Complex sentence: Red based horses, such as chestnut, palomino, and cremello, may "carry" the silver dapple gene, and are capable of passing it on to their offspring, but will not express the gene in their own body color.
Split into simpler sentences: Red based horses (chestnut, palomino, cremello) may "carry" the gene, but will not express it. They can therefore pass it on to their offspring, but the silver dapple gene just won't be visible in their own body color.