Complex sentence: Savage, Jay quoted in Exhibiting sexual dimorphism, female toads were slightly larger than the males, and looked very different: instead of being bright orange, females were colored dark olive to black with scarlet spots encircled by yellow.
Split into simpler sentences: Savage, Jay quoted in Exhibiting sexual dimorphism, female toads were slightly larger than the males, and looked very different. Instead of being bright orange, females were colored dark olive to black with scarlet spots encircled by yellow.