Complex sentence: They appear throughout the level jumping from the water to attack the player and a cluster of them make up the level's boss battle, they were replaced in the Super Nintendo version of the game by the Rat King on a Footski.
Split into simpler sentences: They appear throughout the level jumping from the water to attack the player, and a cluster of them make up the level's boss battle. They were replaced in the Super Nintendo version of the game by the Rat King on a Footski.