{{위키데이터 속성 추적}}
{{생물 분류
|이름=합티스타
|색=진핵생물
|그림=Coccolithus pelagicus.jpg
|그림_설명=
|역=[[진핵생물]]
{{미분류 | 분류=[[식물+HC+SAR 대형군]]}}
{{미분류 | 분류=[[하크로비아]]}}
|미분류_문='''합티스타'''<br/>(Haptista)
|미분류_문_명명=Cavalier-Smith 2015
|하위_링크=아문
|하위=
* [[앙코라키스타 트위스타|알베이다류]] (Alveidia)
* [[육질태양충류]] (Centrohelida)
* [[착편모조류]] (Haptophyta)
|이명=
}}
'''합티스타'''(Haptista)는 제안된 [[원생생물]] 분류군의 하나로 [[육질태양충류]]와 [[착편모조류]] 등으로 이루어져 있다. 계통유전체학 연구 결과 [[SAR 상군]]과 자매군을 형성하는 것으로 추정된다.<ref name=Burki2016>{{인용|last=Burki F, |last2=Kaplan M, |last3=Tikhonenkov DV, |last4=Zlatogursky V, |last5=Minh BQ, |last6=Radaykina LV, |last7=Smirnov A, |last8=Mylnikov AP, |last9=Keeling PJ |year=2016 |title=Untangling the early diversification of eukaryotes: a phylogenomic study of the evolutionary origins of Centrohelida, Haptophyta and Cryptista. |journal=Proc Biol Sci |volume=283 |issue=1823 |pmid=26817772 |doi=10.1098/rspb.2015.2802 |pmc=4795036 |pages=20152802}}</ref><ref>{{저널 인용|pmid=20333193 |year=2009 |month= |author1=Burki, F |author2=Inagaki, Y |author3=Bråte, J |author4=Archibald, J. |author5=Keeling, P. |author6=Cavalier-Smith, T |author7=Sakaguchi, M |author8=Hashimoto, T |author9=Horak, A |author10=Kumar, S |author11=Klaveness, D |author12=Jakobsen, K.S |author13=Pawlowski, J |author14=Shalchian-Tabrizi, K |title=Large-scale phylogenomic analyses reveal that two enigmatic protist lineages, Telonemia and Centroheliozoa, are related to photosynthetic chromalveolates. |volume=1 |issue= |pages=231–8 |issn= |doi=10.1093/gbe/evp022 |journal=Genome Biology and Evolution |url=http://gbe.oxfordjournals.org/cgi/content/short/evp022v1?rss=1 |format=Free full text |pmc=2817417 |url-status=dead |archiveurl = https://archive.today/20120710011541/http://gbe.oxfordjournals.org/cgi/content/short/evp022v1?rss=1 |archivedate=2012-07-10 |df= }}</ref>

== 하위 분류 ==
캐빌리어-스미스(Cavalier-Smith), 차오와 루이스(Chao & Lewis 2015)<ref name=>{{인용|last=Cavalier-Smith |last2=Chao |last3=Lewis |year=2015 |title=Multiple origins of Heliozoa from flagellate ancestors: New cryptist subphylum Corbihelia, superclass Corbistoma, and monophyly of Haptista, Cryptista, Hacrobia and Chromista |journal=Molecular Phylogenetics and Evolution |volume=93 |issue= | pages=331-362 | |pmid= 26234272|doi=10.1016/j.ympev.2015.07.004}}</ref>, 루지에로 등(Ruggiero et al. 2015)<ref>{{인용| date=2015| author = Ruggiero| title =  Higher Level Classification of All Living Organisms | url=http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0119248| volume=10 |issue=4 |pages= e0119248|journal=PLoS ONE | doi=10.1371/journal.pone.0119248 |display-authors=etal | pmid=25923521 | pmc=4418965}}</ref>의 연구에 기초한 하위 분류는 다음과 같다.

* 알베이다아문 (Alveidia) <small>Cavalier-Smith 2018</small>
** 알베이다강 (Alveidea) <small>Cavalier-Smith 2018</small><ref name="Cavalier-Smith 2018">{{저널 인용 | date=17 April 2018 | last=Cavalier-Smith | first=T. | last2=Chao | first2=E. E. |last3=Lewis | first3=R. |title =Multigene phylogeny and cell evolution of chromist infrakingdom Rhizaria: contrasting cell organisation of sister phyla Cercozoa and Retaria | volume= 255|issue= 5|pages=1517–1574 |journal=Protoplasma | doi= 10.1007/s00709-018-1241-1| pmid= 29666938| pmc=6133090}}</ref>
*** [[알베이다목]] (Alveida)
**** [[앙코라키스타 트위스타]] (''Ancoracysta twista'')<ref name="Cavalier-Smith 2018" />
* 육질태양충아문 (Centroheliozoa) <small>Cushman & Jarvis 1929 sensu Durrschmidt & Patterson 1987</small> [Heliozoa <small>Haeckel 1862 stat. n. Margulis 1974 em. Cavalier-Smith 2003</small>]
** [[육질태양충류|육질태양충강]] (Centrohelea) <small>Kuhn 1926 stat. n. Cavalier-Smith 1993</small> [Centroplastiales; Centrohelina <small>Hartmann 1913</small>; Centroplasthelida <small>Febvre-Chevalier 1984</small>]
* [[착편모조류|착편모조아문]] (Haptophytina) Cavalier-Smith 2015 (Haptophyta <small>Hibberd 1976 sensu Ruggerio et al. 2015</small>)
** Rappemonada <small>Kim et al. 2011</small>
*** Rappephyceae <small>Cavalier-Smith 2015</small>
** 착편모조강 (Haptomonada) <small>(Margulis & Schwartz 1998)</small> [Haptophyta Hibberd 1976 emend. Edvardsen & Eikrem 2000; Prymnesiophyta <small>Green & Jordan, 1994</small>; Prymnesiomonada; Prymnesiida Hibberd 1976; Haptophyceae Christensen 1962 ex Silva 1980; Haptomonadida; Patelliferea Cavalier-Smith 1993]
*** [[파블로바과|파블로바강]] (Pavlovophyceae) <small>(Cavalier-Smith 1986) Green & Medlin 2000</small>
*** [[후각편모조강]] (Prymnesiophyceae) <small>Christensen 1962 emend. Cavalier-Smith 1996</small> [Haptophyceae s.s.; Prymnesiophycidae <small>Cavalier-Smith 1986</small>; Coccolithophyceae <small>Casper 1972 ex Rothmaler 1951</small>]

== 계통 분류 ==
{{계통도/진핵생물}}
{{계통도/하크로비아}}

== 각주 ==
{{각주}}

== 외부 링크 ==
* [http://tolweb.org/Hacrobia/124797 생명의 나무: 하크로비아(Hacrobia)]

{{진핵생물}}
{{하크로비아}}

{{분류군 식별자|from=Q21588562}}

[[분류:하크로비아]]
[[분류:토머스 캐빌리어 스미스가 명명한 분류군]]