Complex sentence: However, Orlove noted there is a second derivation of a formula for r which on its own is similarly ad hoc, but the two taken together complement each other, eliminating the ad hoc nature of the formula, and of inclusive fitness theory as well.
Split into simpler sentences: However, Orlove noted there is a second derivation of a formula for r which on its own is similarly ad hoc. Orlove argues that the existence of two independent derivations for r reduces or eliminates the ad hoc nature of the formula, and of inclusive fitness theory as well.