Complex sentence: In both of these latter forms he had the intelligence of an animal, and was unable to access his telepathic powers, but was able to fight evenly with either Wonder Man or the Sub-Mariner.
Split into simpler sentences: In both of these latter forms he had the intelligence of an animal, and was unable to access his telepathic powers. Griffin has been able to fight evenly with either Wonder Man or the Sub-Mariner.