Complex sentence: The validity and circumscription of proposed subfamilies like Garrinae, Labeoninae, or Squaliobarbinae also remains doubtful, although the latter do appear to correspond to a distinct lineage.
Split into simpler sentences: The validity and circumscription of proposed subfamilies like Garrinae, Labeoninae or Squaliobarbinae also remains doubtful. The latter however appear to correspond to a distinct lineage.