Complex sentence: Fingers contain some of the densest areas of nerve endings on the body, and are the richest source of tactile feedback, and have the greatest positioning capability of the body; thus the sense of touch is intimately associated with hands.
Split into simpler sentences: Fingers contain some of the densest areas of nerve endings on the body, and are the richest source of tactile feedback. They also have the greatest positioning capability of the body, thus the sense of touch is intimately associated with hands.