE P-VALUE OF THE SNPS.
 
     
       
         
           
               
               
               
             
               
                 TABLE 5a 
               
               
                   
               
               
                 SNP # 
                 BP # 
                 p 
               
               
                   
               
             
            
               
                 120 
                 10378833 
                 1.97073E−07 
               
               
                 127 
                 10605800 
                 9.79982E−08 
               
               
                 128 
                 11093692 
                 6.40093E−08 
               
               
                 129 
                 11096202 
                 1.83833E−07 
               
               
                 130 
                 11139401 
                 2.43284E−07 
               
               
                 131 
                 11148320 
                 1.32183E−07 
               
               
                 134 
                 11382381 
                 3.11917E−07 
               
               
                 135 
                 11414120 
                 1.67786E−08 
               
               
                 137 
                 11414399 
                 2.55833E−08 
               
               
                 138 
                 11487704 
                 3.11633E−07 
               
               
                 140 
                 11537736 
                 6.95304E−10 
               
               
                 142 
                 11667915 
                  8.7344E−08 
               
               
                 143 
                 11704242 
                 1.87124E−08 
               
               
                 144 
                 11745469 
                  8.4426E−08 
               
               
                 146 
                 12063041 
                 1.18044E−07 
               
               
                 147 
                 12252318 
                 1.20003E−07 
               
               
                   
               
            
           
         
       
     
     
       
         
           
               
               
               
             
               
                 TABLE 5b 
               
               
                   
               
               
                 SNP # 
                 BP # 
                 p 
               
               
                   
               
             
            
               
                 140 
                 11537736 
                 6.95304E−10 
               
               
                 135 
                 11414120 
                 1.67786E−08 
               
               
                 143 
                 11704242 
                 1.87124E−08 
               
               
                 137 
                 11414399 
                 2.55833E−08 
               
               
                 128 
                 11093692 
                 6.40093E−08 
               
               
                 144 
                 11745469 
                  8.4426E−08 
               
               
                 142 
                 11667915 
                  8.7344E−08 
               
               
                 127 
                 10605800 
                 9.79982E−08 
               
               
                 146 
                 12063041 
                 1.18044E−07 
               
               
                 147 
                 12252318 
                 1.20003E−07 
               
               
                 131 
                 11148320 
                 1.32183E−07 
               
               
                 129 
                 11096202 
                 1.83833E−07 
               
               
                 120 
                 10378833 
                 1.97073E−07 
               
               
                 130 
                 11139401 
                 2.43284E−07 
               
               
                 138 
                 11487704 
                 3.11633E−07 
               
               
                 134 
                 11382381 
                 3.11917E−07 
               
               
                   
               
            
           
         
       
         
         SNP# the number refers to the SNP # according to table 1. 
         BP# position of the SNP along chromosome 28. 
         p measure of statistical evidence against a null hypothesis. A smaller p-value indicates stronger evidence substantiating the alternative hypothesis. In practice, the p-value indicates how likely a putative characteristic associated variant is due to random chance. High beta value and low se value will provide a low p-value, while low beta value and high se value will provide a high p-value.