Complex sentence: Studies in "Halothiobacillus neapolitanus" have shown that empty shells of normal shape and composition are assembled in carboxysomal RubisCO-lacking mutants suggesting that carboxysome shell biogenesis and enzyme sequestration are two independent, but functionally linked processes.
Split into simpler sentences: Studies in "Halothiobacillus neapolitanus" have shown that empty shells of normal shape and composition are assembled in carboxysomal RubisCO-lacking mutants. Thus, carboxysome shell biogenesis and enzyme sequestration appear to be two independent, but functionally linked processes.