{{위키데이터 속성 추적}}
{{생물 분류
|이름 = 나노호염고세균
|그림 = 
|그림_설명 = 
|색 = 고균
|계 = [[고세균|고세균계]]
|상문 = [[DPANN군]]
|문 = '''나노호염고세균문'''<br />("Nanohaloarchaeota")
|강 = '''나노호염고세균강'''<br />("Nanohaloarchaea")<br /><small>(sic) Narasingarao et al. 2012</small>
|목 = '''나노호염고세균목'''<br />("Nanohalarchaeales")
|과 = '''나노호염고세균과'''<br />("Nanohalarchaeaceae")
|속 = "Candidatus ''Nanohalarchaeum''"<br /><small>corrig. Hamm et al. 2019</small>
|이명 = "Nanohalarchaeia"<br /><small>corrig. Narasingarao et al. 2012</small>
}}
'''나노호염고세균'''(Nanohaloarchaea)은 작은 [[유전체 (생명과학)|게놈]]과 제한된 대사 능력을 가지는 [[고세균]]의 일부 분기군으로 [[DPANN군|DPANN]] 고세균에 속한다. 이 고세균은 고농도 염분의 서식지에서 극호염성인 [[호염고세균]]과 함께 널리 존재한다. 나노호염고세균은 메타게놈 데이터에서 6개 분기군으로 구성된 미배양 호염성 고세균의 한 부류로 처음으로 확인되어,<ref>{{저널 인용|제목=De novo metagenomic assembly reveals abundant novel major lineage of Archaea in hypersaline microbial communities|저널=ISME J.|성=Narasingarao|날짜=2011|권=6|호=1|쪽=81–93|doi=10.1038/ismej.2011.78|pmc=3246234|pmid=21716304|저자표시=etal}}</ref><ref name="salternRef">{{저널 인용|제목=New Abundant Microbial Groups in Aquatic Hypersaline Environments|저널=Scientific Reports|성=Ghai|이름=Rohit|성2=Lejla Pašić|날짜=10 October 2011|권=1|쪽=135|bibcode=2011NatSR...1E.135G|doi=10.1038/srep00135|pmc=3216616|pmid=22355652|저자표시=4|성3=Ana Beatriz Fernandez|성4=Ana-Belen Martin-Cuadrado|성5=Carolina Megumi Mizuno|성6=Katherine D. McMahon|성7=R. Thane Papke|성8=Ramunas Stepanauskas|성9=Beltran Rodriguez-Brito}}</ref> 곧 [[DPANN군]] 아래의 ''''나노호염고세균 문'''<nowiki/>'(Nanohaloarchaeota) 으로 배치되었다.<ref>{{저널 인용|제목=Insights into the phylogeny and coding potential of microbial dark matter|저널=Nature|성=Rinke C, Schwientek P, Sczyrba A, Ivanova NN, Anderson IJ, Cheng JF, Darling A, Malfatti S, Swan BK, Gies EA, Dodsworth JA, Hedlund BP, Tsiamis G, Sievert SM, Liu WT, Eisen JA, Hallam SJ, Kyrpides NC, Stepanauskas R, Rubin EM, Hugenholtz P, Woyke T|날짜=July 25, 2013|권=499, 431–437 (2013)|호=7459|쪽=431–437|bibcode=2013Natur.499..431R|doi=10.1038/nature12352|pmid=23851394}}</ref> 그러나 나노호염고세균의 [[계통발생학]]적 위치는 여전히 논쟁의 여지가 많아서, '''[[호염고세균]]'''의 자매 계통 또는 '''DPANN 상문'''의 구성원으로 제안되기도 한다.<ref name="petitjean">{{저널 인용|제목=Rooting the domain Archaea by phylogenomic analysis supports the foundation of the new kingdom Proteoarchaeota|저널=Genome Biol. Evol.|성=Petitjean, C.|성2=Deschamps, P.|연도=2014|권=7|호=1|쪽=191–204|doi=10.1093/gbe/evu274|pmc=4316627|pmid=25527841|성3=López-García, P.|성4=Moreira, D.}}</ref><ref name="cavalier-smith">{{저널 인용|제목=Multidomain ribosomal protein trees and the planctobacterial origin of neomura (Eukaryotes, archaebacteria)|저널=Protoplasma|성=Cavalier-Smith|이름=Thomas|성2=Chao|이름2=Ema E-Yung|url=|연도=2020|권=257|호=3|쪽=621–753|doi=10.1007/s00709-019-01442-7|pmc=7203096|pmid=31900730}}</ref><ref>{{저널 인용|제목=Extreme halophilic archaea derive from two distinct methanogen Class II lineages|저널=Molecular Phylogenetics and Evolution|성=Monique Aouad, Najwa Taïb, Anne Oudart, Michel Lecocq, Manolo Gouy,Céline Brochier-Armanet|url=https://hal.archives-ouvertes.fr/hal-01799910/file/aouad2018.pdf|날짜=21 Apr 2018|권=2018|호=27|출판사=Elsevier|쪽=46–54|doi=10.1016/j.ympev.2018.04.011|pmid=29684598}}</ref>

이후에도 오스트레일리아의 탈라소할린 호수,<ref>{{저널 인용|제목=Metagenomic and lipid analyses reveal a diel cycle in a hypersaline microbial ecosystem|저널=ISME J|성=Karen Andrade, Jörn Logemann, Karla B Heidelberg, Joanne B Emerson, Luis R Comolli, Laura A Hug, Alexander J Probst, Angus Keillar, Brian C Thomas, Christopher S Miller, Eric E Allen, John W Moreau, Jochen J Brocks & Jillian F Banfield|날짜=28 April 2015|권=ISME J 9, 2697–2711 (2015)|호=12|쪽=2697–2711|doi=10.1038/ismej.2015.66|pmc=4817636|pmid=25918833}}</ref> 스페인 염전,<ref>{{저널 인용|제목=New Abundant Microbial Groups in Aquatic Hypersaline Environments|저널=Scientific Reports|성=Rohit Ghai, Lejla Pašić, Ana Beatriz Fernández, Ana-Belen Martin-Cuadrado, Carolina Megumi Mizuno, Katherine D. McMahon, R. Thane Papke, Ramunas Stepanauskas, Beltran Rodriguez-Brito, Forest Rohwer, Cristina Sánchez-Porro, Antonio Ventosa & Francisco Rodríguez-Valera|날짜=31 October 2011|권=1|호=Sci Rep 1, 135 (2011)|쪽=135|bibcode=2011NatSR...1E.135G|doi=10.1038/srep00135|pmc=3216616|pmid=22355652}}</ref> 러시아 소다 염수,<ref>{{저널 인용|제목=Metagenomic Insights into the Uncultured Diversity and Physiology of Microbes in Four Hypersaline Soda Lake Brines|저널=Frontiers in Microbiology|성=Vavourakis Charlotte D., Ghai Rohit, Rodriguez-Valera Francisco, Sorokin Dimitry Y., Tringe Susannah G., Hugenholtz Philip, Muyzer Gerard|날짜=25 February 2016|권=7|쪽=211|doi=10.3389/fmicb.2016.00211|pmc=4766312|pmid=26941731}}</ref> 캘리포니아 염전,<ref>{{저널 인용|제목=Cell sorting analysis of geographically separated hypersaline environments|저널=Extremophiles|성=Olga Zhaxybayeva 1, Ramunas Stepanauskas, Nikhil Ram Mohan, R Thane Papke|url=https://www.researchgate.net/publication/235380187|날짜=29 January 2013|권=17,2 (2013)|호=2|쪽=265–275|doi=10.1007/s00792-013-0514-z|pmid=23358730|확인날짜=2 March 2021}}</ref> 및 칠레 암염<ref>{{저널 인용|제목=Functional interactions of archaea, bacteria and viruses in a hypersaline endolithic community|저널=Environmental Microbiology|성=Alexander Crits‐Christoph Diego R. Gelsinger Bing Ma Jacek Wierzchos Jacques Ravel Alfonso Davila M. Cristina Casero Jocelyne DiRuggiero|url=https://doi.org/10.1111/1462-2920.13259|날짜=21 March 2016|권=18|호=2016 v.18 no.6|쪽=2064–2077|doi=10.1111/1462-2920.13259|pmid=26914534|확인날짜=2 March 2021}}</ref>을 포함하는 다양한 고염분 환경의 데이터에서 그 혈통이 확인되었다.

== 하위 분류 ==
현재 인정되는 분류는 [[LPSN]]<ref>{{웹 인용|url=https://lpsn.dsmz.de/phylum/nanohaloarchaeota|제목=Phylum "Candidatus Nanohaloarchaeota"|성=J.P. Euzéby|출판사=[[List of Prokaryotic names with Standing in Nomenclature]] (LPSN)|url-status=live|확인날짜=2011-11-17}}</ref> 및 [[NCBI]]에 기초를 두고 있다.<ref>{{웹 인용|url=https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Tree&id=1051663&lvl=3&lin|제목=Nanohaloarchaea|성=Sayers|출판사=[[National Center for Biotechnology Information]] (NCBI) taxonomy database|확인날짜=2011-06-05|저자표시=etal}}</ref>{{분기도|{{clade|style=font-size:90%;line-height:90%;width:425px
|1={{Clade
  |1="''Candidatus'' [[Nanosalinicola]]"
  |2={{Clade
    |1="''Candidatus'' [[Nanosalina]]"
    |2="''Candidatus'' [[Nanohalobium]]"
     }}
   }}
}}}}Phylum "Nanohaloarchaeota" <small>corrig.</small> <small>Rinkeet al. 2013년</small>

* Class "Nanohaloarchaeia" <small>corrig.</small> <small>Narasingarao et al. 2012</small>
** Order "Nanohaloarchaeales"
*** Family "Nanohaloarchaeaceae"
**** Genus ?"''Candidatus'' [[Nanohaloarchaeum]]" <small>corrig.</small> <small>Hamm et al. 2019</small>
***** "''Ca.'' N. antarcticum" <small>corrig.</small> <small>Hamm et al. 2019</small>
* Class "Nanohalobiia" <small>corrig.</small> <small>La Cono et al. 2020</small>
** Order "Nanohalobiales" <small>La Cono et al. 2020</small><ref>{{저널 인용|제목=Symbiosis between nanohaloarchaeon and haloarchaeon is based on utilization of different polysaccharides|저널=Proc Natl Acad Sci USA|성=La Cono|날짜=2020|권=117|호=33|쪽=20223–20234|bibcode=2020PNAS..11720223L|doi=10.1073/pnas.2007232117|pmc=7443923|pmid=32759215|저자표시=et al.}}</ref> ["Nanosalinales" <small>Rinke et al. 2020</small>]
*** Family "Nanohalobiaceae" <small>La Cono et al. 2020</small> ["Nanosalinaceae" <small>Rinke et al. 2020</small>]
**** Genus ?"''Candidatus'' [[Haloredivivus]]" <small>Ghai et al. 2011</small>
**** Genus ?"''Candidatus'' [[Nanopetraeus]]" <small>corrig.</small> <small>Crits‐Christoph et al. 2016</small> ["Nanopetramus" (sic)]
**** Genus "''Candidatus'' [[Nanohalobium]]" <small>La Cono et al. 2020</small>
***** "''Ca.'' N. constans" <small>La Cono et al. 2020</small>
**** Genus "''Candidatus'' [[Nanosalina]]" <small>Narasingarao et al. 2012</small>
**** Genus "''Candidatus'' [[Nanosalinicola]]" <small>corrig.</small> <small>Narasingarao et al. 2012</small> ["Nanosalinarum" (sic)]

== 계통 분류 ==
{{계통도/고세균}}

== 각주 ==
{{각주}}
* {{저널 인용|제목=New Abundant Microbial Groups in Aquatic Hypersaline Environments|저널=Scientific Reports|성=Ghai|날짜=2011|권=1|호=135|쪽=135|bibcode=2011NatSR...1E.135G|doi=10.1038/srep00135|pmc=3216616|pmid=22355652|저자표시=etal}}
* {{저널 인용|제목=De novo metagenomic assembly reveals abundant novel major lineage of Archaea in hypersaline microbial communities|저널=ISME J.|성=Narasingarao|날짜=2012|권=6|호=1|쪽=81–93|doi=10.1038/ismej.2011.78|pmc=3246234|pmid=21716304|저자표시=etal}}

{{고균}}
{{분류군 식별자|from=Q6964054}}

[[분류:고균]]