Azhdarchidae

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**Azhdarchidae** (from *اژدرها (Aždarha)*, the Persian word for dragon) is a family of pterosaurs known primarily from the late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the early Cretaceous as well (late Berriasian age, about 140Ma ago). Azhdarchids included some of the largest known flying animals of all time. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the azhdarchinae to include the pterosaurs *Azhdarcho*, *Quetzalcoatlus*, and "Titanopteryx" (now known as *Arambourgiania*). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil reccord, but recent studies indicate a wealth in pterosaurian faunas, including pteranodontids, nyctosaurids, tapejarids and several indeterminate forms. Some taxa like *Navajodactylus*, *Bakonydraco* and *Montanazhdarcho* were moved from Azhdarchidae to other clades.

## Description

 File:Life restoration of a group of giant azhdarchids, Quetzalcoatlus northropi, foraging on a Cretaceous fern prairie.png
Azhdarchids are characterized by their long legs and extremely long necks, made up of elongated neck vertebrae which are round in cross section. Most species of azhdarchids are still known mainly from their distinctive neck bones and not much else. The few azhdarchids that are known from reasonably good skeletons include *Zhejiangopterus* and *Quetzalcoatlus*. Azhdarchids are also distinguished by their relatively large heads and long, spear-like jaws. It had been suggested azhdarchids were skimmers, but further research has cast doubt on this idea, demonstrating that azhdarchids lacked the necessary adaptations for a skim-feeding lifestyle, and that they may have led a more terrestrial existence similar to modern storks and ground hornbills.

## Systematics

Azhdarchids were originally classified as close relatives of *Pteranodon* due to their long, toothless beaks. Others have suggested they were more closely related to the toothy Ctenochasmatids (which include filter-feeders like *Ctenochasma* and *Pterodaustro*). Currently it is widely agreed that azhdarchids were closely related to pterosaurs such as *Tupuxuara* and *Tapejara*.

### Taxonomy

Classification after Unwin 2006, except where noted.

 File:Azhdarchfeedingwittonnaish2008.png
* **Family Azhdarchidae**
	+ *Aerotitan*
	+ *Alanqa*
	+ *Aralazhdarcho*
	+ *Arambourgiania*
	+ *Azhdarcho*
	+ ?*Bakonydraco* (possible tapejarid)
	+ *Cretornis*
	+ *Eurazhdarcho*
	+ *Hatzegopteryx*
	+ ?*Montanazhdarcho* (non-azhdarchid azhdarchoid, probably a tapejarid)
	+ "*Ornithocheirus*" *bunzeli*
	+ *Palaeocursornis*
	+ *Phosphatodraco*
	+ *Quetzalcoatlus*
	+ *Volgadraco*
	+ *Zhejiangopterus*
	+ Possible valid azhdarchid genera
		- *Bennettazhia*
		- *Bogolubovia*
		- *Doratorhynchus*
		- *Eoazhdarcho*
		- *Navajodactylus*

## References

1. ↑ Dyke, G., Benton, M., Posmosanu, E. and Naish, D. (2010). "Early Cretaceous (Berriasian) birds and pterosaurs from the Cornet bauxite mine, Romania." *Palaeontology*, published online before print 15 September 2010. Template:Doi
2. ↑ Agnolin, Federico L. and Varricchio, David (2012). "Systematic reinterpretation of *Piksi barbarulna* Varricchio, 2002 from the Two Medicine Formation (Upper Cretaceous) of Western USA (Montana) as a pterosaur rather than a bird" (PDF). *Geodiversitas* **34** (4): 883–894. doi:10.5252/g2012n4a10.
3. ↑ Carroll, N. REASSIGNMENT OF MONTANAZHDARCHO MINOR AS A NON-AZHDARCHID MEMBER OF THE AZHDARCHOIDEA, SVP 2015
4. ↑ 4.0 4.1 Andres, B.; Myers, T. S. (2013). "Lone Star Pterosaurs". *Earth and Environmental Science Transactions of the Royal Society of Edinburgh*: 1. doi:10.1017/S1755691013000303.
5. ↑ Wilton, Mark P. (2013). *Pterosaurs: Natural History, Evolution, Anatomy*. Princeton University Press. ISBN 0691150613.
6. ↑ Nesov, L. A. (1984). "Upper Cretaceous pterosaurs and birds from Central Asia.". *Paleontologicheskii Zhurnal* **1984** (1): 47–57.
7. ↑ Kellner, A. W. A.; Langston, W. (1996). "Cranial remains of *Quetzalcoatlus* (Pterosauria, Azhdarchidae) from Late Cretaceous sediments of Big Bend National Park, Texas". *Journal of Vertebrate Paleontology* **16** (2): 222–231. doi:10.1080/02724634.1996.10011310.
8. ↑ Chatterjee, S.; Templin, R. J. (2004). "Posture, locomotion, and paleoecology of pterosaurs". *Geological Society of America Special Publication* **376**: 1–64. doi:10.1130/0-8137-2376-0.1.
9. ↑ Ősi, A.; Weishampel, D.B.; Jianu, C.M. (2005). "First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary". *Acta Palaeontologica Polonica* **50** (4): 777–787.
10. ↑ Humphries, S.; Bonser, R.H.C.; Witton, M.P.; Martill, D.M. (2007). "Did pterosaurs feed by skimming? Physical modelling and anatomical evaluation of an unusual feeding method" (PDF). *PLoS Biology* **5** (8): e204. doi:10.1371/journal.pbio.0050204.
11. ↑ Witton, Mark P.; Naish, Darren; McClain, Craig R. (28 May 2008). "A Reappraisal of Azhdarchid Pterosaur Functional Morphology and Paleoecology". *PLoS ONE* **3** (5): e2271. PMC 2386974. PMID 18509539. doi:10.1371/journal.pone.0002271.
12. ↑ "Pterosaurs".
13. ↑ Unwin, David M. (2006). *The Pterosaurs: From Deep Time*. New York: Pi Press. p. 273. ISBN 0-13-146308-X.
14. ↑ Ibrahim, N.; Unwin, D.M.; Martill, D.M.; Baidder, L.; Zouhri, S. (2010). Farke, Andrew Allen, ed. "A New Pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco". *PLoS ONE* **5** (5): e10875. PMC 2877115. PMID 20520782. doi:10.1371/journal.pone.0010875.
15. ↑ Averianov, A.O. (2007). "New records of azhdarchids (Pterosauria, Azhdarchidae) from the late Cretaceous of Russia, Kazakhstan, and Central Asia". *Paleontological Journal* **41** (2): 189–197. doi:10.1134/S0031030107020098.
16. ↑ 16.0 16.1 Averianov, A.O. (2010). "The osteology of *Azhdarcho lancicollis* Nessov, 1984 (Pterosauria, Azhdarchidae) from the Late Cretaceous of Uzbekistan." *Proceedings of the Zoological Institute of the Russian Academy of Sciences*, **314**(3): 246-317.
17. ↑ Vremir, M. T. S.; Kellner, A. W. A.; Naish, D.; Dyke, G. J. (2013). Viriot, Laurent, ed. "A New Azhdarchid Pterosaur from the Late Cretaceous of the Transylvanian Basin, Romania: Implications for Azhdarchid Diversity and Distribution". *PLoS ONE* **8**: e54268. doi:10.1371/journal.pone.0054268.
18. ↑ Carroll, N. REASSIGNMENT OF MONTANAZHDARCHO MINOR AS A NON-AZHDARCHID MEMBER OF THE AZHDARCHOIDEA, SVP 2015
19. ↑ Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". *Paleontological Journal* **42** (6): 634–642. doi:10.1134/S0031030108060099.

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