Moschops oweni

Moschops (Greek for "calf face") is an extinct genus of therapsids that lived in the Guadalupian epoch,
 around 265–260 million years ago. They were heavily built plant eaters,
 and they may have lived partly in water, as hippopotamuses do. They had
 short, thick heads and might have competed by head-butting each other.
Their elbow joints allowed them to walk with a more mammal-like gait
rather than crawling. Their remains were found in the Karoo region of South Africa, belonging to the Tapinocephalus Assemblage Zone. Therapsids, such as Moschops, are synapsids, the dominant land animals in the Permian period, which ended 252 million years ago.

Moschops is characterized by a strongly pachyostosed skull with a broad intertemporal region and greatly reduced temporal fossae.
 Two species are known from good fossil record, M. capensis and M.
koupensis. Two other species were assigned (M. whaitsi and M. oweni),
but the validity is doubtful.

Genera regarded as synonyms are Moschoides, Agnosaurus, Moschognathus and Pnigalion.

Another
 taxon, Delphinognathus conocephalus, could have been a synonym also, as
 it could represent a juvenile Moschops. This taxon is known from a
single, moderately pachyostosed skull. It has a conical boss on the
parietal surrounding the pineal foramen.

Moschops was a roughly 2.7 metres (8.9 ft), massively built dinocephalian. It had a short, thick and massive head, which was broad across the orbits. Like other members of Tapinocephalidae, the skull had a tiny opening for the pineal organ. The occiput was broad and deep, but the skull was more narrow in the dorsal border. Furthermore, the pterygoid
 arches and the angular region of the jaw were quite heavy, allowing the
 insertion of strong jaw muscles. Due to that and because it possessed
long-crowned, stout teeth, it is believed that Moschops was a herbivore
feeding on nutrient-poor and tough vegetation, like cycad
 stems. Due to the presumably nutrient-poor food, it very likely had to
feed for a very long time. Its anatomy allowed Moschops to open its
elbow joint more widely, enabling it to move in a more mammal-like way
than the other crawlers in its time. That might have helped it to carry
its massive body more easily while feeding. Very likely, most
dinocephalians were rather slow-moving animals, but capable of raising
themselves, for short bursts. It is also possible that Moschops and
other dinocephalians were semiaquatic, given the heavy build and the
limbs with their spreading hands and feet. The heavy head could have
been useful for diving after food.

Moschops had a thick skull,
prompting speculation that individuals competed with one another by
head-butting. Some doubt whether the Moschops were born with thick
skulls. If they were, then Moschops' short, heavy tail may have
counterbalanced the weight of its head . Its main enemies were very
likely titanosuchids and the larger therocephalians.

Moschops material was discovered for the first time by Robert Broom in the Ecca Group (part of the Karoo Supergroup) in South Africa. The geological horizon dubious, it was referred to that group on the basis of Pareiasaurus remains in near proximity. The material includes a holotype (AMNH 5550) and seven topotypes (AMNH 5551-5557). The degree of pachyostosis
 varies within the skulls of the specimens. According to Broom, it is
because of gender and age variation within the discovered specimens. In
1910, the material was sent to the American Museum of Natural History in New York City and it was described in 1911.