Darwin's Rhea

**Darwin's rhea** (*Rhea pennata*), also known as the **lesser rhea**, is a large flightless bird, but the smaller of the two extant species of rheas. It is found in the Altiplano and Patagonia in South America.

## Contents

* 1Description
* 2Etymology
* 3Taxonomy
* 4Behavior
* 5Distribution and habitat
* 6History of the discovery of the genus *Rhea*
* 7Conservation
* 8Footnotes
* 9References

## Description[edit]

A skeleton at the Cambridge University Museum of Zoology, England
Head of a Darwin's rhea at the Edinburgh Zoo

The lesser rhea stands at 90 to 100 cm (35–39 in) tall. Length is 92 to 100 cm (36–39 in) and weight is 15 to 28.6 kg (33–63 lb). Like most ratites, it has a small head and a small bill, the latter measuring 6.2 to 9.2 cm (2.4 to 3.6 in), but has long legs and a long neck. It has relatively larger wings than other ratites, enabling it to run particularly well. It can reach speeds of 60 km/h (37 mph), enabling it to outrun predators. The sharp claws on the toes are effective weapons. Their feathers are similar to those of ostriches, in that they have no aftershaft. Their plumage is spotted brown and white, and the upper part of their tarsus is feathered. The tarsus is 28 to 32 cm (11 to 13 in) long and has 18 horizontal plates on the front.

## Etymology[edit]

It is known as *ñandú petiso*, or *ñandú del norte*, in Argentina, where the majority live. Other names are *suri* and *choique*. The name *ñandú* comes from the greater rhea's name in Guaraní, *ñandú guazu*, meaning big spider, possibly in relation to their habit of alternately opening and lowering their wings when they run. In English, Darwin's rhea gets its scientific name from Rhea, a Greek goddess, and *pennata*, meaning winged. The specific name was bestowed in 1834 by Darwin's contemporary and rival Alcide d'Orbigny, who first described the bird to Europeans from a specimen from the lower Río Negro south of Buenos Aires, Argentina. As late as 2008, it was classified in the monotypic genus *Pterocnemia*. This word is formed from two Greek words *pteron*, meaning feathers, and *knēmē*, meaning the leg between the knee and the ankle, hence feather-legged, alluding to their feathers that cover the top part of the leg. In 2008, the SACC subsumed *Pterocnemia* into the genus *Rhea*.

## Taxonomy[edit]

Three subspecies have traditionally been recognized:

* *R. p. garleppi* is found in the puna of southeastern Peru, southwestern Bolivia, and northwestern Argentina.
* *R. p. tarapacensis* is found in the puna of northern Chile from the region of Arica and Parinacota to Antofagasta.
* *R. p. pennata* is found in the Patagonian steppes of Argentina and Chile.

The IUCN considers the former two northern taxa *R. p. tarapacensis* and *R. p. garleppi* as a separate species, the puna rhea (*R. tarapacensis*). Both *garleppi* and *tarapacensis* were described by Charles Chubb in 1913. It is possible *garleppi* should be considered a junior synonym of *tarapacensis*.

## Behavior[edit]

Egg, Collection Museum Wiesbaden, Germany
The lesser rhea is mainly a herbivore, with the odd small animal (lizards, beetles, grasshoppers) eaten on occasion. It predominately eats saltbush and fruits from cacti, as well as grasses. They tend to be quiet birds, except as chicks when they whistle mournfully, and as males looking for a female, when they emit a booming call.

The males of this species become aggressive once they are incubating eggs. The females thus lay the later eggs near the nest, rather than in it. Most of the eggs are moved into the nest by the male, but some remain outside, where they rot and attract flies. The male, and later the chicks, eat these flies. The incubation period is 30–44 days, and the clutch size is from 5–55 eggs. The eggs are 87 to 126 mm (3.4–5.0 in) and are greenish yellow. Chicks mature by three years of age. Outside the breeding season, Darwin's rhea is quite sociable: it lives in groups of from 5 to 30 birds, of both sexes and a variety of ages.

## Distribution and habitat[edit]

Darwin's rhea lives in areas of open scrub in the grasslands of Patagonia and on the Andean plateau (the Altiplano), through the countries of Argentina, Bolivia, Chile, and Peru. All subspecies prefer grasslands, brushlands and marshland. However, the nominate subspecies prefers elevations less than 1,500 m (4,900 ft), where the other subspecies typically range from 3,000 to 4,500 m (9,800–14,800 ft), but locally down to 1,220 m (4,000 ft) in the south.

## History of the discovery of the genus *Rhea*[edit]

Illustration of Darwin's rhea, published in 1841 in John Gould's description of birds collected on the voyage of HMS *Beagle*
During the second voyage of HMS *Beagle*, the young naturalist Charles Darwin made many trips on land, and around August 1833 heard from gauchos in the Río Negro area of Northern Patagonia about the existence of a smaller rhea, "a very rare bird which they called the Avestruz Petise". He continued searching fruitlessly for this bird, and the *Beagle* sailed south, putting in at Port Desire in southern Patagonia on 23 December. On the following day, Darwin shot a guanaco which provided them with a Christmas meal, and in the first days of January, the artist Conrad Martens shot a rhea which they enjoyed eating before Darwin realised that this was the elusive smaller rhea rather than a juvenile, and preserved the head, neck, legs, one wing, and many of the larger feathers. As with his other collections, these were sent to John Stevens Henslow in Cambridge. On 26 January, the *Beagle* entered the Straits of Magellan and at St. Gregory's Bay Darwin met Patagonians he described as "excellent practical naturalists". A half Indian, who had been born in the Northern Provinces, told him that the smaller rheas were the only species this far south, while the larger rheas kept to the north. On an expedition up the Santa Cruz River, they saw several of the smaller rheas, which were too wary to be approached closely or caught.

In 1837, Darwin's rhea was described as *Rhea darwinii* (later synomized with *R. pennata*) by the ornithologist John Gould in a presentation to the Zoological Society of London in which he was followed by Darwin reading a paper on the eggs and distribution of the two species of rheas.

When Gould classified Darwin's rhea and the greater rhea as separate species, he confirmed a serious problem for Darwin. These birds mainly live in different parts of Patagonia, but there is also an overlapping zone where the two species coexist. As every living being had been created in a fixed form, as accepted by the science of his time, they could only change their appearance by a perfect adaptation to their way of life, but would still be the same species. But now he had to deal with two different species. This started to form his idea that species were not fixed at all, but that another mechanism might be at work.

## Conservation[edit]

Darwin's rhea is categorized as least concern by the IUCN. The former southern nominate subspecies remains relatively widespread and locally fairly common. Its range is estimated at 859,000 km2 (332,000 sq mi). The situation for the two former northern subspecies is more worrying, with their combined population estimated as being possibly as low as in the hundreds. However, they are classified as *Rhea tarapacensis* by the IUCN, which regards it as being near threatened, with the primary threats being hunting, egg-collecting, and fragmentation of its habitat due to conversion to farmland or pastures for cattle-grazing.