Complex sentence: While nests are all within the territory of the colony, they are not tightly packed like other colony breeders so nests are generally within calling range of neighbor nests, but each nest is within the nesting pairs normal foraging range.
Split into simpler sentences: While nests are all within the territory of the colony, they are not tightly packed like other colony breeders. Nests are generally within calling range of neighbor nests, but each nest is within the nesting pair's normal foraging range.