Complex sentence: All the gods are significant except Neptune, who can be ignored in inland scenarios, and the gods react negatively to descrimination (for example, being given fewer temples and festivals in their honour than the other deities).
Split into simpler sentences: All the gods are significant except Neptune in inland scenarios, who can be ignored. The gods react negatively to descrimination (for example, being given fewer temples and festivals in their honour than the other deities).