Complex sentence: "Puddling" reflects the fact that while engaging in either feeding or mating behavior, i.e. when they are away from home, Spicebush Swallowtails tend to stay in groups.
Split into simpler sentences: Puddling reflects the fact that while engaging in either feeding or mating behavior. When they are away from home, Spicebush Swallowtails tend to stay in groups.