Complex sentence: Espio has the ability to walk upright on walls and ceilings, and instead of curling up into a ball when performing the spin dash, he twists into a tornado, and can destroy unguarded enemies by running headlong into them at full speed.
Split into simpler sentences: Espio has the ability to walk upright on walls and ceilings. He twists into a tornado (instead of curling up into a ball when performing the spin dash), and can destroy unguarded enemies by running headlong into them at full speed.