Complex sentence: The Ordovician marks the last great diversification period amongst the trilobites, very few entirely new patterns of organisation arose post-Ordovician; later evolution in trilobites was largely a matter of variations upon the Ordovician themes.
Split into simpler sentences: The Ordovician marks the last great diversification period amongst the trilobites: very few entirely new patterns of organisation arose post-Ordovician. Later evolution in trilobites was largely a matter of variations upon the Ordovician themes.