Complex sentence: In general there are seldom more than three species occur in the same area, the maximum is fie in the submontane forests of Sumatra, co-occurring species are usually well-spaced on the spectrum of size, to reduce competition.
Split into simpler sentences: In general there are seldom more than three species occurring in the same area, although five species co-occur in the submontane forests of Sumatra. Co-occurring species are usually well-spaced on the spectrum of size, to reduce competition.