Complex sentence: Workers appear to behaviorally have control of the reproductive output of the queen as they have been observed to police the queen by biting and harassing and even removing the queen from the nest, indicating their behavioral control of the nest.
Split into simpler sentences: Workers appear to behaviorally have control of the reproductive output of the queen. Workers have been observed to police the queen by biting and harassing and even removing the queen from the nest, indicating their behavioral control of the nest.