Complex sentence: At the end of 2008, Clarke published a reconstruction of the circumstances which allowed the fossil to remain so unusually well preserved in contrast to other bones found in the same cave, which apparently had been washed over longer periods of time in their final storage location.
Split into simpler sentences: At the end of 2008, Clarke published a reconstruction of the circumstances due to which the fossil could remain so unusually well preserved. In contrast to the other bones found in the same cave, which apparently had been washed over longer periods of time on their final storage location.