Complex sentence: A comprehensive review in October 2012 concluded that the red wolf is a distinct species which diverged from the coyote alongside the closely related eastern wolf 150,000-300,000 years ago, though, this review has not been universally accepted among relevant authorities.
Split into simpler sentences: A comprehensive review in October 2012 concluded that the red wolf is a distinct species which diverged from the coyote alongside the closely related eastern wolf 150,000-300,000 years ago. Nevertheless, this review has not been universally accepted among relevant authorities.