Complex sentence: The largely marine brown and Peruvian pelicans, formerly considered conspecific, are sometimes separated from the others by placement in the subgenus "Leptopelicanus" but in fact species with both sorts of appearance and nesting behavior are found in either.
Split into simpler sentences: The largely marine brown and Peruvian pelicans, formerly considered conspecific, are sometimes separated from the others by placement in the subgenus "Leptopelicanus". However, the species with both sorts of appearance and nesting behavior are found in either.