Echinostoma

## Taxonomy

Anthostoma

Gastrozoa

Tentaclostoma

Neostoma

Lophostoma

Malacoformes

**Echinostoma**

**Echinostoma** (lit. "spiny mouth") is a clade of endoparasitic Malacoformes.

## Description

Echinostomes are a diverse clade of Malacoformes, who live as parasites in the intestines of other terrestrial animals. Echinostomes have diversified in tandem with many of their hosts, having evolved very early on in the history of terrestrial life.

Many Echinostomes are heteroxenous, completing their lifecycle in multiple hosts.

### Biology

Echinostoma anatomy diverges dramatically from that of Eumalacoformes (or any Anthostoma for that matter): their sensory organs, limbs, and most of their internal organs have atrophied completely, while their oviducts have swelled to extend into their spine-covered proboscis. Due to their incredibly low survival rate, Echinostomes are even more R-selective than other Malacoformes, laying several thousands or even millions of eggs.

## Navigation

* **Anthostoma**
	+ Archeanthostoma
	+ Gastrozoa
		- Eugastrozoa
		- Tentaclostoma
			* Lithostraca
				+ Eulithostraca
				+ Anostraca
			* Neostoma
				+ Myzostoma
					- *Pelonomus thomi*
				+ Lophostoma
					- Diplostoma
						* Placostraca
							+ Archaeoplacostraca†
							+ Euplacostraca
								- Euplacostracidae
									* Euplacostracine
									* Echinostracinae†
								- Xerostraca
						* Stenostraca
							+ Eustenostraca
								- Desmostraca
									* Archaeodesmostraca†
										+ Archedesmostracidae†
										+ Notoformes†
											- Archenotoformes†
											- Neonotoformes†
												* Thyreostraca†
												* Campylospondyli†
													+ *Erythrerpeton lequiori*†
									* Eudesmostraca
										+ Eudesmostracidae
											- Eudesmostracinae
											- Myxobolidae
										+ Dolichostraca
											- Pronocanthidae†
												* Eupronocanthidae†
												* Phylactocanthidae†
											- Eudolichostraca
												* Eudolichostracidae
												* Nothocanthidae†
							+ Coleostraca
								- Archaeocoleostraca†
								- Elastospondyli
									* Archelastospondyli
										+ Archelastospondylidae
										+ Kentrodontia
											- Eukentrodontia
												* Archekentrodontia
													+ Archekentrodontidae
													+ Neokentrodontia
														- Trypophora
														- Deinoglossidae
												* Phyllophora†
											- Aspidontia†
												* *Kremarapax mckinleyi*†
									* Opisthoptera
										+ *Eopteryx emmetti*
										+ Archeopisthoptera
											- Latoptera
												* Eulatoptera
												* Pipiodontia
													+ *Corvimimus corvusamousi*
													+ Archepipiodontia
													+ Pterolophidae
											- Picoptera
												* *Gyriphagus oelespi*
										+ Neopisthoptera†
											- Sphenoptera†
												* *Velocialatus alexani*†
											- Magnoptera†
												* Eumagnoptera†
													+ *Skotalatus rauli*†
												* Dyptopterida†
													+ *Tachynectes jocomoli*†
					- Malacoformes
						* **Echinostoma**
						* Eumalacoformes
							+ *unspecified*
								- Ctenognatha
									* Euctenognatha
									* Necroformes
								- *unspecified*
									* Barognatha
									* Aciculognatha
										+ Euaciculognatha
										+ Chlorodermata
							+ *unspecified*
								- *unspecified*
									* Onychognatha
									* Mastigognatha
								- Isognatha
									* Myzognatha
										+ *Vitapatatoria aspiana*
									* Euisognatha
										+ Archeisognatha
										+ Cyanodermata

* **Polypodia**
	+ Tetrabrachia
		- Tachypodia
		- Acanthopodia
			* Orthorhachia
				+ Tanypterida
				+ Remipterida
					- Arheremipterida
					- Synopteridae
			* Hadrorhachia
				+ Ptilopoda†
					- *Bathytitan vendari*
				+ Euhadrorhachia
					- Diplocirridae
						* Archediplocirridae
						* Astrapophora
					- Teleopterida
						* Dorypoda
							+ *Kamakivenator cosmici*
							+ Archedorypoda
							+ Xiphopoda
						* Temnopoda†
							+ *Teratodectes iasoni*
	+ Sarcopodia
		- Coelopoda
			* Eucoelopoda
			* Trichopoda
				+ *Prionorostrum aegisari*
		- Neopoda
			* Metaxypoda†
				+ *Argyroderma piacentini*
			* Osteopoda
				+ Polyschia
					- Platydontia
						* Archeplatydontia
							+ Archeplatydontidae
								- *unspecified*
									* Archeplatydontinae
										+ *Dendronomus vasquezi*
									* Xenodontia†
										+ Brachiocephalia†
										+ Euxenodontia†
											- Streptotarsi†
											- Archexenodontia†
								- Neoplatydontia
									* Pleuroptera
										+ Archepleuroptera
											- Archepleuropteridae†
												* *Anthonomos qbelocki*†
											- Platyptera
												* *Sphenorhynchus sullivanae*
												* Euplatyptera
												* Dermatopoda
													+ Archedermatopoda
													+ Neodermatopoda
														- Rhabdopoda
															* Colobathristes borchmori
														- Compsopoda
										+ Theroptera†
											- Eutheroptera†
												* *Melanopteryx alexanderi*†
											- Chlamyptera†
												* Hypsirhynchidae†
												* Archechlamyptera†
													+ Temnorhynchidae†
														- *Charcharonyx mackrethi*†
													+ Archechlamypteridae†
									* Neoplatydontidae
										+ Scandopoda
											- *Carpophagus xeni*
											- Archescandopoda
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												* Pholcoscelida
												* Brachyscelida
										+ Harpactopoda†
											- Euharpactopoda†
											- Tanybrachidae†
												* *Cephalocarcinus elvari*†
							+ Copetarsi†
								- *Potamodyptes andrewi*†
						* Edaphotheria
							+ Leptopoda
								- Archeleptopoda
									* Euleptopoda†
										+ Euleptopodidae†
										+ Neoleptopoda†
											- Hoplopoda†
											- Camptopoda†
												* Eucamptopoda†
												* Sphenocerata†
									* Amblypoda
								- Thylacopoda†
									* Archethylacopoda†
										+ *Pycnothrix dicksoni*†
									* Allobrachidae†
										+ Euallobrachidae†
											- Corythobrachidae†
												* Archecorythobrachidae†
												* Camarabrachidae†
													+ *Cyanolophus mcgyveri*†
											- Archeallobrachidae†
												* Archeallobrachinae†
													+ *Melanothrix oneiropolos*†
												* Ceratobrachidae†
													+ *Thyreocephalus cryarchon*†
										+ Teleobrachidae†
											- *Therizinyx elliotti†*
							+ Eurycheridae
								- Archeurycheridae†
									* Archeurycheirinae†
										+ *Prolestes solysti*†
									* *Teratocheirus*†
										+ *Teratocheuirus philipi*†
								- Lystrocheiridae
									* Lystrocheirinae
										+ *Psammonyx deschaini*
									* Trypanocheiridae
										+ Archetrypanocheiridae†
										+ Rhamphodontia
											- Eurhamphodontia
												* Archeramphodontia
													+ Apodomorpha
													+ Archeramphodontidae
														- Archeramphodontinae
														- Coryphodontia
															* Archecoryphodontia
															* Septentrioformes
																+ Phractoformes
																+ Cryptonychidae
																	- Archecryptonychidae
																	- Neocryptonychidae
																		* Theristobrachidae
																		* Glyphonychidae
																			+ Archeglyphonychidae
																			+ Archonychidae
												* Acrocheiridae
													+ Euacrocheiridae
														- Archeacrocheiridae
														- Neoacrocheiridae
															* Heterognatha
																+ Distropheognatha
																	- Archedistropheognatha
																	- Neodistropheognatha
																		* Hadronychidae
																		* Daptodontia
																			+ Dryptognatha
																			+ Flagellifera
																+ Procaecoformes
																	- Oligopoda
																	- Euprocaecoformes
																		* Euprocaecoformidae
																		* Megantroformes
															* Panhylotheria
																+ Hylotheria
																	- Thaumatotheriidae
																	- Euhylotheria
																		* Gyrinotheria
																		* Euhylotheriidae
																			+ Euhylotheriinae
																			+ Pelecodontidae
																+ Thrinaxorhynchidae
																	- Archethrinaxorhynchidae
																	- Dictyorhynchidae
													+ Apatocheiridae†
											- Oryctocheiridae
					- Dromaeopoda
						* Archedromaepoda†
							+ *Rhomaleopus*†
							+ Archedromaeopodidae†
								- *Dromovenator setnovi*†
						* Onychodontia
							+ Euonychodontia†
								- Euonychodontidae†
									* Ensidontia†
										+ Euensidontia†
										+ Prionodontia†
									* Euonychodontinae†
										+ Archeonychodontidae†
										+ Cryptodontia†
											- *Raptonyx junae*†
											- Tachydontia†
											- Eucryptodontia†
								- Hadrodontia†
							+ Allodontia
								- Euallodontia†
									* Archallodontia†
									* Eriotherinae†
								- Thecopoda
									* Euthecopoda†
										+ *Leucothrix caliburi*†
									* Amphidontia
										+ Archeamphidontia†
										+ Xenopsidae
											- *polylophodon thomasi*
											- Archexenopsidae
												* Archexenopsinae
												* Euxenopsidae
													+ Ankistrobrachidae
													+ Neoxenopsidae
														- Teleopsidae
														- Chelaphora
															* Stenochelaphora
															* Pachychelaphora
											- Stenopsidae
												* File:StenopsidaeEvo.pngArchestenopsidae
												* Neostenopsidae
													+ Peloropsidae
														- Archepeloropsidae
														- Anaxotheriidae
													+ Ceratodontia
														- Archeceratodontia
														- Neoceratodontia
															* Aratrodontia
															* Isodontia
																+ Kophodontidae
																+ Elaphrodontia
																	- Archelaphrodontia
																	- Neoelaphrodontia
																		* Spathodontia
																		* Xyrodontia
				+ Synischia
					- Megalobrachidae†
						* Archemegalobrachia†
							+ Archemegalobrachidae†
							+ Prenobrachidae†
						* Titanopoda†
							+ Procerapoda†
								- Euprocerapoda†
									* *Megaphagus matti*†
								- Stylopoda†
									* *Hybotitan bluenoti*†
							+ Baropoda†
								- Eubaropoda†
									* *Nanopus proximensis*†
									* Eubaropodidae†
								- Pachypoda†
					- Deinognatha
						* Eudeinognatha†
							+ Eudeinognathidae†
								- *Euoplos gulielmi*†
							+ Malleognatha†
						* Odontognatha
							+ Trachygnatha†
								- *Telmarchos krisi*†
							+ Dolichognatha
								- Eudolichognatha
									* Archedolichognatha†
									* Hybognatha
								- Pelagotheria†
									* Actatheria†
										+ *Eurypus geberi*†
									* Thalattotheria†
										+ Isopterygia†
										+ Anisopterygia†
											- *Pontodrakon nicholsi*†

* **Chemophyta**
	+ Archaeophyta
	+ Xenophyta
		- Caulophyta
			* Neocaulophyta
				+ Eucaulophyta
					- Eucaulophytaceae
						* Eucaulophytoideae
						* Nothodendrales†
					- Reptophyta
				+ Brachyphyta
					- Archebrachyphyta
						* Eubrachyphyta
							+ Eubrachyphytaceae
							+ Desmophyta
								- Carnophyta†
								- Chromatophyta
									* Archechromatophyta
										+ Archechromatophytacea
										+ Aphyllophyta
									* Zygophyta
										+ Zygophytacea
											- Zygodendrales
											- Archezygophytacea
										+ Harpactophyta†
						* Tylophyta
					- Cryptophyta
						* Eucryptophyta
						* Pagophyta
			* Necrophyta
		- Xylophyta
			* Euxylophyta
				+ Archexylophyta
					- Archexylophytacaea
					- Coelophyta
				+ Nodophyta
			* Altiphyta†
				+ *Trilobophyllus axidenti*†

 † = Extinct