Complex sentence: The competitive exclusion of wolves by tigers has been used by Russian conservationists to convince hunters in the Far East to tolerate the big cats, as they limit ungulate populations less than wolves, and are effective in controlling the latter's numbers.
Split into simpler sentences: The competitive exclusion of wolves by tigers has been used by Russian conservationists to convince hunters to tolerate the big cats. Tigers have less impact on ungulate populations than do wolves, and are effective in controlling the latter's numbers.