Complex sentence: He was elected a Fellow of the Royal Society in 1983, by whom he was awarded the Gabor Medal in 1991 for molecular biology, in 1998 the Davy Medal for chemistry and in 2008 the Royal Medal.
Split into simpler sentences: He was elected a Fellow of the Royal Society (FRS) in 1983. The Royal Society awarded him the Gabor Medal in 1991 for molecular biology, in 1998 the Davy Medal for chemistry and in 2008 the Royal Medal.