Complex sentence: As in the novel, the alpha male was killed, mistaken for a dog by a rancher, and the rest of the pack was trapped, sans the one pup that escaped, and held in Kalispell until they could be transferred to Glacier Park.
Split into simpler sentences: As in the novel, the alpha male was killed, mistaken for a dog by a rancher, and the rest of the pack was trapped, except the one pup that escaped. They were held in Kalispell until they could be transferred to Glacier Park.