Complex sentence: Major contributions to the development of coalescent theory have been made by Peter Donnelly, Robert Griffiths, Richard R Hudson and Simon Tavaré, this has included incorporating variations in population size recombination and selection.
Split into simpler sentences: Major contributions to the development of coalescent theory have been made by Peter Donnelly, Robert Griffiths, Richard R Hudson and Simon Tavaré. This has included incorporating variations in population size, recombination and selection.