Complex sentence: While the historical Barbary lion was morphologically distinct, its genetic uniqueness remains questionable; still unclear is the taxonomic status of surviving lions frequently considered as Barbary lions, including those that originated from the collection of the King of Morocco.
Split into simpler sentences: While the historical Barbary lion was morphologically distinct, its genetic uniqueness remains questionable. Still unresolved is the taxonomic status of surviving lions frequently considered as Barbary lions, including those that originated from the collection of the King of Morocco.