Complex sentence: During colony development, the foundress lays primarily all of the eggs within the colony and performs most of the abdominal wagging, both acts usually reserved for the queen, and performed by the highest ranking colony members.
Split into simpler sentences: During colony development, the foundress primarily lays all of the eggs within the colony and performs most of the abdominal wagging. Both acts are usually reserved for the queen and performed by the highest ranking colony members.