Caddis

Many cultures have negative superstitions about cats. An example would be the belief that encountering a black cat ("crossing one's path") leads to bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. The killing of cats in Medieval Ypres, Belgium, is commemorated in the innocuous present-day Kattenstoet (cat parade). In mid-16th century France, cats would be burnt alive as a form of entertainment, particularly during midsummer festivals. According to Norman Davies, the assembled people "shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally carbonized". The remaining ashes were sometimes taken back home by the people for good luck.

According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy, Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives, while in Arabic traditions, the number of lives is six. An early mention of the myth can be found in John Heywood's *The Proverbs of John Heywood* (1546):

> Husband, (quoth she), ye studie, be merrie now,
>
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> And even as ye thinke now, so come to yow.
>
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> Nay not so, (quoth he), for my thought to tell right,
>
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> I thinke how you lay groning, wife, all last night.
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> Husband, *a groning horse and a groning wife*
>
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> *Never faile their master*, (quoth she), for my life.
>
>
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> No wife, *a woman hath nine lives like a cat*.
>
>

The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations.[*citation needed*] Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive righting reflex to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall.

## Lifespan and health

Main articles: Cat health and Aging in cats

The average lifespan of pet cats has risen in recent decades. In the early 1980s, it was about seven years, rising to 9.4 years in 1995 and about 15 years in 2021.[*citation needed*] Some cats have been reported as surviving into their 30s, with the oldest known cat, Creme Puff, dying at a verified age of 38.

Neutering increases life expectancy: one study found castrated male cats live twice as long as intact males, while spayed female cats live 62% longer than intact females. Having a cat neutered confers health benefits, because castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.

### Disease

Main article: List of feline diseases

About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors of metabolism. The high level of similarity among the metabolism of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats as animal models in the study of the human diseases. Diseases affecting domestic cats include acute infections, parasitic infestations, injuries, and chronic diseases such as kidney disease, thyroid disease, and arthritis. Vaccinations are available for many infectious diseases, as are treatments to eliminate parasites such as worms, ticks, and fleas.

## Ecology

### Habitats

The domestic cat is a cosmopolitan species and occurs across much of the world. It is adaptable and now present on all continents except Antarctica, and on 118 of the 131 main groups of islands, even on the isolated Kerguelen Islands. Due to its ability to thrive in almost any terrestrial habitat, it is among the world's most invasive species. It lives on small islands with no human inhabitants. Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas, and wetlands.

The unwantedness that leads to the domestic cat being treated as an invasive species is twofold. On one hand, as it is little altered from the wildcat, it can readily interbreed with the wildcat. This hybridization poses a danger to the genetic distinctiveness of some wildcat populations, particularly in Scotland and Hungary, possibly also the Iberian Peninsula, and where protected natural areas are close to human-dominated landscapes, such as Kruger National Park in South Africa. On the other hand, and perhaps more obviously, its introduction to places where no native felines are present contributes to the decline of native species.

### Ferality

Main article: Feral cat
link=https://en.wikipedia.org/wiki/File:Feral\_cat\_Virginia\_crop.jpg|right|thumb|Feral farm cat
Feral cats are domestic cats that were born in or have reverted to a wild state. They are unfamiliar with and wary of humans and roam freely in urban and rural areas. The numbers of feral cats is not known, but estimates of the United States feral population range from 25 to 60 million. Feral cats may live alone, but most are found in large colonies, which occupy a specific territory and are usually associated with a source of food. Famous feral cat colonies are found in Rome around the Colosseum and Forum Romanum, with cats at some of these sites being fed and given medical attention by volunteers.

### Impact on wildlife

Main article: Cat predation on wildlife

On islands, birds can contribute as much as 60% of a cat's diet. In nearly all cases, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a "mesopredator release" effect; where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are a contributing factor to the decline of many species, a factor that has ultimately led, in some cases, to extinction. The South Island piopio, Chatham rail, and the New Zealand merganser are a few from a long list, with the most extreme case being the flightless Lyall's wren, which was driven to extinction only a few years after its discovery. One feral cat in New Zealand killed 102 New Zealand lesser short-tailed bats in seven days. In the US, feral and free-ranging domestic cats kill an estimated 6.3 – 22.3 billion mammals annually.

In Australia, the impact of cats on mammal populations is even greater than the impact of habitat loss. More than one million reptiles are killed by feral cats each day, representing 258 species. Cats have contributed to the extinction of the Navassa curly-tailed lizard and *Chioninia coctei.*

### History and mythology

Main articles: Cultural depictions of cats and Cats in ancient Egypt

In ancient Egypt, cats were worshipped, and the goddess Bastet often depicted in cat form, sometimes taking on the war-like aspect of a lioness. The Greek historian Herodotus reported that killing a cat was forbidden, and when a household cat died, the entire family mourned and shaved their eyebrows. Families took their dead cats to the sacred city of Bubastis, where they were embalmed and buried in sacred repositories. Herodotus expressed astonishment at the domestic cats in Egypt, because he had only ever seen wildcats.

Ancient Greeks and Romans kept weasels as pets, which were seen as the ideal rodent-killers. The earliest unmistakable evidence of the Greeks having domestic cats comes from two coins from Magna Graecia dating to the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders of Rhegion and Taras respectively, playing with their pet cats. The usual ancient Greek word for 'cat' was *ailouros*, meaning 'thing with the waving tail'. Cats are rarely mentioned in ancient Greek literature. Aristotle remarked in his *History of Animals* that "female cats are naturally lecherous." The Greeks later syncretized their own goddess Artemis with the Egyptian goddess Bastet, adopting Bastet's associations with cats and ascribing them to Artemis. In Ovid's *Metamorphoses*, when the deities flee to Egypt and take animal forms, the goddess Diana turns into a cat.

Cats eventually displaced weasels as the pest control of choice because they were more pleasant to have around the house and were more enthusiastic hunters of mice. During the Middle Ages, many of Artemis's associations with cats were grafted onto the Virgin Mary. Cats are often shown in icons of Annunciation and of the Holy Family and, according to Italian folklore, on the same night that Mary gave birth to Jesus, a cat in Bethlehem gave birth to a kitten. Domestic cats were spread throughout much of the rest of the world during the Age of Discovery, as ships' cats were carried on sailing ships to control shipboard rodents and as good-luck charms.

Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they cannot influence decisions made by humans. In Japan, the *maneki neko* cat is a symbol of good fortune. In Norse mythology, Freyja, the goddess of love, beauty, and fertility, is depicted as riding a chariot drawn by cats. In Jewish legend, the first cat was living in the house of the first man Adam as a pet that got rid of mice. The cat was once partnering with the first dog before the latter broke an oath they had made which resulted in enmity between the descendants of these two animals. It is also written that neither cats nor foxes are represented in the water, while every other animal has an incarnation species in the water. Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated Muhammad had a favorite cat, *Muezza*. He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping on it". The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint, Ahmed ar-Rifa'i, centuries after Muhammad. One of the companions of Muhammad was known as Abu Hurayrah ("father of the kitten"), in reference to his documented affection to cats.

### Vision

link=https://en.wikipedia.org/wiki/File:TapetumLucidum.JPG|right|thumb|Reflection of camera flash from the *tapetum lucidum*
link=https://en.wikipedia.org/wiki/File:Tabby\_cat\_with\_visible\_nictitating\_membrane.jpg|alt=A tabby cat with a third, translucent eyelid covering part of either eye|thumb|A cat's nictitating membrane shown as it blinks
Cats have excellent night vision and can see at only one-sixth the light level required for human vision. This is partly the result of cat eyes having a *tapetum lucidum*, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light. Large pupils are an adaptation to dim light. The domestic cat has slit pupils, which allow it to focus bright light without chromatic aberration. At low light, a cat's pupils expand to cover most of the exposed surface of its eyes. The domestic cat has rather poor color vision and only two types of cone cells, optimized for sensitivity to blue and yellowish green; its ability to distinguish between red and green is limited. A response to middle wavelengths from a system other than the rod cells might be due to a third type of cone. This appears to be an adaptation to low light levels rather than representing true trichromatic vision. Cats also have a nictitating membrane, allowing them to blink without hindering their vision.

### Hearing

The domestic cat's hearing is most acute in the range of 500 Hz to 32 kHz. It can detect an extremely broad range of frequencies ranging from 55 Hz to 79 kHz, whereas humans can only detect frequencies between 20 Hz and 20 kHz. It can hear a range of 10.5 octaves, while humans and dogs can hear ranges of about 9 octaves. Its hearing sensitivity is enhanced by its large movable outer ears, the pinnae, which amplify sounds and help detect the location of a noise. It can detect ultrasound, which enables it to detect ultrasonic calls made by rodent prey. Recent research has shown that cats have socio-spatial cognitive abilities to create mental maps of owners' locations based on hearing owners' voices.

### Smell

Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and a large surface of olfactory mucosa, about 5.8 square centimetres (29⁄32 square inch) in area, which is about twice that of humans. Cats and many other animals have a Jacobson's organ in their mouths that is used in the behavioral process of flehmening. It allows them to sense certain aromas in a way that humans cannot. Cats are sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol, which they use to communicate through urine spraying and marking with scent glands. Many cats also respond strongly to plants that contain nepetalactone, especially catnip, as they can detect that substance at less than one part per billion. About 70–80% of cats are affected by nepetalactone. This response is also produced by other plants, such as silver vine (*Actinidia polygama*) and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.

### Taste

Cats have relatively few taste buds compared to humans (470 or so versus more than 9,000 on the human tongue). Domestic and wild cats share a taste receptor gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness. They, however, possess taste bud receptors specialized for acids, amino acids like protein, and bitter tastes. Their taste buds possess the receptors needed to detect umami. However, these receptors contain molecular changes that make the cat taste of umami different from that of humans. In humans, they detect the amino acids of glutamic acid and aspartic acid, but in cats they instead detect nucleotides, in this case inosine monophosphate and l-Histidine. These nucleotides are particularly enriched in tuna. This has been argued is why cats find tuna so palatable: as put by researchers into cat taste, "the specific combination of the high IMP and free l-Histidine contents of tuna" .. "produces a strong umami taste synergy that is highly preferred by cats". One of the researchers involved in this research has further claimed, "I think umami is as important for cats as sweet is for humans".

Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F) which is similar to that of a fresh kill and routinely rejecting food presented cold or refrigerated (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing).

### Whiskers

link=https://en.wikipedia.org/wiki/File:Kittyply\_edit1.jpg|right|thumb|The whiskers of a cat are highly sensitive to touch.
To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.

Tend to fall asleep lightly for a brief period, with short periods of REM sleep with muscle twitches while they dream. Sleep between 12-16 hours, sometimes 20. Territory ranges from 17-69 acres. Low-light predators. Slightly more active at night.

### Sociability

The social behavior of the domestic cat ranges from widely dispersed individuals to feral cat colonies that gather around a food source, based on groups of co-operating females. Within such groups, one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories. These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands, and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling and, if that does not work, by short but noisy and violent attacks. Despite this colonial organization, cats do not have a social survival strategy or a pack mentality, and always hunt alone.

Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, a cat's human keeper functions as if a mother surrogate. Adult cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny. Their high-pitched sounds may mimic the cries of a hungry human infant, making them particularly difficult for humans to ignore. Some pet cats are poorly socialized. In particular, older cats show aggressiveness toward newly arrived kittens, which include biting and scratching; this type of behavior is known as feline asocial aggression.

Redirected aggression is a common form of aggression which can occur in multiple cat households. In redirected aggression there is usually something that agitates the cat: this could be a sight, sound, or another source of stimuli which causes a heightened level of anxiety or arousal. If the cat cannot attack the stimuli, it may direct anger elsewhere by attacking or directing aggression to the nearest cat, dog, human or other being.

Domestic cats' scent rubbing behavior toward humans or other cats is thought to be a feline means for social bonding.

### Communication

Main article: Cat communication
link=https://en.wikipedia.org/wiki/File:Domestic\_Cat\_Face\_Shot.jpg|thumb|Vocalizing domestic cat
Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and several different forms of meowing. Their body language, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms in cats. A raised tail indicates a friendly greeting, and flattened ears indicate hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate ones. Feral cats are generally silent. Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.

Purring may have developed as an evolutionary advantage as a signaling mechanism of reassurance between mother cats and nursing kittens, who are thought to use it as a care-soliciting signal. Post-nursing cats also often purr as a sign of contentment: when being petted, becoming relaxed, or eating. Even though purring is popularly interpreted as indicative of pleasure, it has been recorded in a wide variety of circumstances, most of which involve physical contact between the cat and another, presumably trusted individual. Some cats have been observed to purr continuously when chronically ill or in apparent pain.

The exact mechanism by which cats purr has long been elusive, but it has been proposed that purring is generated via a series of sudden build-ups and releases of pressure as the glottis is opened and closed, which causes the vocal folds to separate forcefully. The laryngeal muscles in control of the glottis are thought to be driven by a neural oscillator which generates a cycle of contraction and release every 30–40 milliseconds (giving a frequency of 33 to 25 Hz).

### Grooming

link=https://en.wikipedia.org/wiki/File:Cat\_tongue\_macro.jpg|thumb|The hooked papillae on a cat's tongue act like a hairbrush to help clean and detangle fur
Cats are known for spending considerable amounts of time licking their coats to keep them clean. The cat's tongue has backward-facing spines about 500 μm long, which are called papillae. These contain keratin which makes them rigid so the papillae act like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about 2–3 cm (3⁄4–1+1⁄4 in) long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.

### Fighting

link=https://en.wikipedia.org/wiki/File:Gato\_enervado\_pola\_presencia\_dun\_can.jpg|thumb|A domestic cat's arched back, raised fur, and open-mouthed hiss are signs of aggression.
Among domestic cats, males are more likely to fight than females. Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male. Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home. Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to sex hormones.

When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways and hissing or spitting. Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. Cats may also vocalize loudly and bare their teeth in an effort to further intimidate their opponents. Fights usually consist of grappling and delivering powerful slaps to the face and body with the forepaws as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.

Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. Fights for mating rights are typically more severe and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus. Sexually active males are usually involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to their ears and nose. Cats are willing to threaten animals larger than them to defend their territory, such as dogs and foxes.

### Hunting and feeding

See also: Cat food
link=https://en.wikipedia.org/wiki/File:Kot\_z\_mysz%C4%85.jpg|right|thumb|A domestic cat with its prey, a deermouse
The shape and structure of cats' cheeks is insufficient to allow them to take in liquids using suction. Therefore, when drinking they lap with the tongue to draw liquid upward into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upward.

Feral cats and free-fed house cats consume several small meals in a day. The frequency and size of meals varies between individuals. They select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat. Cats reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past. It is also a common misconception that cats like milk/cream, as they tend to avoid sweet food and milk. Most adult cats are lactose intolerant; the sugar in milk is not easily digested and may cause soft stools or diarrhea. Some also develop odd eating habits and like to eat or chew on things like wool, plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten.

Cats hunt small prey, primarily birds and rodents, and are often used as a form of pest control. Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. The strategy used depends on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds. Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.3 to 4.0 billion birds and 6.3 to 22.3 billion mammals annually.

Certain species appear more susceptible than others; in one English village, for example, 30% of house sparrow mortality was linked to the domestic cat. In the recovery of ringed robins (*Erithacus rubecula*) and dunnocks (*Prunella modularis*) in Britain, 31% of deaths were a result of cat predation. In parts of North America, the presence of larger carnivores such as coyotes which prey on cats and other small predators reduces the effect of predation by cats and other small predators such as opossums and raccoons on bird numbers and variety.

Perhaps the best-known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it looks like torture, is that cats often appear to "play" with prey by releasing and recapturing it. This cat and mouse behavior is due to an instinctive imperative to ensure that the prey is weak enough to be killed without endangering the cat.

Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. One explanation is that cats adopt humans into their social group and share excess kill with others in the group according to the dominance hierarchy, in which humans are reacted to as if they are at or near the top. Another explanation is that they attempt to teach their guardians to hunt or to help their human as if feeding "an elderly cat, or an inept kitten". This hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement in raising kittens.

### Play

Main article: Cat play and toys
thumb
Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey. Cats also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals.

Cats also tend to play with toys more when they are hungry. Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest. They become habituated to a toy they have played with before. String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death. Owing to the risks posed by cats eating string, it is sometimes replaced with a laser pointer's dot, which cats may chase.

Ready to mate between early February and August.

Several males, called tomcats, are attracted to a female in heat. They fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually, the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's penis has a band of about 120–150 backward-pointing penile spines, which are about 1 mm (1⁄32 in) long; upon withdrawal of the penis, the spines may provide the female with increased sexual stimulation, which acts to induce ovulation.
link=https://en.wikipedia.org/wiki/File:Radiography\_of\_a\_pregnant\_cat.jpg|left|thumb|Radiography of a pregnant cat. The skeletons of two fetuses are visible on the left and right of the uterus.
After mating, the female cleans her vulva thoroughly. If a male attempts to mate with her at this point, the female attacks him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat. Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.

The morula forms 124 hours after conception. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs. The gestation of queens lasts between 64 and 67 days, with an average of 65 days.
link=https://en.wikipedia.org/wiki/File:1dayoldkitten.JPG|thumb|A newborn kitten
Data on the reproductive capacity of more than 2,300 free-ranging queens were collected during a study between May 1998 and October 2000. They had one to six kittens per litter, with an average of three kittens. They produced a mean of 1.4 litters per year, but a maximum of three litters in a year. Of 169 kittens, 127 died before they were six months old due to a trauma caused in most cases by dog attacks and road accidents. The first litter is usually smaller than subsequent litters. Kittens are weaned between six and seven weeks of age. Queens normally reach sexual maturity at 5–10 months, and males at 5–7 months. This varies depending on breed. Kittens reach puberty at the age of 9–10 months.

Cats are ready to go to new homes at about 12 weeks of age, when they are ready to leave their mother. They can be surgically sterilized (spayed or castrated) as early as seven weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before puberty, at about three to six months. In the United States, about 80% of household cats are neutered.
The **cat** (***Felis catus***) is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the **domestic cat** or **house cat** to distinguish it from the wild members of the family. Cats are commonly kept as house pets but can also be farm cats or feral cats; the feral cat ranges freely and avoids human contact. Domestic cats are valued by humans for companionship and their ability to kill vermin. About 60 cat breeds are recognized by various cat registries.

Strong flexible body, quick reflexes, sharp teeth, retractable claws. Well-developed night vision and sense of smell. Social species but solitary hunter. Crepuscular as a predator. Secretes and perceives pheromones. Vocalizations include meowing, purring, trilling, hissing, growling, and grunting. Have 2-5 children at a time. Have children from spring to late autumn.

### Domestication

See also: Domestication of the cat
link=https://en.wikipedia.org/wiki/File:Tomb\_of\_Nakht\_(7).jpg|thumb|A cat eating a fish under a chair, a mural in an Egyptian tomb dating to the 15th century BC
The earliest known indication for the taming of an African wildcat (*F. lybica*) was excavated close by a human Neolithic grave in Shillourokambos, southern Cyprus, dating to about 7500–7200 BC. Since there is no evidence of native mammalian fauna on Cyprus, the inhabitants of this Neolithic village most likely brought the cat and other wild mammals to the island from the Middle Eastern mainland. Scientists therefore assume that African wildcats were attracted to early human settlements in the Fertile Crescent by rodents, in particular the house mouse (*Mus musculus*), and were tamed by Neolithic farmers. This mutual relationship between early farmers and tamed cats lasted thousands of years. As agricultural practices spread, so did tame and domesticated cats. Wildcats of Egypt contributed to the maternal gene pool of the domestic cat at a later time.

The earliest known evidence for the occurrence of the domestic cat in Greece dates to around 1200 BC. Greek, Phoenician, Carthaginian and Etruscan traders introduced domestic cats to southern Europe. During the Roman Empire they were introduced to Corsica and Sardinia before the beginning of the 1st millennium. By the 5th century BC, they were familiar animals around settlements in Magna Graecia and Etruria. By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a Baltic Sea port in northern Germany.

During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in the wild. Several natural behaviors and characteristics of wildcats may have pre-adapted them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play, and high intelligence. Captive *Leopardus* cats may also display affectionate behavior toward humans but were not domesticated. House cats often mate with feral cats. Hybridisation between domestic and other Felinae species is also possible, producing hybrids such as the Kellas cat in Scotland.

Development of cat breeds started in the mid 19th century. An analysis of the domestic cat genome revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific mutations were selected to develop cat breeds. Most breeds are founded on random-bred domestic cats. Genetic diversity of these breeds varies between regions, and is lowest in purebred populations, which show more than 20 deleterious genetic disorders.

### Skeleton

Cats have seven cervical vertebrae (as do most mammals); 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae (humans have five); three sacral vertebrae (as do most mammals, but humans have five); and a variable number of caudal vertebrae in the tail (humans have only three to five vestigial caudal vertebrae, fused into an internal coccyx). The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis. Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through any space into which they can fit their head.

### Skull

link=https://en.wikipedia.org/wiki/File:Cat\_skull.jpg|right|thumb|Cat skull
link=https://en.wikipedia.org/wiki/File:Cat\_yawn\_with\_exposed\_teeth\_and\_claws.jpg|thumb|A cat with exposed teeth and claws
The cat skull is unusual among mammals in having very large eye sockets and a powerful specialized jaw. Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death. Compared to other felines, domestic cats have narrowly spaced canine teeth relative to the size of their jaw, which is an adaptation to their preferred prey of small rodents, which have small vertebrae.

The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively, and cats are largely incapable of mastication. Cats tend to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar. Nonetheless they are subject to occasional tooth loss and infection.

### Claws

link=https://en.wikipedia.org/wiki/File:Shed\_domestic\_cat\_claw\_sheaths.tiff|thumb|Shed claw sheaths
Cats have protractible and retractable claws. In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows for the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hindfeet. Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading, or for extra traction on soft surfaces. Cats shed the outside layer of their claw sheaths when scratching rough surfaces.

Most cats have five claws on their front paws and four on their rear paws. The dewclaw is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws on the inside of the wrists has no function in normal walking but is thought to be an antiskidding device used while jumping. Some cat breeds are prone to having extra digits ("polydactyly"). Polydactylous cats occur along North America's northeast coast and in Great Britain.

### Ambulation

The cat is digitigrade. It walks on the toes, with the bones of the feet making up the lower part of the visible leg. Unlike most mammals, it uses a "pacing" gait and moves both legs on one side of the body before the legs on the other side. It registers directly by placing each hind paw close to the track of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for hind paws when navigating rough terrain. As it speeds up from walking to trotting, its gait changes to a "diagonal" gait: The diagonally opposite hind and fore legs move simultaneously.

They are fond of perching in high places, as a concealed site from which to hunt, and attack by pouncing from above. It also gives them a better observation point, allowing it to survey its territory.

When falling from a high place, they reflexively twist their bodies to right themselves and land on their feet, using their acute sense of balance and flexibility. They sustain very little damage from any fall higher than 2ft 11in to 9.8 ft at the bare minimum.

The cat family (Felidae) can pass down many colors and patterns to their offsprings. The domestic cat genes *MC1R* and *ASIP* allow for the variety of color in coats. The feline ASIP gene consists of three coding exons. Three novel microsatellite markers linked to ASIP were isolated from a domestic cat BAC clone containing this gene and were used to perform linkage analysis in a pedigree of 89 domestic cats that segregated for melanism.

The **tiger** (***Panthera tigris***) is the largest living cat species and a member of the genus *Panthera*. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on ungulates, such as deer and wild boar. It is territorial and generally a solitary but social predator, requiring large contiguous areas of habitat to support its requirements for prey and rearing of its offspring. Tiger cubs stay with their mother for about two years and then become independent, leaving their mother's home range to establish their own.

The tiger was first scientifically described in 1758. It once ranged widely from the Eastern Anatolia Region in the west to the Amur River basin in the east, and in the south from the foothills of the Himalayas to Bali in the Sunda Islands. Since the early 20th century, tiger populations have lost at least 93% of their historic range and have been extirpated from Western and Central Asia, the islands of Java and Bali, and in large areas of Southeast and South Asia and China. What remains of the range where tigers still roam free is fragmented, stretching in spots from Siberian temperate forests to subtropical and tropical forests on the Indian subcontinent, Indochina and a single Indonesian island, Sumatra.

The tiger is listed as Endangered on the IUCN Red List. As of 2023, the global wild tiger population is estimated to number 5,574 individuals, with most populations living in small isolated pockets . India hosts the largest tiger population. Major reasons for population decline are habitat destruction, habitat fragmentation and poaching. Tigers are also victims of human–wildlife conflict, due to encroachment in countries with a high human population density.

The tiger is among the most recognisable and popular of the world's charismatic megafauna. It featured prominently in the ancient mythology and folklore of cultures throughout its historic range and continues to be depicted in modern films and literature, appearing on many flags, coats of arms and as mascots for sporting teams. The tiger is the national animal of India, Bangladesh, Malaysia and South Korea.

## Etymology

The Middle English *tigre* and Old English *tigras* derive from Old French *tigre*, from Latin *tigris*. This was a borrowing of Classical Greek τίγρις 'tigris', a foreign borrowing of unknown origin meaning 'tiger' and the river Tigris. The origin may have been the Persian word *tigra* ('pointed or sharp') and the Avestan word *tigrhi* ('arrow'), perhaps referring to the speed of the tiger's leap, although these words are not known to have any meanings associated with tigers.

The generic name *Panthera* is derived from the Latin word *panthera* and the Ancient Greek word πάνθηρ ('panther').

## Taxonomy

In 1758, Carl Linnaeus described the tiger in his work *Systema Naturae* and gave it the scientific name *Felis tigris*. In 1929, the British taxonomist Reginald Innes Pocock subordinated the species under the genus *Panthera* using the scientific name *Panthera tigris*.

### Subspecies

link=https://en.wikipedia.org/wiki/File:Tiger\_phylogenetic\_relationships.png|left|thumb|Phylogenetic relationship of tiger populations based on Driscoll et al. (2009).
Following Linnaeus's first descriptions of the species, several tiger specimens were described and proposed as subspecies. The validity of several tiger subspecies was questioned in 1999. Most putative subspecies described in the 19th and 20th centuries were distinguished on basis of fur length and colouration, striping patterns and body size, hence characteristics that vary widely within populations. Morphologically, tigers from different regions vary little, and gene flow between populations in those regions is considered to have been possible during the Pleistocene. Therefore, it was proposed to recognize only two tiger subspecies as valid, namely *P. t. tigris* in mainland Asia, and *P. t. sondaica* in the Greater Sunda Islands.

This two-subspecies proposal was reaffirmed in 2015 by a comprehensive analysis of morphological, ecological and molecular traits of all putative tiger subspecies using a combined approach. The authors proposed recognition of only two subspecies, namely *P. t. tigris* comprising the Bengal, Malayan, Indochinese, South Chinese, Siberian and Caspian tiger populations of continental Asia, and *P. t. sondaica* comprising the Javan, Bali and Sumatran tiger populations of the Sunda Islands. The continental nominate subspecies *P. t. tigris* constitutes two clades: a northern clade composed of the Siberian and Caspian tiger populations, and a southern clade composed of all other mainland populations.

The authors of the 2015 study noted that this two-subspecies reclassification will affect tiger conservation management. It would make captive breeding programs and future re-wilding of zoo-born tigers easier, as one tiger population could then be used to bolster the population of another population. However, there is the risk that the loss of subspecies uniqueness could negatively impact protection efforts for specific populations.

In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group revised felid taxonomy in accordance with the two-subspecies proposal of the comprehensive 2015 study, and recognized the tiger populations in continental Asia as *P. t. tigris*, and those in the Sunda Islands as *P. t. sondaica*.

This two-subspecies view is still disputed by researchers, since the currently recognized nine subspecies can be distinguished genetically. Results of a 2018 whole-genome sequencing of 32 specimens support six monophyletic tiger clades corresponding with the living subspecies and indicate that the most recent common ancestor lived about 110,000 years ago.

The following tables are based on the classification of the species *Panthera tigris* provided in *Mammal Species of the World*, and also reflect the classification used by the Cat Classification Task Force in 2017:

*Panthera tigris tigris* (Linnaeus, 1758)

*Panthera tigris sondaica* (Temminck, 1844)

### Evolution

link=https://en.wikipedia.org/wiki/File:Longdan\_tiger.png|thumb|Restoration of a *Panthera zdanskyi* skull, an extinct tiger relative whose fossil remains were found in northwest China
The tiger's closest living relatives were previously thought to be the *Panthera* species lion, leopard and jaguar. Results of genetic analysis indicate that about 2.88 million years ago, the tiger and the snow leopard lineages diverged from the other *Panthera* species, and that both may be more closely related to each other than to the lion, leopard and jaguar. The geographic origin of the *Panthera* is most likely northern Central Asia. The tiger–snow leopard lineage dispersed in Southeast Asia during the Miocene.

*Panthera zdanskyi* is considered to be a sister taxon of the modern tiger. It lived at the beginning of the Pleistocene about two million years ago, its fossil remains were excavated in Gansu of northwestern China. It was smaller and more "primitive", but functionally and ecologically similar to the modern tiger. It is disputed as to whether it had the striping pattern. Northwestern China is thought to be the origin of the tiger lineage. Tigers grew in size, possibly in response to adaptive radiations of prey species like deer and bovids, which may have occurred in Southeast Asia during the Early Pleistocene.

*Panthera tigris trinilensis* lived about 1.2 million years ago and is known from fossils excavated near Trinil in Java. The Wanhsien, Ngandong, Trinil, and Japanese tigers became extinct in prehistoric times. Tigers reached India and northern Asia in the late Pleistocene, reaching eastern Beringia, Japan, and Sakhalin. Some fossil skulls are morphologically distinct from lion skulls, which could indicate tiger presence in Alaska during the last glacial period, about 100,000 years ago.

Fossil teeth and bones found in Borneo were attributed to the Bornean tiger and date to about 13,745 to 3,000 years ago. It may have accessed Borneo, when the sea level was low during a glaciation period, and may have survived until about 200 years ago. In the Ille Cave on the island of Palawan, two articulated phalanx bones were found amidst an assemblage of other animal bones and stone tools. They were smaller than mainland tiger fossils, possibly due to insular dwarfism. It has been speculated that the tiger parts were either imported from elsewhere, or that the tiger colonised Palawan from Borneo before the Holocene. Fossil remains of tigers were also excavated in Sri Lanka, China, Japan and Sarawak dating to the Late Pliocene, Pleistocene and Early Holocene.

Results of a phylogeographic study indicate that all living tigers had a common ancestor 108,000 to 72,000 years ago. The potential tiger range during the late Pleistocene and Holocene was predicted applying ecological niche modelling based on more than 500 tiger locality records combined with bioclimatic data. The resulting model shows a contiguous tiger range at the Last Glacial Maximum, indicating gene flow between tiger populations in mainland Asia. The Caspian tiger population was likely connected to the Bengal tiger population through corridors below elevations of 4,000 m (13,000 ft) in the Hindu Kush. The tiger populations on the Sunda Islands and mainland Asia were possibly separated during interglacial periods.

The tiger's full genome sequence was published in 2013. It was found to have repeat compositions much as other cat genomes and "an appreciably conserved synteny".

### Hybrids

Further information: Felid hybrid, Panthera hybrid, Liger, and Tigon

Captive tigers were bred with lions to create hybrids called liger and tigon. They share physical and behavioural qualities of both parent species. Breeding hybrids is now discouraged due to the emphasis on conservation. The liger is a cross between a male lion and a tigress. Ligers are typically between 3 and 3.5 m (10 and 12 ft) in length, and weigh between 350 and 450 kg (800 and 1,000 lb) or more. Because the lion sire passes on a growth-promoting gene, but the corresponding growth-inhibiting gene from the female tiger is absent, ligers grow far larger than either parent species.

The less common tigon is a cross between a lioness and a male tiger. Because the male tiger does not pass on a growth-promoting gene and the lioness passes on a growth inhibiting gene, tigons are around the same size as their parents. Some females are fertile and have occasionally given birth to litigons when mated to a male Asiatic lion.

## Description

https://upload.wikimedia.org/wikipedia/commons/thumb/d/d4/Siberian\_Tiger\_by\_Malene\_Th.jpg/140px-Siberian\_Tiger\_by\_Malene\_Th.jpg

Siberian tiger in Aalborg Zoo, Denmark

https://upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Bengal\_Tiger\_Skeleton.jpg/144px-Bengal\_Tiger\_Skeleton.jpg

Bengal tiger skeleton on display at the Museum of Osteology

The tiger has a muscular body with strong forelimbs, a large head and a tail that is about half the length of its body. Its pelage colouration varies between shades of orange with a white underside and distinctive vertical black stripes; the patterns of which are unique in each individual. Stripes are likely advantageous for camouflage in vegetation such as long grass with strong vertical patterns of light and shade. The tiger is one of only a few striped cat species; it is not known why spotted patterns and rosettes are the more common camouflage pattern among felids. The orange colour may also aid in camouflage as the tiger's prey are dichromats, and thus may perceive the cat as green and blended in with the vegetation.

A tiger's coat pattern is still visible when it is shaved. This is not due to skin pigmentation, but to the stubble and hair follicles embedded in the skin. It has a mane-like heavy growth of fur around the neck and jaws and long whiskers, especially in males. The pupils are circular with yellow irises. The small, rounded ears have a prominent white spot on the back, surrounded by black. These spots are thought to play an important role in intraspecific communication.

The tiger's skull is similar to a lion's skull, with the frontal region usually less depressed or flattened, and a slightly longer postorbital region. The lion skull shows broader nasal openings. Due to the variation in skull sizes of the two species, the structure of the lower jaw is a reliable indicator for their identification. The tiger has fairly stout teeth; its somewhat curved canines are the longest among living felids with a crown height of up to 90 mm (3.5 in).

### Size

There is notable sexual dimorphism between male and female tigers, with the latter being consistently smaller. The size difference between them is proportionally greater in the large tiger subspecies, with males weighing up to 1.7 times more than females. Males also have wider forepaw pads, enabling sex to be identified from tracks. It has been hypothesised that body size of different tiger populations may be correlated with climate and be explained by thermoregulation and Bergmann's rule, or by distribution and size of available prey species.

Generally, males vary in total length from 220 to 310 cm (87 to 122 in) and weigh between 90 and 300 kg (200 and 660 lb) with skull length ranging from 295 to 383 mm (11.6 to 15.1 in). The largest tiger on record reportedly weighed 423 kg (933 lb). Females vary in total length from 190 to 275 cm (75 to 108 in), weigh 65 to 167 kg (143 to 368 lb) with skull length ranging from 265 to 318 mm (10.4 to 12.5 in). In either sex, the tail represents about 0.6 to 1.1 m (2 ft 0 in to 3 ft 7 in) of the total length. The Bengal and Siberian tigers are the largest, while the Sumatran tiger is smaller and less heavy, rarely exceeding 142 kg (313 lb) in weight.

### Colour variations

link=https://en.wikipedia.org/wiki/File:Haifa\_White\_Tigers-9129-13.jpg|left|thumb|White tigers in Haifa Zoo
There are three other colour variants – white, golden and nearly stripeless snow white – that are now virtually non-existent in the wild due to the reduction of wild tiger populations, but continue in captive populations. The white tiger has white fur and sepia-brown stripes. The golden tiger has a pale golden pelage with a blond tone and reddish-brown stripes. The snow white tiger is a morph with extremely faint stripes and a pale reddish-brown ringed tail. Both snow white and golden tigers are homozygous for CORIN gene mutations.

The white tiger lacks pheomelanin (which creates the orange colour), and has dark sepia-brown stripes and blue eyes. This altered pigmentation is caused by a mutant gene that is inherited as an autosomal recessive trait, which is determined by a white locus. It is not an albino, as the dark pigments are scarcely affected. The mutation changes a single amino acid in the transporter protein SLC45A2. Both parents need to have the allele for whiteness to have white cubs. Between the early and mid 20th century, white tigers were recorded and shot in the Indian states of Odisha, Bihar, Assam and in the area of Rewa, Madhya Pradesh. The local maharaja started breeding tigers in the early 1950s and kept a white male tiger together with its normal-coloured daughter; they had white cubs. To preserve this recessive trait, only a few white individuals were used in captive breeding, which led to a high degree of inbreeding. Inbreeding depression is the main reason for many health problems of captive white tigers, including strabismus, stillbirth, deformities and premature death. Other physical defects include cleft palate and scoliosis.

The Tiger Species Survival Plan has condemned the breeding of white tigers, alleging they are of mixed ancestry and of unknown lineage. The genes responsible for white colouration are represented by 0.001% of the population. The disproportionate growth in numbers of white tigers points to inbreeding among homozygous recessive individuals. This would lead to inbreeding depression and loss of genetic variability.

There are also records of pseudo-melanic or black tigers which have thick stripes that merge. In Simlipal National Park, 37% of the tiger population has this condition, which has been linked to isolation and inbreeding.

## Distribution and habitat

link=https://en.wikipedia.org/wiki/File:Historical\_tiger\_distribution\_PLoS\_2009.png|thumb|Historical distribution of the tiger
The tiger historically ranged from eastern Turkey and Transcaucasia to the coast of the Sea of Japan, and from South Asia across Southeast Asia to the Indonesian islands of Sumatra, Java and Bali. Since the end of the last glacial period, it was probably restricted by periods of deep snow lasting longer than six months. Currently, it occurs in less than 6% of its historical range, as it has been extirpated from Southwest and Central Asia, large parts of Southeast and East Asia. It now mainly occurs in the Indian subcontinent, the Indochinese Peninsula, Sumatra and the Russian Far East, while its status in the Korean Peninsula is unknown.

The tiger is essentially associated with forest habitats. Tiger populations thrive where populations of wild cervids, bovids and suids are stable. Records in Central Asia indicate that it occurred foremost in Tugay riverine forests along the Atrek, Amu Darya, Syr Darya, Hari, Chu and Ili Rivers and their tributaries. In the Caucasus, it inhabited hilly and lowland forests. Historical records in Iran are known only from the southern coast of the Caspian Sea and adjacent Alborz Mountains. In the Amur-Ussuri region, it inhabits Korean pine and temperate broadleaf and mixed forests, where riparian forests provide food and water, and serve as dispersal corridors for both tiger and ungulates. On the Indian subcontinent, it inhabits mainly tropical and subtropical moist broadleaf forests, moist evergreen forests, tropical dry forests and the swamp forests of the Sundarbans. In the Eastern Himalayas, tigers were documented in temperate forest up to an elevation of 4,200 m (13,800 ft) in Bhutan and of 3,630 m (11,910 ft) in the Mishmi Hills. In Thailand, it lives in deciduous and evergreen forests. In Sumatra, tiger populations range from lowland peat swamp forests to rugged montane forests.

## Ecology and behaviour

### Social and daily activities

https://upload.wikimedia.org/wikipedia/commons/thumb/e/e9/Do\_Not\_Move\_I\_Saw\_You\_%2810915583%29.jpeg/285px-Do\_Not\_Move\_I\_Saw\_You\_%2810915583%29.jpeg

Tigers are comfortable in water and frequently bathe.

https://upload.wikimedia.org/wikipedia/commons/thumb/a/a2/Licking\_tigers.jpg/199px-Licking\_tigers.jpg

Tigers grooming each other

https://upload.wikimedia.org/wikipedia/commons/thumb/7/7d/Queen\_of\_Ranthambore.jpg/173px-Queen\_of\_Ranthambore.jpg

A tigress rubbing her head on a tree

https://upload.wikimedia.org/wikipedia/commons/thumb/e/eb/JenB\_Marking\_Territory.JPG/311px-JenB\_Marking\_Territory.JPG

A tiger scent-marking his territory

When not subject to human disturbance, the tiger is mainly diurnal. It does not often climb trees but cases have been recorded. It is a strong swimmer and often bathes in ponds, lakes and rivers, thus keeping cool in the heat of the day. Individuals can cross rivers up to 7 km (4.3 mi) wide and can swim up to 29 km (18 mi) in a day. During the 1980s, a tiger was observed frequently hunting prey through deep lake water in Ranthambhore National Park.

The tiger is a long-ranging species, and individuals disperse over distances of up to 650 km (400 mi) to reach tiger populations in other areas. Radio-collared tigers in Chitwan National Park started dispersing from their natal areas earliest at the age of 19 months. Four females dispersed between 0 and 43.2 km (0.0 and 26.8 mi), and 10 males between 9.5 and 65.7 km (5.9 and 40.8 mi). None of them crossed open cultivated areas that were more than 10 km (6.2 mi) wide, but moved through forested habitat.

Adult tigers lead largely solitary lives. They establish and maintain territories but have much wider home ranges within which they roam. Resident adults of either sex generally confine their movements to their home ranges, within which they satisfy their needs and those of their growing cubs. Individuals sharing the same area are aware of each other's movements and activities. The size of the home range mainly depends on prey abundance, geographic area and sex of the individual. In India, home ranges appear to be 50 to 1,000 km2 (19 to 386 sq mi) while in Manchuria, they range from 500 to 4,000 km2 (190 to 1,540 sq mi). In Nepal, defended territories are recorded to be 19 to 151 km2 (7.3 to 58.3 sq mi) for males and 10 to 51 km2 (3.9 to 19.7 sq mi) for females.

Young female tigers establish their first territories close to their mother's. The overlap between the female and her mother's territory reduces with time. Males, however, migrate further than their female counterparts and set out at a younger age to mark out their own area. A young male acquires territory either by seeking out an area devoid of other male tigers, or by living as a transient in another male's territory until he is older and strong enough to challenge the resident male. Young males seeking to establish themselves thereby comprise the highest mortality rate (30–35% per year) amongst adult tigers.

To identify his territory, the male marks trees by spraying urine, anal gland secretions, marking trails with feces and marking trees or the ground with their claws. Females also use these "scrapes", urine and fecal markings. Scent markings of this type allow an individual to pick up information on another's identity, sex and reproductive status. Females in oestrus will signal their availability by scent marking more frequently and increasing their vocalisations.

Although for the most part avoiding each other, tigers are not always territorial and relationships between individuals can be complex. An adult of either sex will sometimes share its kill with others, even with unrelated tigers. George Schaller observed a male share a kill with two females and four cubs. Unlike male lions, male tigers allow females and cubs to feed on the kill before the male is finished with it; all involved generally seem to behave amicably, in contrast to the competitive behaviour shown by a lion pride. Stephen Mills described a social feeding event in Ranthambore National Park:

> A dominant tigress they called Padmini killed a 250 kg (550 lb) male nilgai – a very large antelope. They found her at the kill just after dawn with her three 14-month-old cubs, and they watched uninterrupted for the next ten hours. During this period the family was joined by two adult females and one adult male, all offspring from Padmini's previous litters, and by two unrelated tigers, one female the other unidentified. By three o'clock there were no fewer than nine tigers round the kill.
>
>

Male tigers are generally less tolerant of other males within their territories than females are of other females. Territory disputes are usually solved by intimidation rather than outright violence. Several such incidents have been observed in which the subordinate tiger yielded by rolling onto its back and showing its belly in a submissive posture. Once dominance has been established, a male may tolerate a subordinate within his range, as long as they do not live in too close quarters. The most serious disputes tend to occur between two males competing for a female in oestrus, sometimes fighting to the death.

Captive Sumatran tiger roaring

Caged tiger growling and snarling

Facial expressions include the "defense threat", where an individual bares its teeth, flattens its ears and its pupils enlarge. Both males and females show a flehmen response, a characteristic grimace, when sniffing urine markings, but flehmen is more often associated with males detecting the markings made by tigresses in oestrus.

Tigers roar to signal their presence to other individuals over long distances. This vocalisation is forced through an open mouth as it closes and can be heard 3 km (1.9 mi) away. They may roar three or four times in a row, and other tigers may respond in kind. When tense, tigers will moan, a sound similar to a roar but softer and made when the mouth is at least partially closed. Moaning can be heard 400 m (1,300 ft) away. Aggressive encounters involve growling, snarling and hissing. During an attack, an explosive "coughing roar" or "coughing snarl" is emitted through an open mouth and exposed teeth. Chuffing—soft, low-frequency snorting similar to purring in smaller cats—is heard in more friendly situations. Other vocalisations include grunts, woofs and miaows.

### Hunting and diet

https://upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Panthera\_tigris\_-Franklin\_Park\_Zoo%2C\_Massachusetts%2C\_USA-8a\_%281%29.jpg/280px-Panthera\_tigris\_-Franklin\_Park\_Zoo%2C\_Massachusetts%2C\_USA-8a\_%281%29.jpg

An adult tiger showing incisors, canines and part of the premolars and molars

https://upload.wikimedia.org/wikipedia/commons/thumb/1/18/037tiger.jpg/204px-037tiger.jpg

Dentition of tiger above, and of Asian black bear below. The large canines are used for killing and the carnassials for tearing flesh.

Tigers mostly feed on large and medium-sized mammals, particularly ungulates weighing 60–250 kg (130–550 lb). Range-wide, the most selected prey are sambar deer, Manchurian wapiti, barasingha and wild boar. Tigers are capable of taking down larger prey like adult gaur and wild water buffalo, but opportunistically eat much smaller prey, such as monkeys, peafowl and other ground-based birds, hares, porcupines and fish. They also prey on other predators, including dogs, leopards, bears, snakes and crocodiles. Tiger attacks on adult Asian elephants and Indian rhinoceros have also been reported. More often, tigers take the more vulnerable small calves. When in close proximity to humans, tigers sometimes prey on domestic livestock like cattle, horses and donkeys. Although almost exclusively carnivorous, tigers occasionally eat vegetation for dietary fibre such as fruit of the slow match tree.

The tigers is thought to be mainly a nocturnal predator. It generally hunts alone and overpowers its prey from any angle, using its body size and strength to knock the prey off balance. Successful hunts usually require the tiger to almost simultaneously leap onto its quarry, knock it over, and grab the throat or nape with its teeth. Some tigers can reach speeds of about 49–65 km/h (30–40 mph) but only in short bursts; consequently, tigers must be close to their prey before they break cover. If the prey senses the tiger's presence before this, the tiger usually abandons the hunt rather than give chase or battle pre-alerted prey. Horizontal leaps of up to 10 m (33 ft) have been reported, although leaps of around half this distance are more typical. One in 2 to 20 hunts, including stalking near potential prey, ends in a successful kill.

https://upload.wikimedia.org/wikipedia/commons/thumb/6/61/Tiger%27s\_killing\_wild\_boar.jpg/218px-Tiger%27s\_killing\_wild\_boar.jpg

Two tigers working together to kill an Indian boar in Kanha Tiger Reserve

https://upload.wikimedia.org/wikipedia/commons/thumb/f/fe/RANTHAMBORE\_TIGER\_RESERVE.jpg/266px-RANTHAMBORE\_TIGER\_RESERVE.jpg

Bengal tiger attacking a sambar in Ranthambore Tiger Reserve

When hunting larger animals, tigers prefer to bite the throat and use their powerful forelimbs to hold onto the prey, often simultaneously wrestling it to the ground. The tiger remains latched onto the neck until its target dies of strangulation. By this method, tigers killed gaurs and water buffaloes weighing over a ton. Although they can kill healthy adults, tigers often select the calves or infirm of very large species. Healthy adult prey of this type can be dangerous to tackle, as long, strong horns, legs and tusks are all potentially fatal to the tiger. No other extant land predator routinely takes on prey this large on its own.

With small prey such as monkeys and hares, the tiger bites the nape, often breaking the spinal cord, piercing the windpipe, or severing the jugular vein or common carotid artery. Rarely, tigers have been observed to kill prey by swiping with their paws, which are powerful enough to smash the skulls of domestic cattle, and break the backs of sloth bears.

After killing their prey, tigers sometimes drag it to conceal it in vegetation, grasping with their mouths at the site of the killing bite. This, too, can require great physical strength. In one case, after it had killed an adult gaur, a tiger was observed to drag the massive carcass over a distance of 12 m (39 ft). When 13 men simultaneously tried to drag the same carcass later, they were unable to move it. An adult tiger can go for up to two weeks without eating, then gorge on 34 kg (75 lb) of flesh at one time. In captivity, adult tigers are fed 3 to 6 kg (6.6 to 13.2 lb) of meat a day.

### Enemies and competitors

link=https://en.wikipedia.org/wiki/File:Tigerdholes.jpg|left|thumb|An 1807 illustration of dholes attacking a tiger
Tigers usually prefer to eat self-killed prey, but eat carrion in times of scarcity and also steal prey from other large carnivores. Although predators typically avoid one another, if a prize is under dispute or a serious competitor is encountered, displays of aggression are common. If these fail, the conflicts may turn violent; tigers may kill or even prey on competitors such as leopards, dholes, striped hyenas, wolves, bears, pythons, and mugger crocodiles on occasion. Crocodiles, bears, and large packs of dholes may win conflicts with tigers, and crocodiles and bears can even kill them.

The considerably smaller leopard avoids competition from tigers by hunting at different times of the day and hunting different prey. In India's Nagarhole National Park, most prey selected by leopards were from 30 to 175 kg (66 to 386 lb) against a preference for heavier prey by tigers. The average prey weight in the two respective big cats in India was 37.6 kg (83 lb) against 91.5 kg (202 lb). With relatively abundant prey, tigers and leopards were seen to successfully coexist without competitive exclusion or interspecies dominance hierarchies that may be more common to the African savanna, where the leopard lives beside the lion. Golden jackals may scavenge on tiger kills. Tigers appear to inhabit the deep parts of a forest while smaller predators like leopards and dholes are pushed closer to the fringes.

### Reproduction and life cycle

Main article: Life cycle of the tiger

"Tiger cub" redirects here. For other uses, see Tiger Cub.

https://upload.wikimedia.org/wikipedia/commons/thumb/2/20/Tigress\_with\_cubs\_in\_Kanha\_Tiger\_reserve.jpg/242px-Tigress\_with\_cubs\_in\_Kanha\_Tiger\_reserve.jpg

Tiger family in Kanha Tiger Reserve

https://upload.wikimedia.org/wikipedia/commons/thumb/9/9b/Tigers\_of\_Tadoba.jpg/242px-Tigers\_of\_Tadoba.jpg

Tiger family in Tadoba Andhari Tiger Reserve

The tiger mates all year round, but most cubs are born between March and June, with a second peak in September. Gestation ranges from 93 to 114 days, with an average of 103 to 105 days. A female is only receptive for three to six days. Mating is frequent and noisy during that time. The female gives birth in a sheltered location such as in tall grass, in a dense thicket, cave or rocky crevice. The father generally takes no part in rearing. Litters consist of two or three cubs, rarely as many as six. Cubs weigh from 780 to 1,600 g (28 to 56 oz) each at birth, and are born with eyes closed. They open their eyes when they are six to 14 days old. Their milk teeth break through at the age of about two weeks. They start to eat meat at the age of eight weeks. At around this time, females usually shift them to a new den. They make short ventures with their mother, although they do not travel with her as she roams her territory until they are older. Females lactate for five to six months. Around the time they are weaned, they start to accompany their mother on territorial walks and are taught how to hunt.

A dominant cub emerges in most litters, usually a male. The dominant cub is more active than its siblings and takes the lead in their play, eventually leaving its mother and becoming independent earlier. The cubs start hunting on their own earliest at the age of 11 months, and become independent around 18 to 20 months of age. They separate from their mother at the age of two to two and a half years, but continue to grow until the age of five years. Young females reach sexual maturity at three to four years, whereas males at four to five years. Unrelated wandering male tigers often kill cubs to make the female receptive, since the tigress may give birth to another litter within five months if the cubs of the previous litter are lost. The mortality rate of tiger cubs is about 50% in the first two years. Few other predators attack tiger cubs due to the diligence and ferocity of the mother. Apart from humans and other tigers, common causes of cub mortality are starvation, freezing, and accidents. Generation length of the tiger is about eight years. The oldest recorded captive tiger lived for 26 years.

Occasionally, male tigers participate in raising cubs, usually their own, but this is extremely rare and not always well understood. In May 2015, Amur tigers were photographed by camera traps in the Sikhote-Alin Nature Reserve. The photos show a male Amur tiger pass by, followed by a female and three cubs within the span of about two minutes. In Ranthambore, a male Bengal tiger raised and defended two orphaned female cubs after their mother had died of illness. The cubs remained under his care, he supplied them with food, protected them from his rival and sister, and apparently also trained them.

## Conservation

Main article: Tiger conservation

Further information: 21st Century Tiger

In the 1990s, a new approach to tiger conservation was developed: Tiger Conservation Units (TCUs), which are blocks of habitat that have the potential to host tiger populations in 15 habitat types within five bioregions. Altogether 143 TCUs were identified and prioritized based on size and integrity of habitat, poaching pressure and population status. They range in size from 33 to 155,829 km2 (13 to 60,166 sq mi).

In 2016, an estimate of a global wild tiger population of approximately 3,890 individuals was presented during the Third Asia Ministerial Conference on Tiger Conservation. The WWF subsequently declared that the world's count of wild tigers had risen for the first time in a century.

Major threats to the tiger include habitat destruction, habitat fragmentation and poaching for fur and body parts, which have simultaneously greatly reduced tiger populations in the wild. In India, only 11% of the historical tiger habitat remains due to habitat fragmentation. Demand for tiger parts for use in traditional Chinese medicine has also been cited as a major threat to tiger populations. Some estimates suggest that there are fewer than 2,500 mature breeding individuals, with no subpopulation containing more than 250 mature breeding individuals.

India is home to the world's largest population of wild tigers. A 2014 census estimated a population of 2,226, a 30% increase since 2011. On International Tiger Day 2019, the 'Tiger Estimation Report 2018' was released by Prime Minister Narendra Modi. The report estimates a population of 2967 tigers in India with 25% increase since 2014. Modi said "India is one of the safest habitats for tigers as it has achieved the target of doubling the tiger population from 1411 in 2011 to 2967 in 2019". As of 2022, India accounts for 75 percent of global tiger population. The Tiger Census of 2023 reports tiger population in India at 3167.

In 1973, India's *Project Tiger*, started by Indira Gandhi, established numerous tiger reserves. The project was credited with tripling the number of wild Bengal tigers from some 1,200 in 1973 to over 3,500 in the 1990s, but a 2007 census showed that numbers had dropped back to about 1,400 tigers because of poaching. Following the report, the Indian government pledged $153 million to the initiative, set up measures to combat poaching, promised funds to relocate up to 200,000 villagers in order to reduce human-tiger interactions, and set up eight new tiger reserves in India. India also reintroduced tigers to the Sariska Tiger Reserve and by 2009 it was claimed that poaching had been effectively countered at Ranthambore National Park.

In the 1940s, the Siberian tiger was on the brink of extinction with only about 40 animals remaining in the wild in Russia. As a result, anti-poaching controls were put in place by the Soviet Union and a network of protected zones (zapovedniks) were instituted, leading to a rise in the population to several hundred. Poaching again became a problem in the 1990s, when the economy of Russia collapsed. The major obstacle in preserving the species is the enormous territory individual tigers require, up to 450 km (280 mi) needed by a single female and more for a single male. Current conservation efforts are led by local governments and NGO's in concert with international organisations, such as the World Wide Fund for Nature and the Wildlife Conservation Society. The competitive exclusion of wolves by tigers has been used by Russian conservationists to convince hunters to tolerate the big cats. Tigers have less impact on ungulate populations than do wolves, and are effective in controlling the latter's numbers. In 2005, there were thought to be about 360 animals in Russia, though these exhibited little genetic diversity. However, in a decade later, the Siberian tiger census was estimated from 480 to 540 individuals.

In China, tigers became the target of large-scale 'anti-pest' campaigns in the early 1950s, where suitable habitats were fragmented following deforestation and resettlement of people to rural areas, who hunted tigers and prey species. Though tiger hunting was prohibited in 1977, the population continued to decline and is considered extinct in southern China since 2001. Having earlier rejected the Western-led environmentalist movement, China changed its stance in the 1980s and became a party to the CITES treaty. By 1993 it had banned the trade in tiger parts, and this diminished the use of tiger bones in traditional Chinese medicine. The Tibetan people's trade in tiger skins has also been a threat to tigers. The pelts were used in clothing, tiger-skin *chuba* being worn as fashion. In 2006 the 14th Dalai Lama was persuaded to take up the issue. Since then there has been a change of attitude, with some Tibetans publicly burning their chubas.

In 1994, the Indonesian Sumatran Tiger Conservation Strategy addressed the potential crisis that tigers faced in Sumatra. The Sumatran Tiger Project (STP) was initiated in June 1995 in and around the Way Kambas National Park to ensure the long-term viability of wild Sumatran tigers and to accumulate data on tiger life-history characteristics vital for the management of wild populations. By August 1999, the teams of the STP had evaluated 52 sites of potential tiger habitat in Lampung Province, of which only 15 these were intact enough to contain tigers. In the framework of the STP a community-based conservation program was initiated to document the tiger-human dimension in the park to enable conservation authorities to resolve tiger-human conflicts based on a comprehensive database rather than anecdotes and opinions.

The Wildlife Conservation Society and Panthera Corporation formed the collaboration *Tigers Forever*, with field sites including the world's largest tiger reserve, the 21,756 km2 (8,400 sq mi) Hukaung Valley in Myanmar. Other reserves were in the Western Ghats in India, Thailand, Laos, Cambodia, the Russian Far East covering in total about 260,000 km2 (100,000 sq mi).

Tigers have been studied in the wild using a variety of techniques. Tiger population have been estimated using plaster casts of their pugmarks, although this method was criticized as being inaccurate. More recent techniques include the use of camera traps and studies of DNA from tiger scat, while radio-collaring has been used to track tigers in the wild. Tiger spray has been found to be just as good, or better, as a source of DNA than scat.

## Relationship with humans

### Tiger hunting

Main article: Tiger hunting
link=https://en.wikipedia.org/wiki/File:ElephantbackTigerHunt.jpg|thumb|Tiger hunting on elephant-back in India, 1808
The tiger has been one of the most sought after game animals of Asia. Tiger hunting took place on a large scale in the early 19th and 20th centuries, being a recognised and admired sport by the British in colonial India, the maharajas and aristocratic class of the erstwhile princely states of pre-independence India. A single maharaja or English hunter could claim to kill over a hundred tigers in their hunting career. Over 80,000 tigers were slaughtered in just 50 years spanning from 1875 to 1925 in British-ruled India. Tiger hunting was done by some hunters on foot; others sat up on *machans* with a goat or buffalo tied out as bait; yet others on elephant-back. King George V on his visit to Colonial India in 1911 killed 39 tigers in a matter of 10 days One of these is on display at the Royal Albert Memorial Museum.

Historically, tigers have been hunted at a large scale so their famous striped skins could be collected. The trade in tiger skins peaked in the 1960s, just before international conservation efforts took effect. By 1977, a tiger skin in an English market was considered to be worth US$4,250.

### Body part use

link=https://en.wikipedia.org/wiki/File:COLLECTIE\_TROPENMUSEUM\_Een\_groep\_mannen\_en\_kinderen\_poseert\_bij\_een\_pas\_geschoten\_tijger\_te\_Malingping\_in\_Bantam\_West-Java\_TMnr\_10006636.jpg|right|thumb|A hunting party poses with a killed Javan tiger, 1941
Tiger parts are commonly used as amulets in South and Southeast Asia. In the Philippines, the fossils in Palawan were found besides stone tools. This, besides the evidence for cuts on the bones, and the use of fire, suggests that early humans had accumulated the bones, and the condition of the tiger subfossils, dated to approximately 12,000 to 9,000 years ago, differed from other fossils in the assemblage, dated to the Upper Paleolithic. The tiger subfossils showed longitudinal fracture of the cortical bone due to weathering, which suggests that they had post-mortem been exposed to light and air. Tiger canines were found in Ambangan sites dating to the 10th to 12th centuries in Butuan, Mindanao.

Many people in China and other parts of Asia have a belief that various tiger parts have medicinal properties, including as pain killers and aphrodisiacs. There is no scientific evidence to support these beliefs. The use of tiger parts in pharmaceutical drugs in China is already banned, and the government has made some offences in connection with tiger poaching punishable by death.[*which?*] Furthermore, all trade in tiger parts is illegal under the Convention on International Trade in Endangered Species of Wild Fauna and Flora and a domestic trade ban has been in place in China since 1993.

However, the trading of tiger parts in Asia has become a major black market industry and governmental and conservation attempts to stop it have been ineffective to date. Almost all black marketers engaged in the trade are based in China and have either been shipped and sold within in their own country or into Taiwan, South Korea or Japan. The Chinese subspecies was almost completely decimated by killing for commerce due to both the parts and skin trades in the 1950s through the 1970s. Contributing to the illegal trade, there are a number of tiger farms in the country specialising in breeding them for profit. It is estimated that between 5,000 and 10,000 captive-bred, semi-tame animals live in these farms today. However, many tigers for traditional medicine black market are wild ones shot or snared by poachers and may be caught anywhere in the tiger's remaining range (from Siberia to India to the Malay Peninsula to Sumatra). In the Asian black market, a tiger penis can be worth the equivalent of around $300 U.S. dollars. In the years of 1990 through 1992, 27 million products with tiger derivatives were found. In July 2014 at an international convention on endangered species in Geneva, Switzerland, a Chinese representative admitted for the first time his government was aware trading in tiger skins was occurring in China.

### Man-eating tigers

Main article: Tiger attack

Wild tigers that have had no prior contact with humans actively avoid interactions with them. However, tigers cause more human deaths through direct attack than any other wild mammal. Attacks are occasionally provoked, as tigers lash out after being injured while they themselves are hunted. Attacks can be provoked accidentally, as when a human surprises a tiger or inadvertently comes between a mother and her young, or as in a case in rural India when a postman startled a tiger, used to seeing him on foot, by riding a bicycle. Occasionally tigers come to view people as prey. Such attacks are most common in areas where population growth, logging, and farming have put pressure on tiger habitats and reduced their wild prey. Most man-eating tigers are old, missing teeth, and unable to capture their preferred prey. For example, the Champawat Tiger, a tigress found in Nepal and then India, had two broken canines. She was responsible for an estimated 430 human deaths, the most attacks known to be perpetrated by a single wild animal, by the time she was shot in 1907 by Jim Corbett. According to Corbett, tiger attacks on humans are normally in daytime, when people are working outdoors and are not keeping watch. Early writings tend to describe man-eating tigers as cowardly because of their ambush tactics.

Man-eaters have been a particular problem in recent decades in India and Bangladesh, especially in Kumaon, Garhwal and the Sundarbans mangrove swamps of Bengal, where some healthy tigers have hunted humans. Because of rapid habitat loss attributed to climate change, tiger attacks have increased in the Sundarbans. The Sundarbans area had 129 human deaths from tigers from 1969 to 1971. In the 10 years prior to that period, about 100 attacks per year in the Sundarbans, with a high of around 430 in some years of the 1960s. Unusually, in some years in the Sundarbans, more humans are killed by tigers than vice versa. In 1972, India's production of honey and beeswax dropped by 50% when at least 29 people who gathered these materials were devoured. In 1986 in the Sundarbans, since tigers almost always attack from the rear, masks with human faces were worn on the back of the head, on the theory that tigers usually do not attack if seen by their prey. This decreased the number of attacks only temporarily. All other means to prevent attacks, such as providing more prey or using electrified human dummies, did not work as well.

### In captivity

link=https://en.wikipedia.org/wiki/File:Ringling\_Bros\_and\_Barnum\_%26\_Bailey\_Circus\_Gunther\_Gebel-Williams\_1969.jpg|right|thumb|Publicity photo of animal trainer Gunther Gebel-Williams with several of his trained tigers, c. 1969
In Ancient Roman times, tigers were kept in menageries and amphitheatres to be exhibited, trained and paraded, and were often provoked to fight gladiators and other exotic beasts. Since the 17th century, tigers, being rare and ferocious, were sought after to keep at European castles as symbols of their owners' power. Tigers became central zoo and circus exhibits in the 18th century: a tiger could cost up to 4,000 francs in France (for comparison, a professor of the Beaux-Arts at Lyons earned only 3,000 francs a year), or up to $3,500 in the United States, where a lion cost no more than $1,000.

In 2007, over 4,000 captive tigers lived in China, of which 3,000 were held by about 20 larger facilities, with the rest held by some 200 smaller facilities. In 2011, 468 facilities in the USA kept 2,884 tigers. Nineteen US states banned private ownership of tigers, fifteen require a license, and sixteen states have no regulation. Genetic ancestry of 105 captive tigers from fourteen countries and regions showed that forty-nine animals belonged distinctly to five subspecies; fifty-two animals had mixed subspecies origins. Many Siberian tigers in zoos today are actually the result of crosses with Bengal tigers.

## Cultural depictions

Tigers and their superlative qualities have been a source of fascination for mankind since ancient times, and they are routinely visible as important cultural and media motifs. They are also considered one of the charismatic megafauna, and are used as the face of conservation campaigns worldwide. In a 2004 online poll conducted by cable television channel Animal Planet, involving more than 50,000 viewers from 73 countries, the tiger was voted the world's favourite animal with 21% of the vote, narrowly beating the dog.

### Mythology and legend

Further information: Tiger in Chinese culture and Tiger in Korean culture

See also: Tiger worship
link=https://en.wikipedia.org/wiki/File:Bronze\_Tiger\_Tally\_%22Jie%22\_with\_Gold\_Inlay\_from\_Tomb\_of\_Zhao\_Mo.jpg|left|thumb|Tiger-shaped *jie* (badge of authority) with gold inlays, from the tomb of Zhao Mo
In Chinese mythology and culture, the tiger is one of the 12 animals of the Chinese zodiac. In Chinese art, the tiger is depicted as an earth symbol and equal rival of the Chinese dragon – the two representing matter and spirit respectively. The Southern Chinese martial art Hung Ga is based on the movements of the tiger and the crane. In Imperial China, a tiger was the personification of war and often represented the highest army general (or present day defense secretary), while the emperor and empress were represented by a dragon and phoenix, respectively. The White Tiger (Chinese: 白虎; pinyin: *Bái Hǔ*) is one of the Four Symbols of the Chinese constellations. It is sometimes called the White Tiger of the West (Chinese: 西方白虎), and it represents the west and the autumn season.

The tiger's tail appears in stories from countries including China and Korea, it being generally inadvisable to grasp a tiger by the tail. In Korean mythology and culture, the tiger is regarded as a guardian that drives away evil spirits and a sacred creature that brings good luck – the symbol of courage and absolute power. For the people who live in and around the forests of Korea, the tiger considered the symbol of the Mountain Spirit or King of mountain animals. So, Koreans also called the tigers "San Gun" (산군) means Mountain Lord.

In Buddhism, the tiger is one of the Three Senseless Creatures, symbolising anger, with the monkey representing greed and the deer lovesickness. The Tungusic peoples considered the Siberian tiger a near-deity and often referred to it as "Grandfather" or "Old man". The Udege and Nanai called it "Amba". The Manchu people considered the Siberian tiger as "Hu Lin", the king. In Hinduism, the god Shiva wears and sits on tiger skin. The ten-armed warrior goddess Durga rides the tigress (or lioness) Damon into battle. In southern India the god Ayyappan was associated with a tiger. Dingu-Aneni is the god in North-East India is also associated with tiger. The weretiger replaces the werewolf in shapeshifting folklore in Asia; in India they were evil sorcerers, while in Indonesia and Malaysia they were somewhat more benign. In Taiwanese folk beliefs, *Aunt Tiger* portrays the story of a tiger, which turns into an old woman, abducts children at night and devours them to satisfy her appetite. In Greco-Roman tradition, the tiger was depicted being ridden by the god Dionysus.

### Literature and media

See also: Bengal tiger § Literature
link=https://en.wikipedia.org/wiki/File:The\_Tyger\_BM\_a\_1794.jpg|right|thumb|Blake's original printing of *The Tyger*, 1794
In the Hindu epic Mahabharata, the tiger is fiercer and more ruthless than the lion. William Blake's poem in his *Songs of Experience* (1794), titled "The Tyger", portrays the tiger as a menacing and fearful animal. In Rudyard Kipling's 1894 *The Jungle Book*, the tiger Shere Khan is the mortal enemy of the human protagonist Mowgli. Yann Martel's 2001 Booker Prize winning novel *Life of Pi*, features the title character surviving shipwreck for months on a small boat with a large Bengal tiger while avoiding being eaten. The story was adapted in Ang Lee's 2012 feature film of the same name.

Friendly tiger characters include Tigger in A. A. Milne's Winnie-the-Pooh and Hobbes of the comic strip *Calvin and Hobbes*, both represented as stuffed animals come to life. Tony the Tiger is a famous mascot for Kellogg's breakfast cereal Frosted Flakes, known for his catchphrase "They're Gr-r-reat!".

### Heraldry and emblems

See also: Tigers (sports teams)

The tiger is one of the animals displayed on the Pashupati seal of the Indus Valley civilisation. The tiger was the emblem of the Chola Dynasty and was depicted on coins, seals and banners. The seals of several Chola copper coins show the tiger, the Pandyan emblem fish and the Chera emblem bow, indicating that the Cholas had achieved political supremacy over the latter two dynasties. Gold coins found in Kavilayadavalli in the Nellore district of Andhra Pradesh have motifs of the tiger, bow and some indistinct marks. The tiger symbol of Chola Empire was later adopted by the Liberation Tigers of Tamil Eelam and the tiger became a symbol of the unrecognised state of Tamil Eelam and Tamil independence movement. The Bengal tiger is the national animal of India and Bangladesh. The Malaysian tiger is the national animal of Malaysia. The Siberian tiger is the national animal of South Korea.[*citation needed*] The Tiger is featured on the logo of the Delhi Capitals IPL team.

In European heraldry, the tyger, a depiction of a tiger as imagined by European artists, is among the creatures used in charges and supporters. This creature has several notable differences from real tigers, lacking stripes and having a leonine tufted tail and a head terminating in large, pointed jaws. A more realistic tiger entered the heraldic armory through the British Empire's expansion into Asia, and is referred to as the Bengal tiger to distinguish it from its older counterpart. The Bengal tiger is not a common creature in heraldry, but is used as a supporter in the arms of Bombay and emblazoned on the shield of the University of Madras.