Complex sentence: There are two species of chamois in the genus "Rupicapra": "R. rupicapra" (the type species) is replaced in the Pyrenees by the Pyrenean chamois, "R. pyrenaica".
Split into simpler sentences: There are two species of chamois in the genus "Rupicapra". In the Pyrenees the type species, "R. rupicapra" is replaced by the Pyrenean chamois, "R. pyrenaica".