Complex sentence: Iestyn Davies began singing as a boy treble in the choir of St John's College, Cambridge and later, as an undergraduate, he sang bass and then countertenor until he graduated from St John's in Archaeology and Anthropology.
Split into simpler sentences: Iestyn Davies began singing as a boy treble in the choir of St John's College, Cambridge. Later, as an undergraduate at St John's, he sang bass and then countertenor before graduating in Archaeology and Anthropology.