Gomphiobia

For this group of dispeculates the basal condition is as an aquatic predator with 8 lateral fins, though plenty of species had these modified into limbs for life on land.

Gomphiobians are the basalmost dispeculates, yet one of the most successful and diverse. Gomphiobians are older than celernantids, ambulospeculids or panoplovitans, having radiated with little control and dominating the underwater environment and radiating there for various millions of years, until subsequent extinction events allowed the proliferation of marine celernantids and panoplovitans, but nevertheless allowing the gomphiobians to share the domain below the waves. Freshwater and marine gomphiobians exist in abundance. As for adaptations onto land, gomphiobians must be the kings. Although many land-dwelling groups still retain various primtiive adaptations for land-dwelling support, its all the most jaw-dropping to know the sheer amount of lineages of gomphiobians that evolved terrestrial habits independently, and often in such a little period of time. Early on, gomphiobians became one of the most relevant fauna of Wallace II's dry land, forming the most formidable predators and even the largest land-dwelling dispeculates alive today. Flying gomphiobians exist in the form of a few independently evolved clades, being the most important part of one major clade of flying gomphiobians evolved from aquatic creatures. These have proliferated thanks to their adaptability to the water environments or colder environments, avoiding competition with many groups of flying ambulospeculids, though competition between these has been recorded.

## Classification

* Astapiscatoria
	+ Panzerdracoidea
		- **Panzercaria** (Panzercarians are an ancestral group of aquatic gomphiobians, that has diverged from its closest living relatives even before the Tenerian. They are only represented by Panzercaris. That lives in the tropical rivers of Media Insula, Micha, Auth and Terra Septentrionis, though occasionally visiting regions of marine waters, which allowed them to spread across these distant continents. Panzercarians are oviparous.)
		- Xenogalea
			* **Astagaleoidea** (Astagaleoideans are very derived xenogalean gomphiobians, mostly represented for having weird appendages poking out. The clade diverged from latapiscians before the Tenerian, but they themselves evolved in the Scelusian, and since then they have diverged and evolved in radically different forms, but usually not commiting a particularly adamant impact in the Wallacian ecosystem. They are usually marine, and when so, they prefer the colder and temperate waters to avoid competition with other groups, like the astagaleadae living in the northern septentrionian waters, and the remobrof in ths southern virtuan oceans. The jiruth is a highly derived one, that has returned to land, where they climb trees in warmer forests and swamps of Occidentis, waiting for prey. Astagaleoideans are oviparous.)
			* **Latapiscia** (Latapiscians are a very ancient group of aquatic gomphiobians, in fact the oldest "major" clade of gomphiobians, dated back before the Tenerian. They are found in every ocean, although with a striking absence in most of the temperate and tropical Mare Orientalem. A good amount seemingly likes to find an exclusive refuge in the antarctic seas, surrounding Virtu, like Cimex piscis and Galeapistris, which only exist there. Maxillacollum virescens, Aquatilia insidiator and Aquatilia scolopendra are endemic to the Isla Inferno. Profugnathus rojas lives in Medio Mari and Tenebrosa Aqua. The related yellow-winged butterfly fish lives in the southern Medio Mari, northern Tenebrosa Aqua and Vadum Mare. Pseudomandibulatoidea is found only in Vadum Mare. The cephaplodraco is unique for evolving terrestrial habits, a characteristic that gomphiobians seem to have evolved independently countless times throughout their evolution, and this one lives in the secluded lands of Virtu. Latapiscians are oviparous.)
	+ Leorhynchoata
		- **Sahleenodontia** (Sahleenodontians are very basal to the big land-dwelling lineage of astapiscatorians. Sahleenodontians have diverged from the debellatans in the Tenerian. Sahleenodontians are only represented by Sahleenodon piscator, a large semi-aquatic predator, that reaches 20+ meters in length, in many ways, simillar to a *Spinosaurus*. It is only found in Terra Septentrionis, a refuge from elsewhere, where their relatives went extinct. Sahleenodontians are oviparous.)
		- Debellalata
			* **Cetaflumenia** (Cetaflumenians have originated in the Accomodatian and diverged from the neoskiensians back in the Tenerian. These are a largely aquatic clade, but they are descended from an ancient terrestrial lineage that returned to the water. Cetaflumenians are only found in waters surrounding Media Insula and southern Terra Septentrionis, Plagam Maris and the waters surrounding Virtu. Cetaroid thomasi lives in the waters around Media Insula. Caelusaccelestivata is a freshwater clade found in the swamps and marshes of Virtu, that is able to glide out of the water. Debellator flumen is found around Plagam Maris. Vortodorsus is a cetaflumenian that has returned to being able to walk on land again, but is largely aquatic still, living in the coastlines around Vadum Mare, including Media Insula and Micha, as well as in southern Terra Septentrionis. Cetaflumenians are oviparous.)
			* Neoskiensia
				+ **Notonychia** (Notonychians are a group of basal terrestrial gomphiobian predators. They have diverged from soliursans in the Tenerian but have originated in the Scelusian. Leonix is the basalmost and, with the isolation, it has evolved into a formidable apex predator in Terra Septentrionis. The other remaining notonychians, like the felidire and the southern rasadon, are native to Media Insula, with the latter being found in the southern Gabre archipelago with more abundance. Notonychians are basally oviparous, but Leonix and the felifire have independently evolved viviparity.)
				+ Soliursa
					- **Amplectenda** (Amplectendans are a very diverse group of terrestrial gomphiobians that have diverged very early. They have diverged from doryplocephalians in the Tenerian, and themselves also evolved in the late Tenerian. They eventually expanded to most continents, only being absent in Occidentis, after the Compsogene-Recensgene extinction event took a toll on terrestrial astapiscatorians as a whole in the continent. In Auth, there is the bone tree feeding Amplectendo sphenodyptis and the pig-sized carnivorous Discutio bufo. Due to their early divergence, some species are naturally found in Terra Septentrionis, as they have diverged before the continent became isolated, and in that continent, the Ferrumdentia cursora and the fuzzbuckets are native. The sextripodeslacerta reached there through swimming, as many species are semi-aquatic and even fully aquatic, but some seem to have turned into terrestrial creatures once they reached Terra Septentrionis. In Media Insula, only bargruda is native, but seemingly fuzzbuckets have reached the continent through rafting from Terra Septentrionis to Thron in the Compsogene. The Media Insulan Chymera gracilis seems to be part of a Terra Septentrionian lineage that migrated to Media Insula the same way, but eventually that lineage went extinct in the northern continent. In Virtu, the cave bear and the Trirhynchoatus reside. Amplectendans are oviparous.)
					- Doryplocephalia
						* **Solinantia** (Solinantians are weirdly the only terrestrial leorhynchoatans that exist in Occidentis, seemingly being the only ones that survived the aftermath of the Compsogene-Recensgene extinction event. They are also completely exclusive to Occidentis. They likely survived due to their burrowing habits. Solinantians have diverged from the neocephalians in the Tenerian, but themselves have evolved in the Scelusian. Solinantians are viviparous.)
						* Neocephalia
							+ **Carcharasectia** (Carcharasectians are ancient semi-aquatic gomphiobians that are only represented by Carcharasect testudines. Its semi-aquatic habits and evolution are often suggestive that most of the modern leorhynchoatans have developed full terrestriality independently various times, with their common ancestor being just semi-terrestrial, though this is not witgout debate. Carcharasect is unique to the Archa archipelago where they have been evolving in isolation ever since. The carcharasectians have diverged from the neognathoideans since the Tenerian. Carcharasectians are oviparous.)
							+ Neognathoidea
								- **Hastumnasia** (Hastumnasians are only represented by Hastumnasus latus. These are a weird slow moving omnivorous creature that have diverged from acugnathians in the Tenerian. They are only found in Media Insula. Hastumnasians are oviparous.)
								- Acugnathia
									* **Neognathia** (Neognathians are a primitive group of carnivorous gomphiobians, only represented by the Neognathus. They have diverged from the ixodadivolsellioideans in the late Tenerian. They are only found in Terra Septentrionis, where they have evolved in isolation. Neognathians are oviparous.)
									* Ixodadivolsellioidea
										+ **Speculobestia** (Speculobestians have diverged from doryplosaurs just after the Tenerian-Accomodatian extinction event, showing how little terrestrial gomphiobians were affected by that particular mass extinction. Speculobestians themselves have evolved in the late Scelusian, and are currently found in Virtu, Auth and Media Insula. The glass wingdrifter is a small gliding creature found in the forests of Virtu. The acucephal is a social cursorial predator abundant in the open plains of Auth. Ixodidasicut bestia is a small parasitic speculobestian that parasitises the flying celernantid Arenahastama in Media Insula. Speculobestians are oviparous.)
										+ **Doryplosauria** (Doryplosaurians are a very diverse group of terrestrial gomphiobians. A good portion of these are carnivorous. They have evolved in the Accomodatian, in fact most of the modern species have diverged from each other in that period, making most of the species separate from each other for over 200 million years. They are currently found in every continent except Occidentis and Virtu. Media Insula is notorious for its abundance of apex predatory gomphiobians, and doryplosaurs do not let themselves be intimidated by that, though not just predators exist in this clade. In Media Insula, Doryplokamus, the sailback pincer, Xenocarnifex importunus and Gomphiomorph skiensis are native to the continent, with tridoprobozaurians and Tridentigerivenator rex migrating from Terra Septentrionis to Media Insula, through Thron. Terra Septentrionis is, therefore, only present with tridoprobozaurians and the Tridentigerivenator, which have evolved there in isolation and are therefore native there. In Auth, the related Aliepardus loccisoria and Scuamea convolvens lupus are the only natives, but Tridentigerivenator managed to reach Auth, maybe by floating the ocean from Media Insula to Auth, as their exoskeleton allows some buoyancy, and that the colonization might have been accidental in that case. Tridentigerivenator's persistance in conquering landmasses through buoyant spreading as allowed it to spread to the Micha archipelago and some other eastern islands of Media Insula. Doryplosaurus are oviparous.)
* Electrovenatoria
	+ Cochleavenatoria
		- **Candentisia** (Candentisians are a simplistic and basal aquatic clade of gomphiobians that have originated in the Tenerian. They have diverged from insidiocaroids in the early Tenerian too. They are found in every ocean of the globe, specially the cosmopolitan Candentis serpenti. Algamimus flosseti is found both in freshwater and sea water, living around the coasts of Media Insula and surrounding islands, southern Micha, northern and eastern Auth, Prin, southern Occidentis and Phael. Cochleacetaceus northreni is a giant marine creature found in Vadum Mare, the seas that bathe south and northeast of Media Insula, as well as the isolated Perierat Montibus sea. Candentisians are largely oviparous, but Cochleacetus has evolved ovoviviparous habits.)
		- Insidiocaroidea
			* **Insidiocara** (Insidiocarans are the basalmost terrestrial clade of cochleavenatorians, being comparable to some stem-tetrapods, at least in their infancy. They have diverged from turpisloricatans in the Tenerian, but the extant species diverge from each other in the Variagene. Insidiovenator is the most primitive looking, being found in the swamps of central Media Insula. Senzicara has, in turn, been isolated in the Prin island, where they eventually evolved crescent terrestrial habits and became giant and formidable apex predators there. Insidiocarans are oviparous.)
			* Turpisloricata
				+ **Turpiscarnifexia** (Turpiscarnifexians have diverged and evolved back in the Tenerian. They are only represented by the modern Turpiscarnifex mesomelas. This is a primitive carnivore and represents what early true terrestrial cochleavenatorians would have looked like. This creature is only found in the deserts of Media Insula. Turpiscarnifexians are ovoviviparous.)
				+ Ensicrita
					- **Tegroimpeta** (Tegroimpetans have diverged from entomogracilians in the late Tenerian and evolved in the Accomodatian. Tegroimpetus magna is the only one found outside of Occidentis, in Auth. Craniumimmus and Osloricati are found in Occidentis, an example of how ensicritans have surpassed the separately evolved terrestrial leorhynchoatans in their diversity in the Occidentis continent, after the beginning of the Recensgene. Tegroimpetans are ovoviviparous.)
					- **Entomogracilia** (Entomogracilians have first evolved in the Accomodatian. They have diversified in various ways, mostly to avoid competition with other land-dwelling creatures that were diversifying themselves across the history of the planet. Non-flying or marine entomogracilians are found just in Occidentis and Media Insula, being Occidentis the ones they have their most accentuated diversity, maybe due to the absence of relevant leorhynchoatan fauna. In Occidentis, Herocritus and Ensirostrum platygracilis exist, as well as the largest terrestrial dispeculate alive today, the Lupus triplous, a 45 meter long predator that reached these titanic sizes, never seen before in terrestrial dispeculate history, very recently, in the Recensgene, likely as a result of being able to predate even bigger land-dwelling sidobosians, as full adults. In Media Insula, only one, closely related clade, of predators exists, including the layer-jaw, the Pantheramimus ignaticus and the Quadente magna. Entomochelona is the only Terra Septentrionian entomogracilian, having reached there through its aquatic habits, as it can be found in the sea and in freshwater, also being able to crawl onto land. The viditdon sanguinem-lactam is a small flying relative of the Lupus triplous, that has evolved recently in the Recensgene. Its recent evolution has been accompanied by a sudden success across the entire globe, being found in every continent and even isolated islands, in some places even outcompeting small flying dispeculates, like xenopedans. Entomogracilians are oviparous.)
	+ Praedocreatura
		- Praedosmilia
			* **Ornithosmilia** (Ornithosmilians are the basalmost major flying clade of the praedosmilians, also known as the main flying clade of gomphiobians. Ornithosmilids have diverged from exireginans in the Tenerian, indicating that some form of volant, or at least gliding form, existed back then, allowing the different clades to eventually evolve flight independently. They seem to have evolved from aquatic species, though, at first. Currently, ornithosmilids are a very diverse clade filled with different creatures with varying shapes and sizes, spread across most continents and archipelagos, only absent in most of Occidentis, except the northwestern tip, and the entirety of Virtu. Ornithosmilians are oviparous.)
			* Exiregina
				+ **Ichthyowestonia** (Ichthyowestonians are an ancient flying clade that has diverged from praedoreginans in the Accomodatian, and evolved in the Scelusian. They are very derived in their flying techniques, but there seems to be an indication of an aquatic origin for the clade, or at least one in which many members have actively went back to water independently or became dependant of this. Many are still actively terrestrial. This clade is among the most influential group of flying dispeculates on Wallace II, being cosmopolitan and found everywhere in the globe, specially the dracolodontidians, which are accipitriformes-like flying gomphiobians that take predatory-like niches in its majority and are found in every continent and island, as well as the pterolocaridids, which are aquatic flying hunters. Avemaris westonium is an aquatic and aerial basal ichthyowestonian, found in Vadum Mare and the waters that sorround Terra Septentrionis. Ichthypteryx prosilus also flies actively around the waters of Terra Septentrionis, presumably to avoid competition from caedifuremians or Volanmaris. The gomphopteryx is found in Occidentis, while the Papiliocornus magnum, again, refuges itself in Terra Septentrionis. Ichthyowestonians are largely oviparous, with the exception of the fully aquatic Avemaris, which is viviparous.)
				+ **Praedoregina** (Praedoreginans are the close relatives of ichthyowestonians, having first evolved in the Scelusian. They are usually distinguished from their more aquatic-oriented relatives, to being more terrestrially-oriented. Many lineages have become more terrestrial than ever, including one main lineage that has crescently and ultimately became fully flightless by the Compsogene. They are found in every continent except Auth and Terra Septentrionis. The spoiler queen is found in Virtu, as well as the arboreal flightless Simialiena dulcemamma. In Media Insula, the flying Exipteryx exists, as well as the, still flying, yet more terrestrially oriented creature that is informally called the "Dino drawings creature", and the Constantinus theria rostratis victoria, which is fully flightless and is found in Media Insula and Micha. The fully flightless Terracetus gigas is found in Occidentis. Praedoreginans are oviparous.)
		- Formicichthya
			* **Electrapistrida** (Electrapistridans are the basalmost clade of formicichthyans, having diverged from telumorians in the Tenerian and, themselves, having evolved in the Accomodatian. They are very diverse and are found in almost every ocean of the globe, specially the pterapistrids, which are found everywhere that is sea, except the colder northern waters of the planet. Around Media Insula, the largest diversity exists. Exclusively freshwater Media Insulan species include the electric daggerfish and the Ambiguustoma hydroensis. Insidiis dente is a gigantic freshwater species that exists in the cold river deltas of Virtu. Formicirae pinnisphen and Cimetroiani are closely related and are known for some species of these spending sometime out of the water. In the case of Cimetroiani a whole variety of fully aquatic, marine, freshwater, semi-aquatic and even fully terrestrial species have evolved in Media Insula, Micha and Terra Septentrionis, varying greatly in sizes, no matter what specific habitat that statement is referring to. Formicirae is a marine creature that can crawl onto land in occasion being found around Media Insula. Electrapistridans are largely oviparous, but pterapistridans, such as Ambiguustoma and pterapistrids, are viviparous.)
			* Telumoria
				+ **Xenopleurodontia** (Xenopleurodonts are an extremelly ancient group, having existed since the early Tenerian. They are found, with a patchy distribution, in marine waters across the globe, bathing every continent, with the possible exception of Auth. Many seem to have a preference in cold waters. In Virtu, the primitive Celeria denshastia and the more derived Captiormus fermarture exist. Around Terra Septentrionis, the Meditellis fermarture and the Stiriminamyi fermarture live. In more tropical waters, more flashy hunters exist. Around Media Insula, Telumnasus dux exists, while the basal, yet jawes, Xenopleurodon speculatas lives in Vadum Mare and in the marine waters west of Occidentis. Xenopleurodonts are oviparous.)
				+ Velocipiscatoria
					- **Thoramphibia** (Thoramphibians have evolved in the Tenerian, and they have been known to have had a long history of diversity and collapse over millions of years, but currently, only three accepted species are known, onky found around Terra Septentrionis and Media Insula. The wallacean lamprey and the Velocitas gladiodente are fully marine creatures living around Media Insula, while Tesca amphibiarum is a freshwater and semi-aquatic creature that lives in Terra Septentrionis. Thoramphibians are basally oviparous, but the allothorakismenikefaloids, such as the wallacean lamprey and Velocitas are viviparous.)
					- Titanotrigonia
						* **Giraffapiscatoria** (Giraffapiscatorians are a clade of semi-terrestrial, and often even fully terrestrial of gomphiobians, one of the many clades that has evolved terrestriality independently. They diverged from ostrigonians in the Tenerian and evolved in the Accomodatian from aquatic species. Most are still largely aquatic, however. They are only found around Media Insula and Auth. Giraffapiscator is the basalmost, but most iconic as well, found in coastal waters of Media Insula and Micha. In Media Insula, the closely related Plumathong osslug and Intertesta exits, respectively, highly marine and terrestrial species. In Auth, the Procerus titanus exists, as well as the semi-aquatic and predatory Ursasuchus. Giraffapiscatorians are largely oviparous, with the exception of Ursasuchus and Plumathong that have evolved viviparity independently.)
						* **Ostrigonia** (The ostrigonians have first evolved in the Tenerian. Being found in most oceans of the globe, with central Medio Mari being the largely unusual exception. There are many fully marine species, such as the Cornuscopu that surrounds Occidentis, and the related Fortitorsus trigonus that lives in seas surrounding Media Insula. Os oris lives in waters surrounding Occidentis. The related Testa leo and Titanoangulus, the latter a titanic filter feeder, are exclusive to the tropical waters surrounding Media Insula. The Osteosuchus is a large carnivorous aquatic creature that lives in southeastern Vadum Mare but ventures notoriously to the river deltas of Media Insula, into Musa Sapientum Fixa, where it is the aquatic apex predator, so influential that some freshwater celernantids even mimic them for their own benefit. The Diabolus is likely one of the most scary. They are found in Tenebrosa Aqua and Mare Orientalem, and they are gigantic, specially the males, with three times the size of the females. They are even able to inflate and expand their flippers to glide below the water, however young are able to glide very high in the air, once the wind conditions are favorable. What makes these creatures so jaw dropping, is that the largest species can have a wingspan of 50 meters, and a length of 30 meters, though they do not glide at this size. Ostrigonians are oviparous, with the exception of titanoangulatans, such as testa leo and Titanoangulus, which are ovoviviparous.)

## List of Gomphiobians

### Aquatic members

* *Algamimus flossetis* - marsh spear
* *Celeridente hooverii* - Razor Jaws
* *Cochleacetus northreni*
* *Diabolus*
* Iaculidae
* Panzercaris
* *Pseudohypostomus silurimorpha* - Panzercaris
* Phytovorus Mimus
* Pterapistridae
* Scissor Beak (Aquatic when born, Aerial as adult)
* Skint
* *Thorakismenikefali psaria* - Spear Whale
* *Xenopleurodon*
* Yellow-Winged Butterfly Crestfin

### Terrestrial members

* Aquaovium schnorchelii - Water Sheep
* Arborecursor
* Bouncers
* Caelipiscator
* *Daemonium* - Forest Demon
* *Diablotaurus vespa* - Devil bull
* *Exhibuita vulgaris* – "Common Climber"
* Falcatisuchus
* *Saltator* - Bouncers
* *Smilorhinus thisskus* - Walking Blade
* Snowy Shadow
* Uniparrion
* Vrontientomos hespaestus

### **Canon**

Panzercaria (309-0)

* Panzercaris

Astagaleoidea (205-0)

* Astagaleadae
* Jiruth
* Remobrof

Latapiscia (305-0)

* Aquatilia insidiator
* Aquatilia scolopendra
* Cephaplodraco
* Cimex piscis
* Galeapistris
* Maxillacollum virescens
* Profugnathus rojas
* Pseudomandibulatoidea
* Wide-gilled gorgon
* Yellow-winged butterfly crestfin

Sahleenodontia (290-0)

* Sahleenodon piscator

Cetaflumenia (213-0)

* Caelusaccelestivata
* Cetaroid thomasi
* Debellator flumen
* Vortodorsus

Notonychia (186-0)

* Felidire
* Leonix
* The southern rasadon

Amplectenda (257-0)

* Amplectendo sphenodyptis
* Bargruda
* Cave bear
* Chymera gracilis
* Discutio bufo
* Ferrumdentia cursora
* Fuzzbuckets
* Sextripodeslacerta
* Trirhynchoatus

Solinantia (178-0)

* Nix anguis
* Soil swimmer
* Witch's parasol

Carcharasectia (260-0)

* Carcharasect testudines

Hastumnasia (254-0)

* Hastumnasus latus

Neognathia (248-0)

* Neognathus

Speculobestia (162-0)

* Acucephal
* Glass wingdrifter
* Ixodidasicut bestia

Doryplosauria (236-0)

* Aliepardus loccisoria
* Doryplokamus
* Gomphiomorph skiensis
* Sailback pincer
* Scuamea convolvens lupus
* Tridentigerivenator rex
* Tridoprobozauria
* Xenocarnifex inportunus

Candentisia (287-0)

* Algamimus flossetis
* Candentis serpenti
* Cochleacetaceus northreni

Insidiocara (145-0)

* Insidiovenator
* Senzicara

Turpiscarnifexia (285-0)

* Turpiscarnifex mesomelas

Tegroimpeta (234-0)

* Craniummimus
* Osloricati
* Tegroimpetus magna

Entomogracilia (215-0)

* Ensirostrum platygracilis
* Entomochelona
* Herocritus
* Layer-jaw
* Lupus triplous
* Pantheramimus ignaticus
* Quadente magna
* Viditdon sanguinem-lactantem

Ornithosmilia (261-0)

* Ornithosmilidae

Ichthyowestonia (180-0)

* Avemaris westonium
* Dracolodontidia
* Ichthyopteryx prosilitus
* Papiliocornus magnum
* Pterolocarididae
* The gomphogopteryx

Praedoregina (187-0)

* Constantinus theria rostratis victoria
* Dino drawings creature
* Exipteryx
* Simialiena dulcemamma
* Spoiler queen
* Terracetus gigas

Electrapistrida (223-0)

* Ambiguustoma hydroensis
* Cimetroiani
* Electric daggerfish
* Formicirae pinnisphen
* Insidiis dente
* Pterapistridae

Xenopleurodontia (292-0)

* Captiormus fermarture
* Celeria denshastia
* Gladiogomphius
* Meditellis fermarture
* Stiriminamyi fermarture
* Telumnasus dux
* Xenopleurodon speculatas

Thoramphibia (262-0)

* Tesca-amphibiarum
* Thorakismenikefali psaria (now regarded as an ambulospeculid)
* Velocitas gladiodente
* Wallacean lamprey

Giraffapiscatoria (218-0)

* Giraffapiscator
* Intertesta
* Plumathong osslug
* Procerus titanus
* Ursasuchus

Ostrigonia (261-0)

* Cornuscopu
* Diabolus tenebrarum
* Fortitorsus trigonus
* Os oris
* Osteosuchus
* Testa leo
* Titanoangulus