Complex sentence: Although it was originally discovered on the island of Sri Lanka, It is predominantly found in India with studies having been conducted on "T. iridipennis" colonies located in the states of Karnataka, Kerala, and Tamil Nadu.
Split into simpler sentences: Although it was originally discovered on the island of Sri Lanka, it is predominantly found in India. Specifically, studies have been conducted on "T. iridipennis" colonies located in Karnataka, Kerala, and Tamil Nadu.