Complex sentence: In the Biblical tradition, the Karun is to be identified with the Pishon, one of the four rivers of Eden/Paradise, the others being the Tigris, the Euphrates and the Karkheh (the "Gihon" of the Biblical story).
Split into simpler sentences: In the Biblical tradition, Karun is to be identified with Pishon, one of the four rivers of Eden/Paradise. The others being Tigris, Euphrates and the Karkheh ("Gihon" of the Biblical story).