Complex sentence: Throughout the game Croc collects crystals which serve as health in a similar fashion to golden rings in "Sonic the Hedgehog", meaning the player will lose a life if hit without holding any crystals in their inventory.
Split into simpler sentences: Throughout the game Croc collects crystals which serve as health in a similar fashion to golden rings in "Sonic the Hedgehog". Meaning the player will lose a life if hit without holding any crystals in their inventory.