Complex sentence: The earliest of these was Johannes Paulus Lotsy,, who in 1911, building on Wettstein's work, suggested four separate families, Liliaceae, Alliaceae, Agapanthaceae and Gilliesiaceae, within the "Liliifloren".
Split into simpler sentences: The earliest of these was Johannes Paulus Lotsy, (1911) building on Wettstein's work. Lotsy suggested four separate families, Liliaceae, Alliaceae, Agapanthaceae and Gilliesiaceae, within the "Liliifloren".