Complex sentence: Wrangham identified that the primary advantage to cooperation and sociability within female primates is that within environments which contain patchy resources, female-bonded individuals are more likely to succeed in defending the limited resources than those which do not cooperate.
Split into simpler sentences: Wrangham identified that the primary advantage to cooperation and sociability within female primates is that of environments which contain patchy resources. In these conditions female-bonded individuals are more likely to succeed in defending the limited resources than those which do not cooperate.