Ambulospeculida

For this group of dispeculates the basal condition is as a land based creature with 8 legs.

Ambulospeculids are an ancestral group that is basally known to have been terrestrial, or at least semi-terrestrial. Ambulospeculids, due to displaying more adaptations for living on land, are one of the most diverse of the four main clades of dispeculates, adapted to virtually any environment on land, and dominating a good amount of the world, alongside all the other clades that coexist with them at some extent. Although, historically terrestrial, a large portion has turned into aquatic species, and some have even became fully aquatic and marine. Though not as relevant as celernantids, gomphiobians or panoplovitans in the water, their abundance and their importance is still something to envy. Eventually, ambulospeculids, due to their adaptability, have quickly evolved to aerial species. Flying clades of ambulospeculids have evolved independently for a large portion of the history of Wallace II, and are among the oldest fliers of this world. In Wallace II, flying ambulospeculids are spread all across the globe, and are the most abundant flying dispeculates, not too far away from the crazy amount of flying gomphiobians that evolved separately.

## Classification

* Epestredutchiia
	+ **Cetathalassa** (Cetathalassans are the earliest and most basal radiation of fully marine and aquatic ambulospeculids. They are currently the most notorious and abundant marine ambulospeculid clade, originating in the wake of the Accomodatian. However, some secondarily terrestrial species have formed, such as the basal forest-dwelling Epestrepseapotithalassa, from Northern Virtu. Some species became freshwater dwellers, such as the predatorial spinobrachi from Media Insula, and Ventuscibum, a cosmopolitan clade of freshwater and marine creatures, existing in every continent. In the seas, we have Cetesta, swimming in the seas surrounding Occidentis, and the related Maretenuem rex, from Vadum Mare. The ventuscibum relative, the highly successful predatorial and intelligent Thorakismenikefali, swimming in every marine water of the globe, displaying a cosmopolitan distribution. Cetathalassans are basally oviparous, though a large amount has developed viviparity independently, as is the case with Maretenuem, Triacrinasus, Ventuscibum and Thorakismenikefali)
	+ **Maiadorsa** (Maiadorsans are a very basal, yet very successful, radiation of ambulospeculids, that are somewhat convergent to alcyoneuvermians, in the sense they independently evolved mammal-like traits. The only continents that they are absent are on Terra Septentrionis and Virtu, which were inaccessible or too cold for them to reside. They are found in Occidentis with frequence, as evidenced by the very basal bone tree feeding lumbercow, the very tall giraffe-like Makrolaimos, the predatory Cristunguis, and the desert-dwelling Letitrabus portenderenturus and Arenicolus-nanos. A few are present in Media Insula, such as Echinatadorsum and Loricarumversor simia, both relatives of Makrolaimos. Auth is a place for dominance of other more derived maiadorsans, such as the predatory Bullet shrew, the herbivorous Cauda Apidem Bovis and the highly successful jawed maiapeltids, as well as the arboreal Ignavus gigas. Maiadorsans are oviparous)
* Bassodiavermia
	+ **Alcyoneuvermia** (Alcyoneuvermians are much more ancient and less basal ambulospeculids that, simillarly to maiadorsans, evolved a simillar mammal-like bauplan in many species. These evolved much earlier though, in the late Tenerian, while maiadorsans only arrived after, in the Accomodatian. Alcyoneuvermians, therefore, occupy a wider array of niches and geographical distribution. The basalmost is a very derived marine-dwelling worm like creature, the coral worm, which lives in the reefs of the southern archipelago of Media Insula. The terrestrial radiation that lives until today is spread on all continents. The isolated Terra Septentrionis is native to the basalmost terrestrial alcyoneuvermian, Diadematus glacies bestia. The Dpison is a bison-like animal inhabiting the open plains of Media Insula. The crinitusavemians, include the primate-like insulaculians from the forests of southern Media Insula and the highly successful crinitusavems, cursorial creatures that have spreaded in virtually any continent... they seem to have originated in Media Insula, and spreaded into the glacial lands of Terra Septentrionis, which given they can be good swimmers, its distribution into the isolated Occidentis and Auth might be the most justifiable. Duonasus adipemrostrum and Cubiculum inhabit Virtu, while the Altumbarba lives in Auth. Alcyoneuvermians are basally oviparous, though the duobarbaoideans, the native Tenebrian clade, and the Diadematus, are viviparous)
	+ Bassodioidea
		- **Bassodida** (Bassodidans are very weird and basal members of Bassodiavermia, but not as basal as alcyoneuvermians. They seem to represent a transgression between mammalian-like ambulospeculids and arthropod-like ones, or the opposite, which seems more likely given the fossil record. They are only represented by weird hairy arboreal inhabitants of Media Insula. Bassodidans are oviparous)
		- Rimavenatoria
			* **Celestipleura** (Celestipleurans are another basal group of terrestrial ambulospeculids, that have originated in the Tenerian. They are part of the typical "arthropod"-like clade of ambulospeculids, which happens to be the most varied and successful. They are found everywhere except on Meridianam. In Virtu, a good amount exist, such as the basal Dorsostomata, that weirdly walks with its back turned to the ground. In Media Insula, the highly diverse clade of insect-like Thalassopodaridae exists, as aerial, arboreal, aquatic and terrestrial species dominate, and the related millipede-like Troglodytus segmentia, meanwhile the polyspeculate-mimicking Celestiphymimus wanders, as well as the desert hopping Sabulia mobilis. In Auth, the large herbivorous millipede-like Rimatergi wanders in the forest floor. The marine and freshwater living semi-aquatic clade Uncatapleura exists in the waters of Occidentis and the nearby Archa archipelago. The predatory Megalarboranguis exists, isolated, in the largest island of the Micha archipelago. In Terra Septentrionis the diverse Otranax exist, as well as a whole subclade of celestipleurans, including the armored Hoplatarx, the tundra-dwelling Oreduorum frigus and the predatory Semborasaderia, relics of an old past where celestipleurans were much more dominant. Celestipleures are basally oviparous, though Rimatergi has independently evolved viviparous habits, and Otranax are ovoviviparous, with the female laying the eggs inside the male, and the male eventually gives birth to the babies when they hatch)
			* Silvatromouia
				+ Antecaptoria
					- **Silvoidea** (Silvoideans are basal members of the highly successful clade, known as antecaptorians, a clade notorious for its insect and crustacean like species, among others, that spreaded across all continents and islands, although leaving the other sister clade, the aercampians, to rule as major land and air rulers. Silvoideans are the basalmost and are found in Virtu, Auth, Micha and Media Insula. Media Insula has the biggest variety, with the terrestrial Silva spicatas, and the beach-dwelling Vadumspecula, which also exists in the Micha archipelago. Terebrodente, a weird, long-legged cursorial animal, exists in Auth. The primitive and large semi-aquatic swamp-dweller, the Xenotitan, is notorious for its wide distribution in Virtu, Auth and Media Insula. Silvoideans are oviparous)
					- Edurulealia
						* **Edurusconchoidea** (Edurusconchoideans are yet another basal clade, although far more successful, of antecaptorians, very well represented in the fossil record and still today. These creatures are generally armored and often tied to the water's edge, although fully terrestrial ones are not rare. They are found in every continent, except Meridianam and Terra Septentrionis. Auth is home to the basalmost, terrestrial Edurusconchia, and the more derived and terrestrial apex predatory Ictus. Occidentis is inhabited, however, by aquatic creatures, Macca being a freshwater creature, while the living fossil Patella retro densissima, a beach dweller from the southern parts of the continent, simillar to the large beach-dwelling, albeit more aquatic, Flagellum cortice. The Vadum Mare of Micha and Media Insula is home to the weird beach-dwelling Sandhopper tripercute, although terrestrial species exist in the savannas and monsoons of Media Insula, such as Dolabra Cuda, and others in the isolated islands of Micha, such as Aamai oodu. The Paddle Cattle is a marine semi-aquatic inhabitant of an isolated island in the Archa archipelago. The highly derived terrestrial spiral hopper inhabits Virtu. Edurusconchoideans are oviparous)
						* Desererepedea
							+ **Altufonsia** (Altufonsians are a small clade of antecaptorians, these are mostly composed of more terrestrial and more megafaunal antecaptorians, almost all native to Media Insula. The large desert-dwelling Desereregrallatoris, often finds the savanna-dwelling Mantimimus, while the beaches reveal the massive beach combing Gunnaluk. The southern archipelago of the continent reveals the weird Sanguivenandi, that only moves though one pair of limbs. The most successful member is the clade known as the tide springers, with close ties to the beach, are found everywhere that is coastal in Media Insula, reaching the islands of Micha, and even southern Terra Septentrionis and western Auth. Altufonsians are oviparous)
							+ Caesarsolioidea
								- **Ascenderinsecta** (Ascenderinsects are a sister clade to the much more successful and widespread xenopedans. The ascenderinsects are found in almost all continents, except Terra Septentrionis. The arboreal inhabitant of Media Insulan forests, Primas Insectum, is dwarved in geographical distribution, when compared to Ascenderspeculida, found in Occidentis, Virtu, Auth and Media Insula's forests. The highly derived Cascade sun is marine dwelling in the tropical waters of the globe, bathing Occidentis, Media Insula, Archa, Micha and Prin, completely incapable of going on land. Ascenderinsects are basally oviparous, though Primas became viviparous)
								- **Xenopeda** (Xenopedans are basically the "insects" of Wallace II, even though many others take simillar niches. However, these really take the most notorious and most important, and are found virtually everywhere that is land. In Occidentis, the basalmost member, the Antepedea, inhabit the southern portion of the continent, but these species are also found in Phael and northern Virtu. The weirdly large canid-like Canis insectomorfus is wandering in the forest floors and prairies of northern Occidentis, weirdly unlike the typical insect-like bauplan. The related Formicarex inhabits Media Insula. The Spurlites have a almost cosmopolitan distribution, only absent in Meridianam, spending most of their lives in the hide of larger creatures, feeding on their noxious and infesting parasites. The Xenoarachnians, in the other hand, simillarly have a cosmopolitan distribution, yet they are present in Terra Septentrionis and Meridianam, in the role of small spider-like bugs. The blood fly lives in the dense Media Insulan jungles. The hive-living Dracoaranea lives in the southern and northeastern archipelagos of Media Insula. Xenotetrapteryx is a weird grasshopper like creature from Meridianam, which once had relatives living in Occidentis, which are now extinct. The very weird six-winged Octoculus exists in Virtu. The rock-feasting and often flying Petra comedenti is an inhabitant of Terra Septentrionis. The treesap mosquitoe feeds on treesap in the warm climates of Media Insula. The polyspceulate symbiotic tweezer fly behaves like pollinating insects and lives in Occidentis. Xenopedans are oviparous)
				+ Aercampia
					- **Aerotheria** (Aerotherians are the most basal of aercampians. They are composed of uniquely evolving aerial ambulospeculids, and are found nearly everywhere, with the exception of Meridianam and Archa, probably due to competition with flying caedifuremians, gomphiobians and celernantids. The basalmost split, consists of the largest aerial member, the Aervenator, that lives in the regions surrounding Media Insula and Micha, reaching the southern Terra Septentrionis and western Auth. So basal this split is, that flightless species evolved, such as Circumsiliens and Xenotherium, together living in Media Insula. The other split, is much more diverse, adopting various different unique volant shapes, as evidenced by the aerial ganger, an inhabitant of the large eastern island close to Auth, and the forest-dwelling gliding Pteroagapithecini, from the forests of Auth. A more specialized group later developed wings out of feather-like filaments. These creatures never had the chance of growing too big... the Trivolans pulehelus is found in the southern colder regions of Virtu and migrates often to Occidentis, being the only aerotherian in the continent, avoiding competition from other fliers by their sheer numbers, while the more derived and related Hamodactylus and Nimbus Leo inhabit, respectively, Terra Septentrionis and Media Insula. Aerotherians are oviparous)
					- Arborpecusia
						* **Pseudocampia** (Pseudocampians are very basal and very ancient members of the main terrestrial radiation of aercampians. These creatures have historically been associated with the Occidentis continent, as evidenced by the basal Pseudoavis and Brachyphacien pecus. The jumpers are the most successful ones, with a possible Occidentis ancestry, they seem to have migrated to Terra Septentrionis, since Thron broke from Occidentis and passed very close to it, with their small size allowing them to prosper in the regions of the northern continent, but that also allowed them to easily raft into nearby landmasses, and so the jumpers managed to reach to Media Insula and prosper there, when Thron eventually crashed with Cheru, and from that continent, they even managed to reach Auth, through rafting. Pseudocampians are oviparous)
						* Caeditromouia
							+ **Caedifuremia** (Caedifuremians are a bizarre clade, divided in two splits, one a land-dwelling that had its oldest record in the Accomodatian, and a flying one, that represents among the oldest main flying clades of dispeculates, easily evolving and proliferating as early as the Tenerian, not being very affected by the nearing mass extinctons that followed after. The land-dwelling split (nupeuritinians) are found everywhere except Terra Septentrionis. The basalmost arboritines, are weird worm-like creatures, hardly with a visible relation to the rest of its relatives, living in Occidentis. The arboreal Araneaformes are a complex clade only present in the second largest island of the Archa archipelago. The fabacursorids are the most successful of all, conquering niches reminiscent of rodents and lagomorphs of Earth, spreading everywhere, except in the Archa archipelago and Terra Septentrionis. The Caedicenos is a very successful megafaunal animal from Media Insula and the largest island of the Micha archipelago. Nupeus ciqurun, a relative of caedicenos, lives in the open plains of Virtu. The flying clade is naturally an ancient one. Toxicus furem might be the most basal, from Media Insula, and its a relative of Malefirax, another inhabitant of Media Insula, that is the last surviving member of a prehistoric clade that included the extinct Tenerian and Accomodatian luminocollarians. The ambulovolant flies in the northern skies of Terra Septentrionis, and the Paper hawk in the southern ones, avoiding the polar regions of Virtu, preferring the temperate regions of that continent, as well, as Phael, southern Occidentis, southern Media Insula, Auth and Meridianam. The Moebia deepoli is part of a clade consisting one of the largest flying caedifuremians, inhabiting the coasts of Micha and Media Insula that are bathed by Vadum Mare, though other members of its family, the Cancropteridae, live in Occidentis as well. The Osverspertilio flies in Media Insula and the closely related Sporovenator flies in Micha, Media Insula and western Auth. Caedifuremians are ancestrally oviparous, though the moebiahoneians, the clade that includes Moebia, Sporovenator and Osverspertilio, is viviparous)
							+ Glossasegardia
								- **Adriaeanatia** (Adriaeanatians are largely megafaunal creatures closely tied to the water. Thanks to that they are found in every continent. The littoral segarde is found in marine and freshwater courses of Media Insula, capable of swimming on water and walking on land, and also found in the central lake Musa Sapientum Fixa, where it has long colonies of green polyspeculates growing in their long neck hairs. The related terrestrial Tree drinker is found in Media Insula as well. The weird aquatic Pitcher duck, is found in marine and freshwater aquatic environments and also able to go on land when needed, and lives in Occidentis, and its unique strategy of hunting is deserving of a whole another topic. The terrestrial Clavatherium, share a simillar niche to the maiadorsan Makrolaimos, and lives in the same continent, Occidentis, and only does not overlap its niche due to its preference in lower, softer browse instead of the higher, tougher browse of Makrolaimos. The highly aquatic, marine-dwelling, yet still walking-capable, Titanothalassus, lives in the virgin waters of Terra Septentrionis, and its ancestors seemingly reached the continent through swimming. The swamp probers live in swampy areas of Media Insula, preferring the hunting of the amphibious celernantid talon jaws. In the rivers of Virtu, the crocodilian-like Loricatorum salamandra lives there. The Cristoceragnath ruptus, from Auth, is a unique relative that convergently evolved with large predatory and armored terrestrial gomphiobians, and is a descendant of aquatic loricatorum-like creatures, and adopts sharp jaws, and an armored body. Adriaeanatians are oviparous)
								- Captioperforossumia
									* **Polyraptoria** (Polyraptorians are among the most successful clades of terrestrial ambulospeculids. These evolved and are endemic to Occidentis, Virtu and Auth, being largely absent in Media Insula, with the exception of one species, which would otherwise been a place for prosperity to the related oromollabians. Occidentis displays the wildest diversity, with basal predators like Glossa tromou, the omnivorous Osteoraptor (which is also present in Phael and Virtu), the related Polystomatatherium, the weird bipedal Unoculum nocticursor, the egg-feasting Ovicleptes and the tree browser Intusfistula longuscolii. In Virtu, the osteoraptorids inhabit. Auth displays yet another faunal composition of polyraptorians, namely the volcanic plain dwelling Terrarus, and the closely related Armagnus ossea and Culumstansbestia. In Media Insula, the only polyraptorian present is the Distratopodi reypallius, a gigantic relative of the much smaller Unoculum, that seems to have reached to the continent through island hopping from Auth, and from there, evolved to massive sizes. Polyraptorians are ancestrally oviparous, though Osteoraptorids are ovoviviparous)
									* Probosphagia
										+ **Oscanida** (Oscanids are a derived, yet reduced, group of captioperforossumians, more related to oromollabians. These creatures are only represnted by the Media Insulan Doryonyx venatos and the Occidentian Red-bellied bonedog. They have originated in the early Compsogene. Oscanids are oviparous)
										+ Captionemioidea
											- **Captionemia** (Captionemians are poorly studied Occidentian creatures, only composed by one species, Captionem, and is the basalmost captionemioid. and diverged from the mollabiaugerians as early as the mid-Variagene. They are large semi-aquatic creatures that use their legs as highly sensitive organs to detect prey and eventually trap them. Captionemians are viviparous)
											- Mollabiaugeria
												* **Soliaugeria** (Soliaugerians are basal mollabiaugerians, sister clading oromollabians. They are composed of the terrestrial Proboscusacutidae from Media Insula and Micha, the terrestrial Celeraugere from Occidentis, and the fully aquatic Sun eater, swimming in the waters surrounding Media Insula. Soliaugerians are oviparous)
												* **Oromollabia** (Oromollabians are a very successful group of ambulospeculids, mostly native to Media Insula, but they managed to spread to the regions of Auth and Virtu, and in some exceptional cases, to Terra Septentrionis, being absent altogether in the polyraptorian dominated Occidentis. The basalmost radiation is composed by mostly arthropod like creatures, with only Thorax laneus being found in Media Insula. The grazing Plain mower is found in Virtu. In Auth, the closely related and derived Arthropanthera, coexists with the cold climate inhabitant Trunked golianth. The other more successful radiation is endemic to Media Insula, with the exception of a few that reached Terra Septentrionis and Auth, but those will be talked in a bit. The basalmost rainforest-dwelling Mollab lives in the Media Insulan rainforests. An entire clade of oromollabians, the oroursans, have managed to conquer Terra Septentrionis, most likely rafting there and diversifying there when their ancestors were still very small in the late Compsogene and early Recensgene. Oroursans include the mountain chipracka, the tardibear and the mud lurker. The colorful hunter Mantiswolf, lives in the southern archipelago of the continent, as other hunters get used with the mainland, such as Sobekerius and Sauromimus, with menacing jaws. The great forest devourer, meanwhile, is a titanic herbivore, with large proboscis surrounded by skin that they use to strip plants and take down the forest, remodelling the region, in a simillar fashion to how other titans do on Earth. Although with a menacing look, Gitanonychus is a herbivore, extremelly closely related with the great forest devourer, and prefers the deserts and drylands of Media Insula. However, these can be extremelly good swimmers, when floods occur, and they have been observed floating in the sea, many miles away, and that is the main theory behind their otherwise mysterious appearance in Terra Septentrionis and Auth. Oromollabians are oviparous)

## List of Ambulospeculida

* Arafa Ultima
* Arborserpereidae
* Bestia Captiosus
* Callicorp
* Falcoptidae
* Fleaboscis?
* Lizardbug
* Mega equus
* Parasitic Stick
* Pellisvagusidae
* Plesiohippo Aquatilium
* *Renoungulus*
* Rock eater
* Sicut avis
* Trap Cricket
* Troglobite predator
* Western Clad Runner?

### **Canon**

Cetathalassa (220-0)

* Cetesta
* Epestrepseapotithalassa
* Maretenuem rex
* Spinobrachi velocivenatus
* Thorakismenikefali psaria?
* Triacrinasus dutchii
* Ventuscibum

Maiadorsa (222-0)

* Arenicolus-nanos
* Cauda apidem bovis
* Cristunguis bestia
* Echinatadorsum
* Ignavus gigas
* Letitrabus portenderenturus
* Loricarumeversor simia
* Lumbercow
* Maiapeltidae
* Makrolaimos stomacheiri
* The bullet shrew

Alcyoneuvermia (254-0)

* Altumbarba
* Coral worm
* Crinitusavem
* Cubiculum
* Diadematus glacies bestia
* Dpison
* Duonasus adipemrostrum
* Insulaculian

Bassodida (294-0)

* Bassodidae

Celestipleura (290-0)

* Celestiphymimus eruendi
* Dorsostomata
* Hoplatarx
* Megalarboranguis
* Oreduorum (genus)
* Otranax
* Rimatergi
* Sabulia mobilis
* Semborasaderia
* Thalassopódaridae
* Troglodytus segmentia
* Uncatapleura

Silvoidea (282-0)

* Silva spicatas
* Terebrodente
* Vadumspecula spp.
* Xenotitan eolania

Edurusconchoidea (275-0)

* Aamai oodu
* Dolabra cuda
* Edurusconchia insectumcaptor
* Flagellum cortice
* Ictus
* Macca
* Paddle cattle
* Patella retro densissima
* Sandhopper tripercute
* Spiral hopper

Altufonsia (187-0)

* Desereregrallatoris
* Gunnaluk
* Mantimimus savahni
* Sanguivenandi pictacaerulealis
* Tide springers

Ascenderinsecta (146-0)

* Ascenderespeculida
* Cascade Sun
* Primas insectum

Xenopeda (187-0)

* Antepedea
* Blood fly
* Canis insectomorfus
* Dracoaranea mellivora
* Equitembestia
* Formicarex oscomedentis bustosi
* Octoculus bocamanus
* Petram comedenti
* Treesap mosquitoe
* Tweezer fly
* Xenoarachnia
* Xenotetrapteryx clavocrura

Aerotheria (240-0)

* Aerial ganger
* Aervenator
* Circumsiliens vulturis
* Hamodactylus carpasinus
* Nimbus leo
* Pteroagapithecini
* Southern Shiver Bird
* Xenotherium

Pseudocampia (215-0)

* Blue-backed jumper
* Brachyphacien pecus
* Pseudoavis
* Scrub Jumper
* Tundra Jumper

Caedifuremia (282-0)

* Ambulovolant
* Araneaformes
* Arboritinae
* Caedicenos
* Fabacursor amicus
* Luminocollaris lanceae
* Moebia deepoli
* Malefirax
* Nupeus ciqurun
* Osverspertilio
* Paper hawk
* Sporovenator honei
* Toxicus furem

Adriaeanatia (199-0)

* Clavatherium
* Cristoceragnath ruptus?
* Littoral segarde
* Loricatorum salamandra
* Pelorotopsidae
* Pitcher duck
* Titanothalassus
* Tree drinker

Polyraptoria (168-0)

* Armagnus
* Culumstansbestia
* Distratopodi reypallius
* Intusfistula longuscolii
* Osteoraptor
* Ovicleptes
* Polystomatatherium
* Terrarus
* Tongue of Horror
* Unoculum nocticursor

Oscanida (100-0)

* Bonedog
* Doryonyx venatus

Captionemia (129-0)

* Captionem

Soliaugeria (75-0)

* Celeraugere
* Proboscusacutidae perforossum
* Sun eater

Oromollabia (95-0)

* Arthropanthera
* Gitanonychus
* Mantiswolf
* Mollab
* Mountain chipracka
* Plain mower
* Sauromimus
* Sobekerius occultatum
* Tardibear
* The great forest devourer
* The mud lurker
* Thorax laneus
* Trunked golianth