Complex sentence: As a Vespidae wasp, "Polistes exclamans" nests are typically made up of paper with a single layer of cells with open combs; although nest size varies, the upper limit is around 500 cells.
Split into simpler sentences: As a Vespidae wasp, "Polistes exclamans" nests are typically made up of paper with a single layer of cells with open combs. Although nest size varies, the upper limit is around 500 cells.