Complex sentence: When an adult is being carried by another or colonies are migrating, it takes on a pupal position until the ant is let go; both queens and workers may be dragged by others and males are seen riding on queens without trying to copulate.
Split into simpler sentences: When an adult is being carried by another or colonies are migrating, it takes on a pupal position until the ant is let go. Both queens and workers may be dragged by others and males are seen riding on queens without trying to copulate.