Complex sentence: There were only four digits on the manus ('hand'), as opposed to the five found in most other ornithischians (including all other ceratopsians), while the four-toed hindfoot was very similar to many other small ornithischians.
Split into simpler sentences: There were only four digits on the manus ('hand'), as opposed to the five found in most other ornithischians (including all other ceratopsians). Overall, the four-toed hindfoot was very similar to many other small ornithischians.