Complex sentence: It forms a superspecies with Jackson's Francolin, Handsome Francolin, Chestnut-naped Francolin, and Erckel's Francolin; it is particularly closely related to the latter two species and is intermediate to them in both location and morphological features.
Split into simpler sentences: It forms a superspecies with Jackson's Francolin, Handsome Francolin, Chestnut-naped Francolin, and Erckel's Francolin. It is particularly closely related to the latter two species and is intermediate to them in both location and morphological features.