Complex sentence: The other issue that marked his career was the Funder vs. Readjuster debate, which culminated in the end of the Conservative Party in Virginia and the formation of the Readjuster Party, with William Mahone as its boss, and the Virginia Democratic Party.
Split into simpler sentences: The other issue that marked his career was the Funder vs. Readjuster debate. The controversy culminated in the end of the Conservative Party in Virginia and the formation of the Readjuster Party, with William Mahone as its boss, and the Virginia Democratic Party.