Complex sentence: A second edition was published by Darwin's son in 1889, without several revisions wanted by Darwin; these were not published until the third edition of 1999 (edited by Paul Ekman).
Split into simpler sentences: A second edition was published by Darwin's son in 1889. It did not contain several revisions wanted by Darwin, which were not published until the third edition of 1999 (edited by Paul Ekman).