Bohemiatupus elegans

Bohemiatupus is an extinct genus
of griffenfly in the family Meganeuridae and containing a single species
 Bohemiatupus elegans. The species is known only from the Late
Carboniferous, Bolsovian stage, Kladno Formation near the village of
Radnice in the Radnice Basin, Czech Republic.

*Bohemiatupus elegans* is known only from one fossil, the holotype, specimen number "M00485" which is composed of an isolated fore-wing and hind-wing. The wings are preserved as a negative imprint fossil in a sedimentary tuff. The fossil specimen is from outcrops of the Kladno Formation exposed in the Ovčín opencast mine. *Bohemiatupus elegans* is the first described occurrence of large griffenflies found in the continental basins of the Bohemian Massif. The type specimen is currently preserved in the collections housed in the West Bohemian Museum, located in Plzeň, Czech Republic. *Bohemiatupus* was first studied by Jakub Prokop of Charles University in Prague, Czech Republic and André Nel of the Muséum national d'histoire naturelle in Paris, France. Their 2008 type description of the genus and species was published in the journal *Annales de la Société Entomologique de France*. The generic name was coined by Prokop and Nel from a combination of the "Bohemia" in reference to the historical central European region and the meganeurid genus *Tupus* to which *Bohemiatupus* is related. The etymology of the specific epithet *elegens* is Latin and refers to the "magnificent state of preservation" displayed by the holotype.

The ecosystem in which *Bohemiatupus elegans* lived is interpreted as a shallow lake that developed into a peat-mire due to sediment infill from volcanic ash during the Bolsovian. *Bohemiatupus elegans* shared this environment with the notably large insect species *Bojophlebia prokopi* and *Carbotriplura kukalovae*, which were described from the Kladno Formation earlier.

The holotype wings are mostly complete with the apical section of the
fore-wing folded and indistinct while only the basal section of the
hind-wing is present. The wings shows no indications of any color pattering that may have been present in life and are interpreted to have been hyaline
 in nature. The preserved segment of the fore-wing is 115 millimetres
(4.5 in) giving an estimated full length for the wing of approximately
260 millimetres (10 in). An adult *Bohemiatupus elegans* would
have had a full wingspan of around 520 millimetres (20 in). The overall
 size larger than the 150 millimetres (5.9 in) wingspan of the older
genus *Sinomeganeura huangheensis*. However this is still smaller than other members of the subfamily such as *Meganeuropsis permiana* which is noted for a wingspan exceeding 700 millimetres (28 in). Several key vein structure features present in the fore-wing show *Bohemiatupus elegans* to be a member of the meganeurid subfamily Tupinae close to the genus *Tupus*.
 The AA1 vein possesses several rearward branches, and the CuP vein has
 a number of concave branches, while the braces CuP and CuA are oblique
but not fused into a single vein. Overall the hind-wing is distinctly
wider in the cubito-anal area than that of *Tupus*, where the hind and fore wings are similar in width. *B. elegans* can be separated from other genera in the subfamily by various aspects of the fine vein structuring in the wing.