Complex sentence: This removed the need for the reversal manoeuvre for Heidelberg bound trains, and Preston suburban service also ran via the new line, while Whittlesea trains continued to use the Inner Circle, albeit only from Spencer Street.
Split into simpler sentences: This removed the need for the reversal manoeuvre for Heidelberg-bound trains, and the Preston suburban service also ran via the new line. However Whittlesea trains continued to use the Inner Circle, albeit only from Spencer Street.