Dryptosaurus

*Dryptosaurus Aquilunguis*

***Dryptosaurus*** (dryp·to·saur·us/pronounced drip-toe-SORE-us) (from Greek ***drypto/δρύπτω***, "to tear" and ***sauros/σαυρος*** "lizard" ) was a genus of primitive tyrannosaur that lived in Eastern North America during the middle Maastrichtian stage of the Late Cretaceous period, around 67 million years ago.

Although largely unknown now outside of academic circles, a famous painting of the genus by Charles R. Knight made it one of the more widely known dinosaurs of its time, in spite of its poor fossil record. Its specific name aquilunguis is Latin for "having claws like an eagle's".

## Description

Dryptosaurus is estimated to have been 7.5 metres (25 ft) long and to have weighed 1.5 to 1.7 tons, although this is based on partial remains of one individual. Like its relative Eotyrannus, Dryptosaurus seems to have had relatively long arms compared to, for example, Tyrannosaurus, and the hands are believed to have had three fingers. Each of these fingers was tipped by a talonlike 8 inch claw. These claws lend a meaning for the type species aquilunguis: eagle-clawed.

## Discovery and species

In 1866, an incomplete skeleton (ANSP 9995) was found in New Jersey by quarry workers in rock belonging to the upper part of the New Egypt Formation. Paleontologist E.D. Cope described the remains, naming the creature "Lælaps" ("storm wind", after the dog in Greek mythology that never failed to catch what it was hunting). Lælaps became one of the first dinosaurs described from North America (following Hadrosaurus, Aublysodon and Trachodon). Subsequently, it was discovered that the name Lælaps had already been given to a genus of mite, and Cope's lifelong rival O.C. Marsh changed the name in 1877 to Dryptosaurus.

During the 19th century, numerous theropod species in North America were assigned to the genus Dryptosaurus (often as Lælaps or Laelaps), only to be reclassified. Only one potential second species is sometimes recognized today, D. macropus, known from a partial hind limb found in the Navesink Formation (probably roughly equivalent to the New Eagle Formation in time). Joseph Leidy originally referred the specimen to Coelosaurus.

Cope later recognized it as a distinct species, referring to it as Lælaps macropus, differing from the specimen now known as Dryptosaurus by its longer toes. Subsequently, most scientists had concluded that it probably belonged to Coelosaurus after all, but in 2004 paleontologist Thomas R. Holtz listed it as an indeterminate tyrannosauroid, possibly referable to the genus Dryptosaurus.

## Ecology

Although certainly a carnivore, the paucity of known Cretaceous East Coast dinosaurs make ascertaining the specific diet of Dryptosaurus difficult. Hadrosaurids are known from the same time and place as Dryptosaurus, the island continent of Appalachia, and they may have been a prominent part of its diet. Nodosaurs were also present, although less likely to be hunted due to their armor plating.

## Classification

Dryptosaurus was the only large carnivore known in eastern North America before the discovery of Appalachiosaurus in 2005.[5] During that time Dryptosaurus was classified in a number of theropod families. Originally considered a megalosaurid by Cope, it was later assigned to its own family (Dryptosauridae) by Marsh, and later found (through phylogenetic studies of the 1990s) to be a coelurosaur, though its exact placement within that group remained uncertain. The discovery of the closely related (and more complete) Appalachiosaurus made it clear that Dryptosaurus was a primitive tyrannosauroid.

The fossil material assigned to Dryptosaurus was reviewed by Ken Carpenter in 1997 in light of the many different theropods discovered since Cope's day. He felt that due to some unusual features it couldn't be placed in any existing family and warranted placement in its own family, Dryptosauridae.

### Dubious species:

    Numerous species have been assigned to Dryptosaurus based on very fragmentary material, and probably belong to various other types of theropod. These include:

* D. macropus (Cope, 1868)
* D. falculus (Cope, 1876)
* D. hazenianus (Cope, 1876)

Another dubious species was Laelaps trihedrodon by Cope in 1877. Although the type specimen included a partial dentary, all material—except for a collection of five damaged partial tooth crowns (AMNH 5780)—has been lost. AMNH 5780 has many features in common with Allosaurus and is probably referable to that genus. However some of the Allosaurus-like characters of the tooth are primitive to theropods as a whole and may have been present in the less-studied or poorly preserved [Morrison Formation theropod species. Consequently the synonymization of L. trihedrodon with Allosaurus is tentative, despite its high likelihood.

## JPInstitute.com Description

Edward Drinker Cope discovered Dryptosaurus in 1866. His find was of a poorly preserved partial skeleton, which may not have been fully-grown.

The relationship of Dryptosaurus to other carnivorous dinosaurs is the subject of great debate, but it is clear Dryptosaurus was a large, meat-eating beast that walked on two legs and lived near the end of the age of dinosaurs.

## Dinosaur Field Guide Description

Dryptosaurus ("tearing lizard") has an important place in the history of dinosaur science. Before this species was discovered in 1866, paleontologists had never found the arms and legs of the same species of a meat-eating dinosaur: Because of this, they thought that meat-eaters were four-legged hunters, like giant lizards or bears. However, when Dryptosaurus was found, scientists saw that meat-eating dinosaurs had arms that were much shorter than their legs, and realized they had to be bipedal, or two-footed. Dryptosaurus was named for its enormous 8-inch (21-cm) claw. Its discoverer, Edward Drinker Cope, thought that this claw came from the foot, but it was later realized that this was actually a hand claw. Unfortunately, not much of the skeleton of Dryptosaurus was ever found. Paleontologists must guess at what it may have looked like.

### Fun Facts

famous painting by artist Charles R. Knight shows two Dryptosaurus fighting, with one of them leaping high in the air to pounce on the ather This is an exciting image but an unlikely scenario for animals af their size!

### Trivia

Dryptosaurus Was named selene It wee chen discovered that the name had already been used far a type of trick!

## Links

http://web.archive.org/web/20040214164709fw\_/http://www.jpinstitute.com/dinopedia/dinocards/dc\_drypt.html

## References

* http://www.smithsonianmag.com/science-nature/how-dryptosaurus-got-its-name-68864150/?no-ist
* https://digitallibrary.amnh.org/handle/2246/6117
* https://www.researchgate.net/publication/281043562\_The\_Anatomy\_of\_Dryptosaurus\_aquilunguis\_Dinosauria\_Theropoda\_and\_a\_Review\_of\_Its\_Tyrannosauroid\_Affinities
* https://web.archive.org/web/20070310231340/http://www.vertpaleo.org/jvp/17-561-573.html
* https://www.tandfonline.com/doi/abs/10.1080/00222936708679723
* https://archive.org/details/mesozoicvertebra0000unse/page/10/mode/2up
* https://www.gbif.org/dataset/13c8f8f0-3ea5-4a40-afb1-5292f8380faa
* https://www.researchgate.net/publication/318485947\_Teihivenator\_gen\_nov\_a\_new\_generic\_name\_for\_the\_Tyrannosauroid\_Dinosaur\_Laelaps\_macropus\_Cope\_1868\_preoccupied\_by\_Koch\_1836
* https://www.biodiversitylibrary.org/part/238885
* https://repository.si.edu/handle/10088/22800
* https://digital.lib.ecu.edu/ncpi/view/10005
* https://www.researchgate.net/publication/233904673\_A\_New\_Genus\_And\_Species\_Of\_Tyrannosauroid\_From\_The\_Late\_CretaceousMiddle\_Campanian\_Demopolis\_Formation\_Of\_Alabama
* https://www.app.pan.pl/article/item/app20110141.html
* https://dx.doi.org/10.1098/rsos.210127
* https://www.jstor.org/stable/1300989
* https://www.researchgate.net/publication/232690303\_Landman\_N\_H\_R\_O\_Johnson\_and\_L\_E\_Edwards\_Cephalopods\_from\_the\_CretaceousTertiary\_boundary\_interval\_on\_the\_Atlantic\_Coastal\_Plain\_with\_a\_description\_of\_the\_highest\_ammonite\_zones\_in\_North\_America\_Part\_2