Complex sentence: It has important beneficial characteristics including tolerance of high mountain conditions and the grain's high protein content and lack of the saponins which complicate quinoa use, but its domestication is not complete and non-uniformity of grain ripening is a limitation.
Split into simpler sentences: It has important beneficial characteristics including: tolerance of high mountain conditions, high protein content, and a lack of the saponins which complicate quinoa use. However, its domestication is not complete and non-uniformity of grain ripening is a limitation.