Neognathus

The large predator, Neognathus (new-jaw) lives 300 million years after its predecessor, Gomphiobia. Since its humble beginnings as a shark-like super predator of the sea, it has since evolved to become a capable land predator. Over the lengthy period of time separating itself from its ancestor, it has become almost unrecognizable. The creature has specialized most of its characteristics.
One such characteristic is that of its senses. The compound eye once owned by Gomphiobia has fused into a more efficient eye suited to land-based predation, and has moved to the top of the back of the head. This allows for binocular vision, a very important aspect of most predatory animals' hunting strategies. The olfactory sense has also greatly improved; the once simple siphons the creature used to gather chemical stimuli in its environment have since evolved into sensory pits covering most of the face, with small osseoplumes allowing for a greater gathering of chemicals into its leather-like pores to help it find easy prey.
Another incredible adaptation is that of its limbs. What were once bony protrusions have evolved into limbs that allow it to not only shamble upon the dirt of Wallace-II, but to run several times faster than most of its prey. The creature lost two of its limbs due to their uselessness over time, however, it has also gained the ability of manipulating its environment with its mid-limbs. They are neutral-facing limbs, so they're not quite suited to tool use yet, but they could possibly gain the ability to pronate and supinate.
Neognathus also has an adaptation that serves only as a display mechanism; a thin membranous growth creates a fan-like protrusion used to attract mates. It is brightly colored, and is likely to evolve to become more and more ornate as time further progresses this species' growth.
The habitat that this creature inhabits is that of the beaches and grasslands of Wallace-II, where prey is often abundant. Both watering holes and beachfronts are where ambush predation is most useful, so it does so with great success. It will usually run its prey down, tiring it out, and will then approach it, making sure to do so carefully. If the prey is fiesty enough, its armor plating usually protects it from attacks. Once the prey is exhausted, the Neognathus will attempt to pounce upon its prey, launching killing blows at the flanks (with its manipulatory limbs) and head/neck region (with its "jaw" and strong neck muscles). Once the prey is dispatched, it will either swallow it whole if it is of small enough size, or will tear it apart using its strong neck muscles and jaw.
The mating habits of this creature are also quite interesting. It will often search for many Wallace-months looking for a mate, as they are solitary creatures. Once the Neognathus finds the scent of a female or females, it will follow the scent until it reaches its mate. Even with the solitary nature of the species, the females are often picky about the males they choose. It is only very rarely that the male will impress the female enough to mate with her. The male will flush his fan with blood, creating brilliant patterns of blue and red. The female then expresses her thoughts about the fan with click-like sounds from her abdominal lungs, snapping out their air and sucking it back in. When successful, the two copulate, and conjoin side to side, rather than what would likely be a messy top to bottom. The female's egg cell then becomes fertilized, and she produces a calcium-based egg, from which her offspring will hatch. The male then leaves the female, likely off to find yet another mate to spread his genes. The female, however, will protect the offspring at all costs. The species' reproductive habits leave only one viable heir to their lineage at a time, and this strengthens the bond between mother and child. One is often seen with the other for most of the mother's life. She'll even protect her child from males who wish to mate with them, teaching them not to mess with mama.
This protective bond is serving as somewhat of a transition point into sapience; an increase in mutations causing the mother to not instinctively drive her offspring away has begun to cause packs to form, a matriarchical one at that. Somewhat like a herd of elephants, a single, dominant female leads a group of both females and their offspring. The males still stay solitary, but the pack structure is increasing their intelligence as well. The pack coordinates in killing prey, in ambush formation, much like wolves. They help to problem solve with each other to both find food and water, in situations that may be perilous. Soon enough, they will likely evolve from wolf/badger-like organisms into thinking beings, with thoughts, emotions, art, science, and the like. Who knows; Wallace's natural evolutionary forces are unbeknownst to them, and they could be taken down an entirely different path in evolutionary history.