Complex sentence: Imitation often involves the copying of an observed behaviour of another individual, but memes may transmit from one individual to another through a copy recorded in an inanimate source, such as a book or a musical score.
Split into simpler sentences: Imitation often involves the copying of an observed behaviour of another individual. Communication may be direct or indirect, where memes transmit from one individual to another through a copy recorded in an inanimate source, such as a book or a musical score.