Complex sentence: This study suggested that there is an age related dominance hierarchy based on the observation that adolescent dogs frequently submitted to their elders and young pups were submissive to the adolescents as well as the adults.
Split into simpler sentences: This study suggested that there is an age related dominance hierarchy based on the observation that adolescent dogs frequently submitted to their elders. Young pups were submissive to adolescents as well as the adults.