Leopard

Template:Other uses
Template:Pp-move-indef
Template:Speciesbox

The **leopard** (*Panthera pardus*) Template:IPAc-en is one of the five "big cats" in the genus *Panthera*. It is a member of the family Felidae with a wide range in sub-Saharan Africa and parts of Asia. Fossil records suggest that in the Late Pleistocene it occurred in Europe and Japan.

Compared to other members of Felidae, the leopard has relatively short legs and a long body with a large skull. It is similar in appearance to the jaguar, but has a smaller, lighter physique. Its fur is marked with rosettes similar to those of the jaguar, but the leopard's rosettes are smaller and more densely packed, and do not usually have central spots as the jaguar's do. Both leopards and jaguars that are melanistic are known as black panthers.

The leopard is distinguished by its well-camouflaged fur, opportunistic hunting behaviour, broad diet, and strength (which it uses to move heavy carcasses into trees), as well as its ability to adapt to various habitats ranging from rainforest to steppe, including arid and montane areas, and its ability to run at speeds of up to Template:Convert.

The leopard is listed as vulnerable on the IUCN Red List because leopard populations are threatened by habitat loss and fragmentation, and are declining in large parts of the global range. In Hong Kong, Singapore, Kuwait, Syria, Libya, Tunisia and most likely in Morocco, leopards have already been extirpated.
Leopards are hunted illegally, and their body parts are smuggled in the wildlife trade for medicinal practices and decoration.

## Etymology

The common name "leopard" Template:IPAc-en is a Greek compound of Template:Lang *leōn* ("lion") and Template:Lang *pardos* ("male panther"). The name reflects the fact that in antiquity, a leopard was believed to be a hybrid of a lion and a panther. The Greek word is related to Sanskrit Template:Lang Template:IAST ("snake", "tiger" or "panther"), and probably derives from a Mediterranean language, such as Egyptian. The name was first used in the 13th century. Other vernacular names for the leopard include graupanther, panther and several regional names such as *tendwa* in India. The term "black panther" refers to leopards with melanistic genes.

The scientific name of the leopard is *Panthera pardus*. The generic name *Panthera* derives from Latin via Greek πάνθηρ (*pánthēr*). The term "panther", whose first recorded use dates back to the 13th century AD, generally refers to the leopard, and less often to the cougar and the jaguar. Alternative origins suggested for *Panthera* include an Indo-Iranian word meaning "white-yellow" or "pale". In Sanskrit, this could have been derived from पाण्डर *pāṇḍara* ("tiger"), which in turn comes from पुण्डरीक *puṇḍárīka* (with the same meaning). The specific name *pardus* is derived from the Greek Template:Lang (*pardos*) ("male panther").

## Taxonomy

 File:Two cladograms for Panthera (including Neofelis).png
The leopard is one of the five extant species of the genus *Panthera*, which also includes the jaguar (*P. onca*), the lion (*P. leo*), the snow leopard (*P. uncia*; sometimes placed in *Uncia*, a separate genus of its own) and the tiger (*P. tigris*). This genus, along with the genus *Neofelis* - which consists of the clouded leopard (*N. nebulosa*) and the Sunda clouded leopard (*N. dardi*) - forms the subfamily Pantherinae of the Felidae. The leopard was first described by Swedish zoologist Carl Linnaeus in the 10th edition of *Systema Naturae* (1758). Linnaeus named the leopard as *Felis pardus*, placing it in the genus *Felis* along with the domestic cat, the jaguar, the Eurasian lynx, the lion, the ocelot and the tiger. In the 18th and 19th centuries, most naturalists and taxonomists followed his example. In 1816, Lorenz Oken proposed a definition of the genus *Panthera*, with a subgenus Panthera using *F. pardus* as a type species. Oken's classification, however, was not widely accepted, and until the early 20th century continued using *Felis* or *Leopardus* when describing leopard subspecies. In 1916, British zoologist Reginald Innes Pocock accorded *Panthera* generic rank defining *Panthera pardus* as species.

The leopard is part of the *Panthera* lineage, one of the eight lineages of Felidae. This lineage comprises the species of *Panthera* and *Neofelis*. The clouded leopard diverged first from the lineage, followed by the snow leopard. Subsequent branching began two to three million years ago, but the details of this are disputed.
 A 2006 phylogenetic study by Warren E. Johnson (of the National Cancer Institute) and colleagues, based on nDNA and mtDNA analysis, showed that the leopard is sister to two clades within *Panthera* - one consisting of the tiger and the snow leopard, and the other of the lion and the jaguar. This was seconded by a 2009 study by Lars Werdelin and colleagues. However, the results obtained in a 2010 study by Brian W. Davis (of the Texas A&M University) and colleagues and a 2011 study by Ji H. Mazák (of the Shanghai Science and Technology Museum) and colleagues showed a swapping between the leopard and the jaguar in the cladogram. Results of a 2001 phylogenetic analysis of chemical secretions amongst cats suggested, however, that the leopard is closely related to the lion.

### Subspecies

 File:Panthera pardus subspecies map.png
As many as 27 leopard subspecies were subsequently described by naturalists from 1794 to 1956. Since 1996, only eight subspecies have been considered valid on the basis of mitochondrial analysis. Later analysis revealed a ninth valid subspecies, the Arabian leopard. However, this number might be an underestimation; limited sampling of the African leopards means that different subspecies may not have been resolved.

The nine subspecies recognised by the International Union for Conservation of Nature (IUCN) are summarised in the following table.

Template:Clear

A morphological analysis of characters of leopard skulls implies the validity of two more subspecies:

* Anatolian leopard (*P. p. tulliana*) (Valenciennes, 1856) inhabited western Turkey until the 1970s
* Balochistan leopard (*P. p. sindica*) (Pocock, 1930) possibly evolved in southern Iran, southern Afghanistan and south-western Pakistan being separated from the northern population by Dasht-e Kavir and Dasht-e Lut deserts

Template:Clear
The table below shows the 27 leopard metapopulations that are either currently recognised as subspecies or recently amalgamated into other subspecies due to reassessments based on molecular analysis of the leopard's population genetics.

The smallest leopard subspecies is the Arabian leopard. Adult females weigh as little as Template:Convert. Large subspecies, in which males weigh up to Template:Convert, are the Sri Lankan leopard and the Persian leopard. Such larger leopards inhabit areas which lack tigers and lions, so that leopards are at the top of the food chain with no competitive restriction from large prey.

## Evolution

The last common ancestor of the *Panthera* and *Neofelis* species is believed to have occurred about 6.37 million years ago. The clouded leopard was the first to diverge from the rest of the *Panthera* lineage, followed by the snow leopard. The genus *Panthera* is believed to have emerged in Asia, from where they subsequently emigrated to Africa. The tiger-snow leopard clade diverged from the rest of *Panthera* around 2.9 million years ago. Johnson and colleagues suggest that the leopard diverged next, and followed by the lion-jaguar clade.

Fossils of ancestors of the leopard have been found in East Africa and South Asia, dating back to the Pleistocene between 2 and 3.5 million years ago. The modern leopard is suggested to have evolved in Africa 0.5 to 0.8 million years ago and to have radiated across Asia 0.2 to 0.3 million years ago.

In Europe, the leopard is known at least since the Pleistocene. Fossil bones and teeth dating from the Pliocene were found in Perrier in France, northeast of London, and in Valdarno (Italy). Similar fossils dating back to the Pleistocene were excavated mostly in loess and caves at 40 sites in the continent - from near Lisbon, near Gibraltar, and Santander Province in northern Spain to several sites in France, Switzerland, Italy, Austria, Germany, in the north up to Derby in England, in the east to Přerov in the Czech Republic, and the Baranya in southern Hungary. The Pleistocene leopards of Europe can be divided into four subsequent subspecies. The first European leopard subspecies *P. p. begoueni* is known from the beginning of the early Pleistocene and was replaced about 0.6 million years ago by *P. p. sickenbergi*, which in turn was replaced by *P. p. antiqua* around 0.3 million years ago. The most recent form, the Late Pleistocene Ice Age leopard (*P. p. spelaea*), appeared at the beginning of the Late Pleistocene and survived until about 24,000 years ago in several parts of Europe.

Pleistocene fossils have also been excavated in the Japanese Archipelago.

## Genetics

The diploid number of chromosomes in the leopard is 38, the same as in any other felid, save for the ocelot and the margay, whose diploid number of chromosomes is 36. The chromosomes include four acrocentric, five metacentric, seven submetacentric and two telocentric pairs.

### Hybrids

 File:Pumapard-1904.jpg
Template:Main article
Crossbreeding between the leopard and the other members of the *Panthera* has been documented. In 1953, a lioness and a male leopard were mated in the Hanshin Park in Nishinomiya, Japan. The first litter from this pairing was born on 2 November 1959, consisting of a male and a female. Another litter was born in 1961, in which all the offspring were spotted and bigger than a juvenile leopard. The hybrid came to be known as "leopon". Unsuccessful attempts were made to mate a leopon with a tigress.

Although lions and leopards may come into contact in sub-Saharan Africa, they are generally not known to interbreed naturally. However, there have been anecdotal reports of felids larger than the cheetah but smaller than the lion, with a lion-like face, from the Central African Republic, Kenya, Rwanda and Uganda. This animal, known as the marozi and by several other names, is covered with grayish spots or rosettes on the back, the flanks and the legs. However, there have been no confirmed sightings of the marozi since the 1930s.

A pumapard is a hybrid animal resulting from a mating between a leopard and a puma (a member of the genus *Puma*, not the genus *Panthera*). Three sets of these hybrids were bred in the late 1890s and early 1900s by Carl Hagenbeck at his animal park in Hamburg, Germany. While most of these animals did not reach adulthood, one of these was purchased in 1898 by the Berlin Zoo. A similar hybrid in the Berlin Zoo purchased from Hagenbeck was a cross between a male leopard and a female puma. A specimen in the Hamburg Zoo (in the photo at right) was the reverse pairing, fathered by a puma bred to an Indian leopardess. The pumapard is characterised by a long body like the puma's, but with shorter legs. The hybrid is in general a dwarf, smaller than either parent. The coat is variously described as sandy, tawny or greyish with brown, chestnut or faded rosettes.

### Variant colouration

 File:Blackleopard.JPG
Template:Main article
Melanistic leopards are known, like melanistic jaguars, as "black panthers". Pseudomelanism (abundism) also occurs in leopards.
Melanism in leopards is inherited as a trait relatively recessive to the spotted form. Interbreeding in melanistic leopards produces a significantly smaller litter size than is produced by normal pairings.

The black panther is common in the equatorial rainforest of Malaya and the tropical rainforest on the slopes of some African mountains such as Mount Kenya.
Between January 1996 and March 2009, Indochinese leopards were photographed at 16 sites in the Malay Peninsula in a sampling effort of more than 1000 trap nights. Of the 445 photographs of melanistic leopards, 410 came from study sites south of the Kra Isthmus, where the non-melanistic morph was never photographed. This data suggests the near fixation of the dark allele in the region. The expected time for the fixation of this recessive allele due to genetic drift alone ranged from about 1,100 years to about 100,000 years.

A rare "strawberry" leopard was photographed in South Africa's Madikwe Game Reserve. This condition was probably caused by erythrism, a little-understood genetic condition that causes either an overproduction of red pigments or an underproduction of dark pigments. Pseudomelanism has also been reported in leopards.

## Characteristics

 File:Leopard africa.jpg
The leopard is a big cat distinguished by its robust build and muscular but relatively shorter limbs, a broad head and a coat covered by spots arranged in rosettes. Males stand Template:Convert at the shoulder, while females are Template:Convert tall. The head-and-body length is typically between Template:Convert. While males weigh Template:Convert, females weigh Template:Convert; these measurements vary geographically. The maximum recorded weight for a leopard is Template:Convert. Sexually dimorphic, males are larger and heavier than females.

Basically pale yellow to yellowish brown or golden (except for the melanistic forms), the coat is spotted and rosetted; spots fade toward the white underbelly and the insides and lower parts of the legs. Rosettes are most prominent on the back, flanks and hindquarters. The pattern of the rosettes is unique to each individual. Juveniles have woolly fur, and appear dark due to the densely arranged spots. The white-tipped tail, Template:Convert long, white underneath, displays rosettes except toward the end, where the spots form incomplete bands.

The texture and colour of the fur varies by climate and geography; leopards in forests are observed to be darker than those in deserts. The guard hairs (the layer of hairs that protect the basal hairs) are the shortest (Template:Convert) on the face and the head, and increase in length toward the flanks and the underparts (Template:Convert). The fur is generally soft and thick; the fur on the underparts is notably softer than that on the back. A few geographical variations have been noted in the colour and texture of the fur. Leopards in forests tend to be darker than those in deserts; the fur tends to grow longer in populations living in colder climates. The rosettes, circular in eastern African populations, tend to be squarish in southern Africa and larger in Asian populations. Their yellow coat tends to be more pale and cream coloured in desert populations, more gray in colder climates, and of a darker golden hue in rainforest habitats.

The leopard is often confused with the cheetah; however, the cheetah is marked with small round spots instead of the larger rosettes. Moreover, the leopard lacks the facial tear streaks characteristic of the cheetah. Other similar species are the clouded leopard and jaguar. The clouded leopard can be told apart by the diffuse "clouds" of spots compared to the smaller and distinct rosettes of the leopard, longer legs and thinner tail. The jaguar has rosettes that typically have spots within them, while those of leopards often do not. Moreover, the jaguar has larger and rounder foot pads and a larger and stronger skull.

## Distribution and habitat

 File:Sousse museum Smirat-retouched.jpg
 File:A leopard on the tree in the Serengeti Plain.JPG
Leopards have the largest distribution of any wild cat, occurring widely in Africa as well as eastern and southern Asia, although populations have shown a declining trend, and are fragmented outside of sub-Saharan Africa. Within sub-Saharan Africa, the species is still numerous and even thriving in marginal habitats where other large cats have disappeared. Populations in North Africa may be extinct. Data on their distribution in Asia are not consistent. Populations in southwest and central Asia are small and fragmented; in the northeast, they are critically endangered. In the Indian subcontinent, Southeast Asia, and China, leopards are still relatively abundant. Of the species as a whole, its numbers are greater than those of other *Panthera* species, all of which face more acute conservation concerns.

Leopards are exceptionally adaptable, although associated primarily with savanna and rainforest. Populations thrive anywhere in the species range where grasslands, woodlands, and riverine forests remain largely undisturbed. In the Russian Far East, they inhabit temperate forests where winter temperatures reach a low of Template:Convert. They are equally adept surviving in some of the world's most humid rainforests and even semi-arid desert edges.

Leopards in west and central Asia try to avoid deserts, areas with long-duration snow cover and areas that are near urban development. In India, leopard populations sometimes live quite close to human settlements and even in semi-developed areas. Although occasionally adaptable to human disturbances, leopards require healthy prey populations and appropriate vegetative cover for hunting for prolonged survival and thus rarely linger in heavily developed areas. Due to the leopard's superlative stealthiness, people often remain unaware that big cats live in nearby areas.

## Ecology and behaviour

 File:Leopard on the tree.jpg
Leopards, like lions and tigers, tend to be nocturnal (active mainly at night). However, leopards in western African forests have been observed to be largely diurnal and hunt during twilight, when their prey animals are active; activity patterns may even vary by season. Leopards generally are active mainly from dusk till dawn, and rest for most of the day and for some hours at night in thickets, among rocks or over tree branches. Leopards have been observed walking Template:Convert across their range at night; they may even wander up to Template:Convert if disturbed.

Leopards are known for their ability to climb and have been observed resting on tree branches during the day, dragging their kills up trees and hanging them there, and descending from trees headfirst. They are powerful swimmers, although are not as disposed to swimming as some other big cats, such as the tiger. They are very agile, and can run at over Template:Convert, leap over Template:Convert horizontally, and jump up to Template:Convert vertically.

### Social spacing

Template:Multiple image

The leopard is solitary and territorial, as are several other felids; individuals associate appreciably only in the mating season, though mothers may continue to interact with their offspring even after weaning. Mothers have been observed sharing kills with their offspring when they can not obtain any meal. In Kruger National Park, most leopards tend to keep Template:Convert apart. Fathers may interact with their partners and cubs at times. Aggressive encounters are rare, typically limited to defending territories from intruders. In a South African reserve, a male was wounded in a male–male territorial battle over a carcass. A few instances of cannibalism have been reported.

Leopards communicate with each other in tall grass using white spots on their ears and tails. They also produce a number of vocalisations, including growls, snarls, meows and purrs. The roaring sequence in leopards consists mainly of grunts and is also known called "sawing", having been described as resembling the sound of sawing wood. Cubs are known to call their mother with a *urr-urr* sound.

Males occupy territories that often overlap with a few smaller female territories, probably as a strategy to enhance access to females. A radio-collar analysis in the Ivory Coast found a female home range completely enclosed within a male's. Female live with their cubs in territories that overlap extensivelyTemplate:Sndprobably due to the association between mothers and their offspring. There may be a few other fluctuating territories, belonging to young individuals. It is not clear if male territories tend to overlap among themselves as much as those of females do. Individuals will try to drive away intruders of the same sex.

A study of leopards in the Namibian farmlands showed that the size of territories was not significantly affected by sex, rainfall patterns or season; it concluded that the higher the prey availability in an area, the greater the population density of leopards and the smaller the size of territories, but territories tend to expand if there is human interference (which has been notably high in the study area). Territorial sizes vary geographically; they can be as small as Template:Convert for males and Template:Convert for females in forests and rocky terrain (such as in the Serengeti or Kruger National Park), or as large as Template:Convert for males and Template:Convert for females in northeastern Namibia (they might be even larger in deserts and montane areas). Territories recorded in Nepal, Template:Convert for males and Template:Convert for females, are smaller than those generally observed in Africa.

### Hunting and diet

Template:Multiple image

The leopard is a carnivore that prefers medium-sized prey with a body mass ranging from Template:Convert. A study noted that prey animals in this weight range tend to occur in dense habitat, form small herds and can be easily captured by the leopard; on the other hand, animals that prefer open areas and have developed significant anti-predator strategies are hardly preferred. Prey as heavy as Template:Convert (such as greater kudu and giraffe) may be hunted if larger carnivores such as lions or tigers are absent; the largest prey killed by a leopard was reportedly a Template:Convert male eland. Leopards can feed on a broad variety of prey, mainly antelopes, deer and rodents; these include: cattle, chital, duiker, dung beetle, hartebeest, hyrax, impala, gazelle, muntjac, nyala, porcupine, primates, rat, reedbuck, springbok, sambar, squirrel, waterbuck, young zebras, warthog and wildebeest. Females with cubs primarily target smaller prey.

A study at Wolong Reserve in China demonstrated variation in the leopards' diet over time; over the course of seven years, the vegetative cover receded, and the animals opportunistically shifted from primarily consuming tufted deer to pursuing bamboo rats and other smaller prey. A study estimated average daily consumption rates at Template:Convert for males and Template:Convert for females. A study of leopards in the southern Kalahari showed that water requirements are met by the bodily fluids of the prey, succulent plants and water bodies; they drink water every two to three days, and feed infrequently on moisture-rich plants such as gemsbok cucumbers (*Acanthosicyos naudinianus*), tsamma melon (*Citrullus lanatus*) and Kalahari sour grass (*Schmidtia kalahariensis*). A few instances of cannibalism have been reported.

The leopard depends mainly on its acute sense of hearing and vision for hunting. Hunting is primarily a nocturnal activity in most areas, though leopards in western African forests and Tsavo have been observed hunting by the day. It will stalk the prey and try to approach as close as possible (typically within Template:Convert) to the target, and finally pounce on it with its forepaws (unlike the lion, that pounces as the prey starts escaping) and kill it by suffocation. Small prey are killed with a bite on the back of the neck, while larger animals are held strongly by the throat and strangled.

Small kills are eaten immediately, while larger carcasses are dragged over several hundred metres and safely cached to be consumed later on trees, in bushes or even caves. The way the kill is stored to be consumed later depends on the local topography and individual preferences; while trees are preferred in Kruger National Park, bushes are preferred in the plain terrain of the Kalahari. Kills are cached up to Template:Convert apart. Although they are smaller than most other members of its genus, leopards are able to take large prey due to their massive skulls that facilitate powerful jaw muscles. Leopards are strong enough to drag carcasses heavier than themselves up trees; an individual was seen to haul a young giraffe, nearly Template:Convert, up Template:Convert into a tree.

### Enemies and competitors

Leopards must compete for food and shelter with other large predators such as tigers, lions, cheetahs, spotted hyenas, striped hyenas, brown hyenas, up to five species of bear and both African and Asiatic wild dogs. These animals may steal the leopard's kill, devour its young or even kill adult leopards. Leopards co-exist alongside these other large predators by hunting for different types of prey and by avoiding areas frequented by them. Leopards may also retreat up a tree in the face of direct aggression from other large carnivores but leopards have been seen to either kill or prey on competitors such as black-backed jackal, African wild cat and the cubs of lions, cheetahs, hyenas, and wild dogs.

 File:Lioness vs Leopard 9 July 2016 Latest Sightings 1.png
Resource partitioning occurs where leopards share their range with tigers. Leopards tend to take smaller prey, usually less than Template:Convert, where tigers are present. In areas where the leopard is sympatric with the tiger, coexistence is reportedly not the general rule, with leopards being few where tigers are numerous. The mean leopard density decreased significantly (from 9.76 to 2.07 animals per 100 km2) when the mean density of tigers increased (from 3.31 animals/100km2 to 5.81 animals/100km2) from 2004–5 to 2007–8 in the Rajaji National Park in India following the relocation of pastoralists out of the park. There, the two species have high dietary overlap, and an increase in the tiger population resulted in a sharp decrease in the leopard population and a shift in the leopard diet to small prey (from 9% to 36%) and domestic prey (from 6.8% to 31.8%). In the Primore region of the Russian Far East, Amur leopards were absent or very rarely encountered at places where Siberian tigers reside. However, in the Chitwan National Park in Nepal, both species coexist because there is a large prey biomass, a large proportion of prey is of the smaller sizes, and dense vegetation exists. Here leopards killed prey ranging from less than Template:Convert to Template:Convert in weight with most kills in the Template:Convert range; tigers killed more prey in the Template:Convert range. There were also differences in the microhabitat preferences of the individual tiger and leopard followed over 5-month (December to April) period in this study - the tiger used roads and (except in February) forested areas more frequently, while the leopard used recently burned areas and open areas more frequently  Usually when a tiger began to kill baits at sites formerly frequented by leopards, the leopards would no longer come and kill there. In the tropical forests of India's Nagarhole National Park, tigers selected prey weighing more than Template:Convert, whereas leopards selected prey in the Template:Convert range. In tropical forest they do not always avoid the larger cats by hunting at different times. With relatively abundant prey and differences in the size of prey selected, tigers and leopards seem to successfully coexist without competitive exclusion or inter-species dominance hierarchies that may be more common to the leopard's co-existence with the lion in savanna habitats. In areas with high tiger populations, such as in the central parts of India's Kanha National Park, leopards are not permanent residents, but transients. They were common near villages at the periphery of the park and outside the park.

In some areas of Africa, troops of large baboon species (potential leopard prey themselves) will kill and sometimes eat leopard young if they discover them. Occasionally, Nile crocodiles may prey on leopards of any age. For example, one large adult leopard was grabbed and consumed by a large crocodile while attempting to hunt along a bank in Kruger National Park. Mugger crocodiles may also on rare occasions kill an adult leopard in India. Lions are occasionally successful in climbing trees and fetching leopard kills, but leopards are also known to kill or prey on cubs of lions. In the Kalahari desert, leopards frequently lose kills to the brown hyena, if the leopard is unable to move the kill into a tree. Single brown hyenas have been observed charging at and displacing male leopards from kills. Burmese pythons have been known to prey on leopards, with an adult cat having been recovered from the stomach of a Template:Convert specimen.

Two cases of leopards killing cheetahs have been reported in 2014.

### Reproduction and life cycle

Depending on the region, leopards may mate all year round. In Manchuria and Siberia, they mate during January and February. The estrous cycle lasts about 46 days and the female usually is in heat for 6–7 days. Gestation lasts for 90 to 105 days. Cubs are usually born in a litter of 2–4 cubs. Mortality of cubs is estimated at 41–50% during the first year.

Females give birth in a cave, crevice among boulders, hollow tree, or thicket to make a den. Cubs are born with closed eyes, which open four to nine days after birth. The fur of the young tends to be longer and thicker than that of adults. Their pelage is also more gray in colour with less defined spots. Around three months of age, the young begin to follow the mother on hunts. At one year of age, leopard young can probably fend for themselves, but remain with the mother for 18–24 months.

The average typical life span of a leopard is between 12 and 17 years. The oldest recorded spotted leopard was a female named Roxanne living in captivity at McCarthy's Wildlife Sanctuary in The Acreage, Palm Beach County, Florida. She died August 8, 2014 at the age of 24 years, 2 months and 13 days. This has been verified by the Guinness World Records. Previously, the oldest recorded leopard was a female named Bertie living in captivity in Warsaw Zoo. She died in December 2010 at the age of 24. The oldest recorded male leopard was Cezar, who reached the age of 23. He also lived at Warsaw Zoo and was Bertie's lifelong companion.

Template:Anchor

## Leopards and humans

 File:Benin, acquamanile a forma di leopardo, XVIII sec.JPG
Leopards have been known to humans throughout history, and have featured in the art, mythology, and folklore of many countries where they have historically occurred, such as ancient Greece, Persia, and Rome, as well as some where they have not existed for several millennia, such as England. The modern use of the leopard as an emblem for sport or a coat of arms is much more restricted to Africa, though numerous products worldwide have used the name. During the Benin Empire, the leopard was commonly represented on engravings and sculptures and was used to symbolise the power of the king or *oba*; since the leopard was considered the king of the forest. Leopard were also kept and paraded as mascots, totems and sacrifices to deities. As a result of their association with kings in Africa, the leopard's pelt is often seen today as a symbol of aristocratic rank, chiefs using it as a part of their traditional regalia.

 File:Flag of Shropshire.svg
The lion passant guardant or leopard is a frequently used charge in heraldry, most commonly appearing in groups of three. The heraldic leopard lacks spots and sports a mane, making it visually almost identical to the heraldic lion, and the two are often used interchangeably. These traditional lions passant guardant appear in the coat of arms of England and many of its former colonies; more modern naturalistic (leopard-like) depictions appear on the coat of arms of several African nations including Benin, Malawi, Somalia, the Democratic Republic of the Congo and Gabon, which uses a black panther.

Leopard domestication has also been recorded—several leopards were kept in a menagerie established by King John at the Tower of London in the 13th century; around 1235, three of these animals were given to Henry III by Holy Roman Emperor Frederick II.

### Tourism

 File:Leopard near driver.jpg
In protected areas of several countries, wildlife touring programs and safari ventures offer sightings of leopards in their natural habitat. While luxury establishments may boast the fact that wild animals can be seen at close range on a daily basis, the leopard's camouflage and propensity to hide and stalk prey typically make leopard sightings rare. In Sri Lanka's Yala National Park, leopards have been ranked by visitors to be among the least visible of all animals in the park despite their high concentration in the reserve.

In South Africa, safaris are offered in the Sabi Sand Game Reserve. In Sri Lanka, wildlife tours are available in the Yala and Wilpattu National Parks. In India, safaris are offered in the Madhya Pradesh and Uttarakhand national parks as well as in the Pali district of western Rajasthan.

### Man-eating

Template:Main article

 File:PanarManeater.jpg
Most leopards avoid people, but humans may occasionally be targeted as prey. Most healthy leopards prefer wild prey to humans, but injured, sickly, or struggling cats or those with a shortage of regular prey may resort to hunting humans and become habituated to it. Although usually slightly smaller than humans, an adult leopard is much more powerful and easily capable of killing them. Two extreme cases occurred in India: the first leopard, "the Leopard of Rudraprayag", killed more than 125 people; the second, the "Panar Leopard", was believed to have killed more than 400. Both were killed by the renowned hunter and conservationist Jim Corbett. Man-eating leopards are considered bold and difficult to track by feline standards and may enter human settlements for prey, more so than lions and tigers. Author and big game hunter Kenneth Anderson had first-hand experience with many man-eating leopards, and described them as far more threatening than tigers:

Template:Quote

Template:Quote

## See also

* List of solitary animals

Template:Portal bar

## Further reading

Template:Refbegin

* Template:Cite book
* Template:Cite journal
* Template:Cite journal
* Template:Cite web
* Template:Cite book
* Template:Cite journal
* Template:Cite book
* Template:Cite journal
* Template:Cite book
* Template:Cite journal

Template:Refend

## External links

Template:Wikispecies
Template:Commons+cat

* IUCN/SSC Cat Specialist Group : *Panthera pardus* in Africa and *Panthera pardus* in Asia
* Leopard Anthology: Research and conservation of leopards in Asia
* Asian Leopard Specialist Society: Research, conservation and management of Asian leopard subspecies
* The Animal Files: Leopard

Template:Carnivora
Template:Authority control

1. ↑ Template:MSW3 Wozencraft
2. ↑ Template:Cite journal
3. ↑ 3.0 3.1 Template:Cite bookTemplate:Page needed
4. ↑ 4.0 4.1 4.2 4.3 4.4 4.5 Template:Cite book
5. ↑ Template:Cite journal
6. ↑ Template:Cite journal
7. ↑ 7.0 7.1 7.2 Cite error: Invalid `<ref>` tag; no text was provided for refs named `IUCN`
8. ↑ Bergin, D. and Nijman, V. (2014) Open, Unregulated Trade in Wildlife in Morocco's Markets. TRAFFIC Bulletin
9. ↑ Template:Cite journal
10. ↑ 10.0 10.1 Template:MerriamWebsterDictionary
11. ↑ Template:Cite book
12. ↑ 12.0 12.1 Template:Cite book
13. ↑ 13.0 13.1 13.2 13.3 13.4 13.5 13.6 13.7 13.8 Template:Cite journal
14. ↑ 14.0 14.1 Template:MerriamWebsterDictionary
15. ↑ 15.0 15.1 Template:Cite book
16. ↑ 16.0 16.1 16.2 Template:Cite journal
17. ↑ 17.0 17.1 Template:Cite journal
18. ↑ 18.0 18.1 18.2 Template:Cite journal
19. ↑ 19.0 19.1 19.2 Template:Cite journal
20. ↑ Template:MSW3
21. ↑ Template:ITIS
22. ↑ Template:Cite book
23. ↑ Template:Cite book
24. ↑ Template:Cite book
25. ↑ Template:Cite journal
26. ↑ Template:Cite journal
27. ↑ 27.0 27.1 Template:Cite journal
28. ↑ 28.0 28.1 28.2 28.3 28.4 Template:Cite journal
29. ↑ 29.0 29.1 29.2 Template:Cite journal
30. ↑ Template:Cite journal
31. ↑ Spalton, J.A. and H.M. Al Hikmani (2006). The Leopard in the Arabian Peninsula – Distribution and Subspecies Status. *Cat News Special Issue* Special Issue 1: 4–8.
32. ↑ Template:Cite book
33. ↑ Template:Cite journal
34. ↑ Template:Cite journal
35. ↑ Template:Cite book
36. ↑ Template:Cite journal
37. ↑ Template:Cite book
38. ↑ Template:Cite book
39. ↑ Template:Cite web
40. ↑ Template:Cite book
41. ↑ Template:Cite journal
42. ↑ Template:Cite journal
43. ↑ Template:Cite book
44. ↑ Template:Cite journal
45. ↑ Template:Cite journal
46. ↑ Template:Cite news
47. ↑ Template:Cite book
48. ↑ 48.0 48.1 48.2 48.3 48.4 48.5 48.6 48.7 48.8 48.9 Template:Cite book
49. ↑ 49.0 49.1 49.2 49.3 49.4 Template:Cite book
50. ↑ 50.0 50.1 Template:Cite book
51. ↑ Template:Cite book
52. ↑ 52.0 52.1 52.2 52.3 Template:Cite book
53. ↑ 53.0 53.1 53.2 Template:Cite book
54. ↑ Template:Cite book
55. ↑ 55.0 55.1 Template:Cite book
56. ↑ Template:Cite book
57. ↑ Template:Cite book
58. ↑ Template:Cite book
59. ↑ Template:Cite journal
60. ↑ Template:Cite journal
61. ↑ 61.0 61.1 61.2 Template:Cite journal
62. ↑ 62.0 62.1 62.2 Athreya, Vidya (2012-08-16) Living with Leopards Outside Protected Areas in India. conservationindia.org
63. ↑ Template:Cite book
64. ↑ Template:Cite book
65. ↑ Template:Cite journal Template:Open access
66. ↑ Template:Cite journal
67. ↑ Template:Cite journal
68. ↑ 68.0 68.1 Template:Cite web
69. ↑ Template:Cite web
70. ↑ 70.0 70.1 Template:Cite book
71. ↑ Template:Cite journal
72. ↑ 72.0 72.1 Template:Cite journal
73. ↑ Template:Cite journal
74. ↑ Cite error: Invalid `<ref>` tag; no text was provided for refs named `sandiego`
75. ↑ Template:Cite book
76. ↑ Template:Cite journal
77. ↑ Template:Cite journal Template:Open access
78. ↑ Template:Cite journal
79. ↑ Template:Cite journal
80. ↑ Template:Cite journal
81. ↑ Template:Cite journal
82. ↑ 82.0 82.1 82.2 Template:Cite journal
83. ↑ Template:Cite journal
84. ↑ 84.0 84.1 Template:Cite book
85. ↑ Template:Cite journal
86. ↑ Template:Cite journal
87. ↑ Template:Cite journal
88. ↑ 88.0 88.1 Template:Cite book
89. ↑ Template:Cite journal Template:Open access
90. ↑ Template:Cite book
91. ↑ 91.0 91.1 Template:Cite thesis
92. ↑ 92.0 92.1 Template:Cite book
93. ↑ Template:Cite book
94. ↑ Template:Cite journal
95. ↑ 95.0 95.1 Template:Cite journal
96. ↑ Hepnter, V.G.; and Sluskii, A.A. (1992) *Mammals of the Soviet Union*. Vol 2, Part 2. (Carnivores: Hyaenas and Cats) New Delhi: Amerind Publishing. pp. 266–7.
97. ↑ 97.0 97.1 Template:Cite journal
98. ↑ Template:Cite journal
99. ↑ 99.0 99.1 Template:Cite journal
100. ↑ Template:Cite journal
101. ↑ Template:Cite journal
102. ↑ Leopard left for dead by baboon troop. wilderness-safaris.com (2006-10-25).
103. ↑ Template:Cite web
104. ↑ Bailey, T. N. (1993). *The African leopard: ecology and behavior of a solitary felid*. Columbia University Press.
105. ↑ Kingdon, J., Happold, D., Butynski, T., Hoffmann, M., Happold, M., & Kalina, J. (2013). *Mammals of Africa (Vol. 1)*. A&C Black.
106. ↑ Template:Cite journal
107. ↑ Template:Cite book
108. ↑ Template:Cite journal
109. ↑ Template:Cite book
110. ↑ Perowne, J. (2014). Blog: Leopard kills cheetah in the Mara and hoists it up a tree. The Safari Collection, Nairobi.
111. ↑ Hagen, M. (2014). Leopard kills Cheetah. youtube.com
112. ↑ Template:Cite journal
113. ↑ Template:Cite book
114. ↑ Template:Cite journal
115. ↑ Template:Cite web
116. ↑ Salisbury, Susan (2014-08-09) Roxanne, oldest spotted leopard in captivity, dies at Acreage preserve. *The Palm Beach Post*
117. ↑ Template:Cite web
118. ↑ Template:Cite web
119. ↑ Template:Cite web
120. ↑ Template:Cite book
121. ↑ Template:Cite book
122. ↑ Template:Cite web
123. ↑ Template:Cite news
124. ↑ Weerasinghe, U.M.I.R.K., Kariyawasm, D. and M. De Zoysa (2003). Ruhuna (Yala) National Park in Sri Lanka: Visitors, Visitation, and Eco-Tourism. Contribution to the XII World Forestry Congress, Quebec.
125. ↑ Template:Cite news
126. ↑ Template:Cite book
127. ↑ Template:Cite book