Complex sentence: He conducted studies at marine biology stations in Naples, Villefranche-sur-Mer, Helgoland and Monaco, subsequently earning his doctorate in 1912 with a dissertation on the vascular system of Octopoda.
Split into simpler sentences: He conducted studies at marine biology stations in Naples, Villefranche-sur-Mer, Helgoland and Monaco. In 1912 he obtained his doctorate with a dissertation on the vascular system of Octopoda.