Complex sentence: Yet sharks, fig wasps, and villagers all cooperate, and it has been a vexatious problem in evolutionary studies to explain how such cooperation should evolve, let alone persist, in a world of self-maximizing egoists.
Split into simpler sentences: Yet sharks, fig wasps, and villagers all cooperate. It has been a vexatious problem in evolutionary studies to explain how such cooperation should evolve, let alone persist, in a world of self-maximizing egoists.