Complex sentence: The smaller Spanish red deer ("Cervus elaphus hispanicus ") is more gray, while the Norwegian red deer ("Cervus elaphus atlanticus") is smaller in size and paler in color.
Split into simpler sentences: The smaller Spanish red deer ("Cervus elaphus hispanicus ") is grayer. While the Norwegian red deer ("Cervus elaphus atlanticus") is smaller in size and paler in color.