Complex sentence: The great majority of these units were light infantry battalions and independent companies designated "caçadores", which were deployed in grid along the theatre of operations, with each one being responsible for an area of responsibility.
Split into simpler sentences: The great majority of these units were light infantry battalions and independent companies designated "caçadores". They were deployed in grid ("quadrilha") along the theatre of operations, with each one being responsible for a given area of responsibility.