Complex sentence: Dr. Harlow began his career with nonhuman primate research working with the primates at the Henry Vilas Zoo, where he developed the Wisconsin General Testing Apparatus (WGTA) to study learning, cognition, and memory.
Split into simpler sentences: Dr. Harlow began his career with nonhuman primate research. He worked with the primates at Henry Vilas Zoo, where he developed the Wisconsin General Testing Apparatus (WGTA) to study learning, cognition, and memory.