Complex sentence: Mark Richardson joined Skunk Anansie to replace their original drummer, and when the band folded, he joined Feeder following the loss of their drummer Jon Lee and is now an established member of the band.
Split into simpler sentences: Mark Richardson joined Skunk Anansie to replace their original drummer. When that band folded, he joined Feeder, following the loss of their original drummer Jon Lee, and is now an established member of the band.