Complex sentence: Cysteine proteases play multi-faceted roles, virtually in every aspect of physiology and development, in plants such as in the growth and development, in senescence and apoptosis (programmed cell death), in accumulation and mobilization of storage proteins such as in seeds.
Split into simpler sentences: Cysteine proteases play multi-faceted roles, virtually in every aspect of physiology and development. In plants they are important in growth and development and in accumulation and mobilization of storage proteins such as in seeds.