Complex sentence: Science writer Matt Ridley popularized the term in connection with sexual selection in his 1993 book "The Red Queen", in which he discussed the debate in theoretical biology over the adaptive benefit of sexual reproduction to those species in which it appears.
Split into simpler sentences: Science writer Matt Ridley popularized the term in connection with sexual selection in his 1993 book "The Red Queen". In the book, Ridley discussed the debate in theoretical biology over the adaptive benefit of sexual reproduction to those species in which it appears.