Complex sentence: Also, the ancient vertebrates, which made the Coconino tracks were not salamanders, differed significantly enough from salamanders that salamanders are not appropriate models in terms of behavior or tracking making processes.
Split into simpler sentences: Also, the ancient vertebrates, which made the Coconino tracks were not salamanders. They differ significantly enough from salamanders that salamanders are inappropriate models in terms of behavior for the Coconino animals or their tracking making processes.