Complex sentence: Secondary sex characteristics are features that appear at sexual maturity in animals and during puberty in humans, especially the sexually dimorphic phenotypic traits that distinguish the sexes of a species, but that, unlike the sex organs, are not directly part of the reproductive system.
Split into simpler sentences: Secondary sex characteristics are features that appear during puberty in humans, and at sexual maturity in other animals. These are particularly evident in the sexually dimorphic phenotypic traits that distinguish the sexes of a species, but unlike the sex organs, are not directly part of the reproductive system.