Tero the Triceratops

Tero the Triceratops

## Tero

It is a Yellow Triceratops with Brown and White and Red Stripes It is a 20 years old Male Tero Has a Girlfriend Named Mallow Who Live Together like a Human Being Tero Horn in his Head 3 Horn today news of Fighting

## Information

***Triceratops 'three-horned face') is a genus of chasmosaurine ceratopsian dinosaur that lived during the late Maastrichtian age of the Late Cretaceous period, about 68 to 66 million years ago in what is now western North America. It was one of the last-known non-avian dinosaurs and lived until the Cretaceous–Paleogene extinction event 66 million years ago. The name Triceratops, which literally means 'three-horned face', is derived from the Greek words trí- (τρί-) meaning 'three', kéras (κέρας) meaning 'horn', and ṓps (ὤψ) meaning 'face'.***

***Bearing a large bony frill, three horns on the skull, and a large, four-legged body, exhibiting convergent evolution with bovines and rhinoceroses, Triceratops is one of the most recognizable of all dinosaurs and the best-known ceratopsian. It was also one of the largest, up to 8–9 metres (26–30 ft) long and 5–9 metric tons (5.5–9.9 short tons) in body mass. It shared the landscape with and was most likely preyed upon by Tyrannosaurus, though it is less certain that two adults would battle in the fanciful manner often depicted in museum displays and popular media. The functions of the frills and three distinctive facial horns on its head have inspired countless debates. Traditionally, these have been viewed as defensive weapons against predators. More recent interpretations find it probable that these features were primarily used in species identification, courtship, and dominance display, much like the antlers and horns of modern ungulates.***

***Triceratops was traditionally placed within the "short-frilled" ceratopsids, but modern cladistic studies show it to be a member of Chasmosaurinae, which usually have long frills. Two species, T. horridus and T. prorsus, are considered valid today. Seventeen different species, however, have been named throughout history. Research published in 2010 concluded that the contemporaneous Torosaurus, a ceratopsid long regarded as a separate genus, represents Triceratops in its mature form. This view has still been highly disputed and much more data is needed to settle this ongoing debate.***

***Triceratops has been documented by numerous remains collected since the genus was first described in 1889 by American paleontologist Othniel Charles Marsh. Specimens representing life stages from hatchling to adult have been found. As the archetypal ceratopsian, Triceratops is one of the most beloved, popular dinosaurs and has been featured in numerous films, postage stamps, and many other types of media.***

## ***Discovery and identification[edit]***

***See also: Timeline of ceratopsian research***

***The first named fossil specimen now attributed to Triceratops is a pair of brow horns attached to a skull roof that were found by George Lyman Cannon near Denver, Colorado, in the spring of 1887. This specimen was sent to Othniel Charles Marsh, who believed that the formation from which it came from dated from the Pliocene and that the bones belonged to a particularly large and unusual bison, which he named Bison alticornis. He realized that there were horned dinosaurs by the next year, which saw his publication of the genus Ceratops from fragmentary remains, but he still believed B. alticornis to be a Pliocene mammal. It took a third and much more complete skull to fully change his mind.***

***Although not confidentially assignable, fossils possibly belonging to Triceratops were described as two taxa, Agathaumas sylvestris and Polyonax mortuarius, in 1872 and 1874, respectively, by Marsh's archrival Edward Drinker Cope. Agathaumas was named based on a pelvis, several vertebrae, and a few ribs collected by Fielding Bradford Meek and Henry Martyn Bannister near the Green River of southeastern Wyoming from layers coming from the Maastrichtian Lance Formation. Due to the fragmentary nature of the remains, it can only confidently be assigned to Ceratopsidae. Polyonax mortuarius was collected by Cope himself in 1873 from northeastern Colorado, possibly coming from the Maastrichtian Denver Formation. The fossils only consisted of fragmentary horn cores, 3 dorsal vertebrae, and fragmentary limb elements. Polyonax has the same issue as Agathaumas, with the fragmentary remains non-assignable beyond Ceratopsidae.***

***The Triceratops holotype, YPM 1820, was collected in 1888 from the Lance Formation of Wyoming by fossil hunter John Bell Hatcher, but Marsh initially described this specimen as another species of Ceratops. Cowboy Edmund B. Wilson had been startled by the sight of a monstrous skull poking out of the side of a ravine. He tried to recover it by throwing a lasso around one of the horns. When it broke off, the skull tumbling to the bottom of the cleft, Wilson brought the horn to his boss. His boss was rancher and avid fossil collector Charles Arthur Guernsey, who just so happened to show it to Hatcher. Marsh subsequently ordered Hatcher to locate and salvage the skull. The holotype was first named Ceratops horridus. When further preparation uncovered the third nose horn, Marsh changed his mind and gave the piece the new generic name Triceratops (lit. 'three horn face'), accepting his Bison alticornis as another species of Ceratops. It would, however, later be added to Triceratops. The sturdy nature of the animal's skull has ensured that many examples have been preserved as fossils, allowing variations between species and individuals to be studied. Triceratops remains have subsequently been found in Montana and South Dakota (and more in Colorado and Wyoming), as well as the Canadian provinces of Saskatchewan and Alberta.***

### ***Species[edit]***

***After Triceratops was described, between 1889 and 1891, Hatcher collected another thirty-one of its skulls with great effort. The first species had been named T. horridus by Marsh. Its specific name was derived from the Latin word horridus meaning "rough" or "rugose", perhaps referring to the type specimen's rough texture, later identified as an aged individual. The additional skulls varied to a lesser or greater degree from the original holotype. This variation is unsurprising, given that Triceratops skulls are large three-dimensional objects from individuals of different ages and both sexes that which were subjected to different amounts and directions of pressure during fossilization.***

***In the first attempt to understand the many species, Richard Swann Lull found two groups, although he did not say how he distinguished them. One group composed of T. horridus, T. prorsus, and T. brevicornus ('the short-horned'). The other composed of T. elatus and T. calicornis. Two species (T. serratus and T. flabellatus) stood apart from these groups. By 1933, alongside his revision of the landmark 1907 Hatcher–Marsh–Lull monograph of all known ceratopsians, he retained his two groups and two unaffiliated species, with a third lineage of T. obtusus and T. hatcheri ('Hatcher's') that was characterized by a very small nasal horn. T. horridus–T. prorsus–T. brevicornus was now thought to be the most conservative lineage, with an increase in skull size and a decrease in nasal horn size. T. elatus–T. calicornis was defined by having large brow horns and small nasal horns. Charles Mortram Sternberg made one modification by adding T. eurycephalus ('the wide-headed') and suggesting that it linked the second and third lineages closer together than they were to the T. horridus lineage.***

***With time, the idea that the differing skulls might be representative of individual variation within one (or two) species gained popularity. In 1986, John Ostrom and Peter Wellnhofer published a paper in which they proposed that there was only one species, Triceratops horridus. Part of their rationale was that there are generally only one or two species of any large animal in a region. To their findings, Thomas Lehman added the old Lull–Sternberg lineages combined with maturity and sexual dimorphism, suggesting that the T. horridus–T. prorsus–T. brevicornus lineage was composed of females, the T. calicornis–T. elatus lineage was made up of males, and the T. obtusus–T. hatcheri lineage was of pathologic old males.***

***These findings were contested a few years later by paleontologist Catherine Forster, who reanalyzed Triceratops material more comprehensively and concluded that the remains fell into two species, T. horridus and T. prorsus, although the distinctive skull of T. ("Nedoceratops") hatcheri differed enough to warrant a separate genus. She found that T. horridus and several other species belonged together and that T. prorsus and T. brevicornus stood alone. Since there were many more specimens in the first group, she suggested that this meant the two groups were two species. It is still possible to interpret the differences as representing a single species with sexual dimorphism.***

***In 2009, John Scannella and Denver Fowler supported the separation of T. prorsus and T. horridus, noting that the two species are also separated stratigraphically within the Hell Creek Formation, indicating that they did not live together at the same time.***

#### ***Valid species[edit]***

* ***T. horridus (Marsh, 1889) Marsh, 1889 (originally Ceratops) (type species)***
* ***T. prorsus Marsh, 1890***

#### ***Synonyms and doubtful species[edit]***

***Some of the following species are synonyms, as indicated in parentheses ("=T. horridus" or "=T. prorsus"). All the others are each considered a nomen dubium (lit. 'dubious name') because they are based on remains too poor or incomplete to be distinguished from pre-existing Triceratops species.***

* ***T. albertensis C. M. Sternberg, 1949***
* ***T. alticornis (Marsh 1887) Hatcher, Marsh, and Lull, 1907 [originally Bison alticornis, Marsh 1887, and Ceratops alticornis, Marsh 1888]***
* ***T. brevicornus Hatcher, 1905 (=T. prorsus)***
* ***T. calicornis Marsh, 1898 (=T. horridus)***
* ***T. elatus Marsh, 1891 (=T. horridus)***
* ***T. eurycephalus Schlaikjer, 1935***
* ***T. flabellatus Marsh, 1889 (= Sterrholophus Marsh, 1891) (=T. horridus)***
* ***T. galeus Marsh, 1889***
* ***T. hatcheri (Hatcher & Lull 1905) Lull, 1933 (contentious; see Nedoceratops below)***
* ***T. ingens Marsh vide Lull, 1915***
* ***T. maximus Brown, 1933***
* ***T. mortuarius (Cope, 1874) Kuhn, 1936 (nomen dubium; originally Polyonax mortuarius)***
* ***T. obtusus Marsh, 1898 (=T. horridus)***
* ***T. serratus Marsh, 1890 (=T. horridus)***
* ***T. sulcatus Marsh, 1890***
* ***T. sylvestris (Cope, 1872) Kuhn, 1936 (nomen dubium; originally Agathaumas sylvestris)***

## ***Description[edit]***

### ***Size[edit]***

***Triceratops was a very large animal, reaching 8–9 metres (26–30 ft) in length and weighing 6–12 metric tons (6.6–13.2 short tons). A specimen of T. horridus named Kelsey measured 6.7–7.3 meters (22–24 ft) long, has a 2-meter (6.5 ft) skull, stood about 2.3 meters (7.5 ft) tall, and was estimated by the Black Hills Institute to weigh approximately 5.4 metric tons (6.0 short tons).***

### ***Skull[edit]***

***Like all chasmosaurines, Triceratops had a large skull relative to its body size, among the largest of all land animals. The largest-known skull, specimen MWC 7584 (formerly BYU 12183), is estimated to have been 2.5 meters (8.2 ft) in length when complete and could reach almost a third of the length of the entire animal.***

***The front of the head was equipped with a large beak in front of its teeth. The core of the top beak was formed by a special rostral bone. Behind it, the premaxillae bones were located, embayed from behind by very large, circular nostrils. In chasmosaurines, the premaxillae met on their midline in a complex bone plate, the rear edge of which was reinforced by the "narial strut". From the base of this strut, a triangular process jutted out into the nostril. Triceratops differs from most relatives in that this process was hollowed out on the outer side. Behind the toothless premaxilla, the maxilla bore thirty-six to forty tooth positions, in which three to five teeth per position were vertically stacked. The teeth were closely appressed, forming a "dental battery" curving to the inside. The skull bore a single horn on the snout above the nostrils. In Triceratops, the nose horn is sometimes recognisable as a separate ossification, the epinasal.***
***The skull also featured a pair of supraorbital "brow" horns approximately 1 meter (3.3 ft) long, with one above each eye. The jugal bones pointed downward at the rear sides of the skull and were capped by separate epijugals. With Triceratops, these were not particularly large and sometimes touched the quadratojugals. The bones of the skull roof were fused and by a folding of the frontal bones, a "double" skull roof was created. In Triceratops, some specimens show a fontanelle, an opening in the upper roof layer. The cavity between the layers invaded the bone cores of the brow horns.***

***At the rear of the skull, the outer squamosal bones and the inner parietal bones grew into a relatively short, bony frill, adorned with epoccipitals in young specimens. These were low triangular processes on the frill edge, representing separate skin ossifications or osteoderms. Typically, with Triceratops specimens, there are two epoccipitals present on each parietal bone, with an additional central process on their border. Each squamosal bone had five processes. Most other ceratopsids had large parietal fenestrae, openings in their frills, but those of Triceratops were noticeably solid, unless the genus Torosaurus represents mature Triceratops individuals, which it most likely does not. Under the frill, at the rear of the skull, a huge occipital condyle, up to 106 millimeters (4.2 in) in diameter, connected the head to the neck.***

***The lower jaws were elongated and met at their tips in a shared epidentary bone, the core of the toothless lower beak. In the dentary bone, the tooth battery curved to the outside to meet the battery of the upper jaw. At the rear of the lower jaw, the articular bone was exceptionally wide, matching the general width of the jaw joint. T. horridus can be distinguished from T. prorsus by having a shallower snout.***

### ***Postcranial skeleton[edit]***

***Chasmosaurines showed little variation in their postcranial skeleton. The skeleton of Triceratops is markedly robust. Both Triceratops species possessed a very sturdy build, with strong limbs, short hands with three hooves each, and short feet with four hooves each. The vertebral column consisted of ten neck, twelve back, ten sacral, and about forty-five tail vertebrae. The front neck vertebrae were fused into a syncervical. Traditionally, this was assumed to have incorporated the first three vertebrae, thus implying that the frontmost atlas was very large and sported a neural spine. Later interpretations revived an old hypothesis by John Bell Hatcher that, at the very front, a vestige of the real atlas can be observed, the syncervical then consisting of four vertebrae. The vertebral count mentioned is adjusted to this view. In Triceratops, the neural spines of the neck are constant in height and don't gradually slope upwards. Another peculiarity is that the neck ribs only begin to lengthen with the ninth cervical vertebra.***

***The rather short and high vertebrae of the back were, in its middle region, reinforced by ossified tendons running along the tops of the neural arches. The straight sacrum was long and adult individuals show a fusion of all sacral vertebrae. In Triceratops the first four and last two sacrals had transverse processes, connecting the vertebral column to the pelvis, that were fused at their distal ends. Sacrals seven and eight had longer processes, causing the sacrum to have an oval profile in top view. On top of the sacrum, a neural plate was present formed by a fusion of the neural spines of the second through fifth vertebrae. Triceratops had a large pelvis with a long ilium. The ischium was curved downwards. The foot was short with four functional toes. The phalangeal formula of the foot is 2-3-4-5-0.***

***Although certainly quadrupedal, the posture of horned dinosaurs has long been the subject of some debate. Originally, it was believed that the front legs of the animal had to be sprawling at a considerable angle from the thorax in order to better bear the weight of the head. This stance can be seen in paintings by Charles Knight and Rudolph Zallinger. Ichnological evidence in the form of trackways from horned dinosaurs and recent reconstructions of skeletons (both physical and digital) seem to show that Triceratops and other ceratopsids maintained an upright stance during normal locomotion, with the elbows flexed to behind and slightly bowed out, in an intermediate state between fully upright and fully sprawling, comparable to the modern rhinoceros.***

***The hands and forearms of Triceratops retained a fairly primitive structure when compared to other quadrupedal dinosaurs, such as thyreophorans and many sauropods. In those two groups, the forelimbs of quadrupedal species were usually rotated so that the hands faced forward with palms backward ("pronated") as the animals walked. Triceratops, like other ceratopsians and related quadrupedal ornithopods (together forming the Cerapoda), walked with most of their fingers pointing out and away from the body, the original condition for dinosaurs. This was also retained by bipedal forms, like theropods. In Triceratops, the weight of the body was carried by only the first three fingers of the hand, while digits 4 and 5 were vestigial and lacked claws or hooves. The phalangeal formula of the hand is 2-3-4-3-1, meaning that the first or innermost finger of the forelimb has two bones, the next has three, the next has four, etc.***

### ***Skin[edit]***

***Preserved skin from Triceratops is known. This skin consist of large scales, some of which exceed 100 millimetres (3.9 in) across, which have conical projections rising from their center. A preserved piece of skin from the frill of a specimen is also known, which consists of small polygonal basement scales.***

## ***Classification[edit]***

Triceratops is the best-known genus of Ceratopsidae, a family of large, mostly North American ceratopsians. The exact relationship of Triceratops among the other ceratopsids has been debated over the years. Confusion stemmed mainly from the combination of a short, solid frill (similar to that of Centrosaurinae), with long brow horns (more akin to Chasmosaurinae). In the first overview of ceratopsians, R. S. Lull hypothesized the existence of two lineages, one of Monoclonius and Centrosaurus leading to Triceratops, the other with Ceratops and Torosaurus, making Triceratops a centrosaurine as the group is understood today. Later revisions supported this view when Lawrence Lambe, in 1915, formally describing the first, short-frilled group as Centrosaurinae (including Triceratops), and the second, long-frilled group as Chasmosaurinae.

In 1949, Charles Mortram Sternberg was the first to question this position, proposing instead that Triceratops was more closely related to Arrhinoceratops and Chasmosaurus based on skull and horn features, making Triceratops a chasmosaurine ("ceratopsine" in his usage) genus. He was largely ignored, with John Ostrom and later David Norman placing Triceratops within the Centrosaurinae.

Subsequent discoveries and analyses, however, proved the correctness of Sternberg's view on the position of Triceratops, with Thomas Lehman defining both subfamilies in 1990 and diagnosing Triceratops as "ceratopsine" on the basis of several morphological features. Apart from the one feature of a shortened frill, Triceratops shares no derived traits with centrosaurines. Further research by Peter Dodson, including a 1990 cladistic analysis and a 1993 study using resistant-fit theta-rho analysis, or RFTRA (a morphometric technique which systematically measures similarities in skull shape), reinforces Triceratops' placement as a chasmosaurine.

The cladogram below follows Longrich (2014), who named a new species of Pentaceratops, and included nearly all species of chasmosaurine.

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