Archaeocoleostraca

## Taxonomy

Anthostoma

Gastrozoa

Tentaclostoma

Neostoma

Lophostoma

Diplostoma

Stenostraca

Coleostraca

**Archaeocoleostraca**

***Archaeocoleostraca*** (lit. "ancient Coleostracans") are an extinct branch of Coleostraca. They were the first macropredators to have ever existed.

## Description

Emerging around 4185ma, the *Archaeocoleostracans* were the first clade of terrestrial macropredators, feeding upon other clades of Diplostomes. *Archaeocoleostracans* measured around 50cm in length, and possessed short, barbed oral tentacles. Their gladius helped them obtain speeds greater than other Diplostomes. The evolution of the *Archaeocoleostracans* led to the emergence of the Desmostraca clade, and led Placostracans to evolve thicker armor.

### Extinction

The *Archaeocoleostracans* persisted for over 15 million years after the emergence of the Osteopods. However, they were eventually driven to extinction by the first Dromaeopods, whose greater speed made them superior hunters.

## Navigation

* **Anthostoma**
	+ Archeanthostoma
	+ Gastrozoa
		- Eugastrozoa
		- Tentaclostoma
			* Lithostraca
				+ Eulithostraca
				+ Anostraca
			* Neostoma
				+ Myzostoma
					- *Pelonomus thomi*
				+ Lophostoma
					- Diplostoma
						* Placostraca
							+ Archaeoplacostraca†
							+ Euplacostraca
								- Euplacostracidae
									* Euplacostracine
									* Echinostracinae†
								- Xerostraca
						* Stenostraca
							+ Eustenostraca
								- Desmostraca
									* Archaeodesmostraca†
										+ Archedesmostracidae†
										+ Notoformes†
											- Archenotoformes†
											- Neonotoformes†
												* Thyreostraca†
												* Campylospondyli†
													+ *Erythrerpeton lequiori*†
									* Eudesmostraca
										+ Eudesmostracidae
											- Eudesmostracinae
											- Myxobolidae
										+ Dolichostraca
											- Pronocanthidae†
												* Eupronocanthidae†
												* Phylactocanthidae†
											- Eudolichostraca
												* Eudolichostracidae
												* Nothocanthidae†
							+ Coleostraca
								- **Archaeocoleostraca**†
								- Elastospondyli
									* Archelastospondyli
										+ Archelastospondylidae
										+ Kentrodontia
											- Eukentrodontia
												* Archekentrodontia
													+ Archekentrodontidae
													+ Neokentrodontia
														- Trypophora
														- Deinoglossidae
												* Phyllophora†
											- Aspidontia†
												* *Kremarapax mckinleyi*†
									* Opisthoptera
										+ *Eopteryx emmetti*
										+ Archeopisthoptera
											- Latoptera
												* Eulatoptera
												* Pipiodontia
													+ *Corvimimus corvusamousi*
													+ Archepipiodontia
													+ Pterolophidae
											- Picoptera
												* *Gyriphagus oelespi*
										+ Neopisthoptera†
											- Sphenoptera†
												* *Velocialatus alexani*†
											- Magnoptera†
												* Eumagnoptera†
													+ *Skotalatus rauli*†
												* Dyptopterida†
													+ *Tachynectes jocomoli*†
					- Malacoformes
						* Echinostoma
						* Eumalacoformes
							+ *unspecified*
								- Ctenognatha
									* Euctenognatha
									* Necroformes
								- *unspecified*
									* Barognatha
									* Aciculognatha
										+ Euaciculognatha
										+ Chlorodermata
							+ *unspecified*
								- *unspecified*
									* Onychognatha
									* Mastigognatha
								- Isognatha
									* Myzognatha
										+ *Vitapatatoria aspiana*
									* Euisognatha
										+ Archeisognatha
										+ Cyanodermata

* **Polypodia**
	+ Tetrabrachia
		- Tachypodia
		- Acanthopodia
			* Orthorhachia
				+ Tanypterida
				+ Remipterida
					- Arheremipterida
					- Synopteridae
			* Hadrorhachia
				+ Ptilopoda†
					- *Bathytitan vendari*
				+ Euhadrorhachia
					- Diplocirridae
						* Archediplocirridae
						* Astrapophora
					- Teleopterida
						* Dorypoda
							+ *Kamakivenator cosmici*
							+ Archedorypoda
							+ Xiphopoda
						* Temnopoda†
							+ *Teratodectes iasoni*
	+ Sarcopodia
		- Coelopoda
			* Eucoelopoda
			* Trichopoda
				+ *Prionorostrum aegisari*
		- Neopoda
			* Metaxypoda†
				+ *Argyroderma piacentini*
			* Osteopoda
				+ Polyschia
					- Platydontia
						* Archeplatydontia
							+ Archeplatydontidae
								- *unspecified*
									* Archeplatydontinae
										+ *Dendronomus vasquezi*
									* Xenodontia†
										+ Brachiocephalia†
										+ Euxenodontia†
											- Streptotarsi†
											- Archexenodontia†
								- Neoplatydontia
									* Pleuroptera
										+ Archepleuroptera
											- Archepleuropteridae†
												* *Anthonomos qbelocki*†
											- Platyptera
												* *Sphenorhynchus sullivanae*
												* Euplatyptera
												* Dermatopoda
													+ Archedermatopoda
													+ Neodermatopoda
														- Rhabdopoda
															* Colobathristes borchmori
														- Compsopoda
										+ Theroptera†
											- Eutheroptera†
												* *Melanopteryx alexanderi*†
											- Chlamyptera†
												* Hypsirhynchidae†
												* Archechlamyptera†
													+ Temnorhynchidae†
														- *Charcharonyx mackrethi*†
													+ Archechlamypteridae†
									* Neoplatydontidae
										+ Scandopoda
											- *Carpophagus xeni*
											- Archescandopoda
											- Neoscandopoda
												* Pholcoscelida
												* Brachyscelida
										+ Harpactopoda†
											- Euharpactopoda†
											- Tanybrachidae†
												* *Cephalocarcinus elvari*†
							+ Copetarsi†
								- *Potamodyptes andrewi*†
						* Edaphotheria
							+ Leptopoda
								- Archeleptopoda
									* Euleptopoda†
										+ Euleptopodidae†
										+ Neoleptopoda†
											- Hoplopoda†
											- Camptopoda†
												* Eucamptopoda†
												* Sphenocerata†
									* Amblypoda
								- Thylacopoda†
									* Archethylacopoda†
										+ *Pycnothrix dicksoni*†
									* Allobrachidae†
										+ Euallobrachidae†
											- Corythobrachidae†
												* Archecorythobrachidae†
												* Camarabrachidae†
													+ *Cyanolophus mcgyveri*†
											- Archeallobrachidae†
												* Archeallobrachinae†
													+ *Melanothrix oneiropolos*†
												* Ceratobrachidae†
													+ *Thyreocephalus cryarchon*†
										+ Teleobrachidae†
											- *Therizinyx elliotti†*
							+ Eurycheridae
								- Archeurycheridae†
									* Archeurycheirinae†
										+ *Prolestes solysti*†
									* *Teratocheirus*†
										+ *Teratocheuirus philipi*†
								- Lystrocheiridae
									* Lystrocheirinae
										+ *Psammonyx deschaini*
									* Trypanocheiridae
										+ Archetrypanocheiridae†
										+ Rhamphodontia
											- Eurhamphodontia
												* Archeramphodontia
													+ Apodomorpha
													+ Archeramphodontidae
														- Archeramphodontinae
														- Coryphodontia
															* Archecoryphodontia
															* Septentrioformes
																+ Phractoformes
																+ Cryptonychidae
																	- Archecryptonychidae
																	- Neocryptonychidae
																		* Theristobrachidae
																		* Glyphonychidae
																			+ Archeglyphonychidae
																			+ Archonychidae
												* Acrocheiridae
													+ Euacrocheiridae
														- Archeacrocheiridae
														- Neoacrocheiridae
															* Heterognatha
																+ Distropheognatha
																	- Archedistropheognatha
																	- Neodistropheognatha
																		* Hadronychidae
																		* Daptodontia
																			+ Dryptognatha
																			+ Flagellifera
																+ Procaecoformes
																	- Oligopoda
																	- Euprocaecoformes
																		* Euprocaecoformidae
																		* Megantroformes
															* Panhylotheria
																+ Hylotheria
																	- Thaumatotheriidae
																	- Euhylotheria
																		* Gyrinotheria
																		* Euhylotheriidae
																			+ Euhylotheriinae
																			+ Pelecodontidae
																+ Thrinaxorhynchidae
																	- Archethrinaxorhynchidae
																	- Dictyorhynchidae
													+ Apatocheiridae†
											- Oryctocheiridae
					- Dromaeopoda
						* Archedromaepoda†
							+ *Rhomaleopus*†
							+ Archedromaeopodidae†
								- *Dromovenator setnovi*†
						* Onychodontia
							+ Euonychodontia†
								- Euonychodontidae†
									* Ensidontia†
										+ Euensidontia†
										+ Prionodontia†
									* Euonychodontinae†
										+ Archeonychodontidae†
										+ Cryptodontia†
											- *Raptonyx junae*†
											- Tachydontia†
											- Eucryptodontia†
								- Hadrodontia†
							+ Allodontia
								- Euallodontia†
									* Archallodontia†
									* Eriotherinae†
								- Thecopoda
									* Euthecopoda†
										+ *Leucothrix caliburi*†
									* Amphidontia
										+ Archeamphidontia†
										+ Xenopsidae
											- *polylophodon thomasi*
											- Archexenopsidae
												* Archexenopsinae
												* Euxenopsidae
													+ Ankistrobrachidae
													+ Neoxenopsidae
														- Teleopsidae
														- Chelaphora
															* Stenochelaphora
															* Pachychelaphora
											- Stenopsidae
												* File:StenopsidaeEvo.pngArchestenopsidae
												* Neostenopsidae
													+ Peloropsidae
														- Archepeloropsidae
														- Anaxotheriidae
													+ Ceratodontia
														- Archeceratodontia
														- Neoceratodontia
															* Aratrodontia
															* Isodontia
																+ Kophodontidae
																+ Elaphrodontia
																	- Archelaphrodontia
																	- Neoelaphrodontia
																		* Spathodontia
																		* Xyrodontia
				+ Synischia
					- Megalobrachidae†
						* Archemegalobrachia†
							+ Archemegalobrachidae†
							+ Prenobrachidae†
						* Titanopoda†
							+ Procerapoda†
								- Euprocerapoda†
									* *Megaphagus matti*†
								- Stylopoda†
									* *Hybotitan bluenoti*†
							+ Baropoda†
								- Eubaropoda†
									* *Nanopus proximensis*†
									* Eubaropodidae†
								- Pachypoda†
					- Deinognatha
						* Eudeinognatha†
							+ Eudeinognathidae†
								- *Euoplos gulielmi*†
							+ Malleognatha†
						* Odontognatha
							+ Trachygnatha†
								- *Telmarchos krisi*†
							+ Dolichognatha
								- Eudolichognatha
									* Archedolichognatha†
									* Hybognatha
								- Pelagotheria†
									* Actatheria†
										+ *Eurypus geberi*†
									* Thalattotheria†
										+ Isopterygia†
										+ Anisopterygia†
											- *Pontodrakon nicholsi*†

* **Chemophyta**
	+ Archaeophyta
	+ Xenophyta
		- Caulophyta
			* Neocaulophyta
				+ Eucaulophyta
					- Eucaulophytaceae
						* Eucaulophytoideae
						* Nothodendrales†
					- Reptophyta
				+ Brachyphyta
					- Archebrachyphyta
						* Eubrachyphyta
							+ Eubrachyphytaceae
							+ Desmophyta
								- Carnophyta†
								- Chromatophyta
									* Archechromatophyta
										+ Archechromatophytacea
										+ Aphyllophyta
									* Zygophyta
										+ Zygophytacea
											- Zygodendrales
											- Archezygophytacea
										+ Harpactophyta†
						* Tylophyta
					- Cryptophyta
						* Eucryptophyta
						* Pagophyta
			* Necrophyta
		- Xylophyta
			* Euxylophyta
				+ Archexylophyta
					- Archexylophytacaea
					- Coelophyta
				+ Nodophyta
			* Altiphyta†
				+ *Trilobophyllus axidenti*†

 † = Extinct