{{위키데이터 속성 추적}}
{{생물 분류
| 이름      = 꼭두서니아과
| 화석_범위 = 
| 그림      = Rubia tinctorum flowers.jpg
| 그림_설명 = ''Rubia tinctorum''
| 상태      = 
| 상태_기준 = 
| 상태_출처 = 
| 색        = 식물
| 역        = 
| 계        = [[식물|식물계]]
| 미분류_문 = [[속씨식물|속씨식물군]]
| 미분류_강 = [[진정쌍떡잎식물|진정쌍떡잎식물군]]
| 미분류_목 = [[국화군]]
| 목        = [[용담목]]
| 과        = [[꼭두서니과]]
| 아과      = '''꼭두서니아과'''<br>Rubioideae
| 아과_명명 = [[버나드 버드코트|Verdc.]], 1958
| 지도      = 
| 지도_설명 = 
| 모식속    = [[꼭두서니속]](''Rubia'' <small>L.</small>)
| 하위_링크 = [[속 (생물학)|속]]
| 하위      = * [[#하위 분류|본문 참조]]
}}
'''꼭두서니아과'''(----亞科, <small>[[학명]]:</small> {{lang|la|Rubioideae|루비오이데아이}})는 [[꼭두서니과]]의 [[아과]]이다.<ref>Verdcourt, Bernard. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 280. 1958.</ref> 26개 족, 약 7600여 종으로 이루어져 있다.<ref>Juss., 1789. ''Genera Plantarum'' 196.</ref><ref>B. Bremer and J.-F. Manen, ''Phylogeny and classification of the subfamily Rubioideae (Rubiaceae)'' Plant Syst. Evol. 225:43-72 (2000) [http://www.bergianska.se/pub/publikationer/Bremer/Bremer_and_Manen_2000.pdf PDF].</ref> [[갈퀴덩굴]], [[계요등]], [[꼭두서니]], [[노니]], [[민둥갈퀴]], [[솔나물]], [[수정목]], [[호자나무]] 등을 포함하고 있다.

== 하위 분류 ==
{{columns-list|colwidth=20em|
; [[가이르트네라족]](Gaertnereae <small>[[슈테판 라디슬라우스 엔틀리허|Endl.]]</small>)<ref>Endlicher, Stephan Friedrich Ladislaus. Genera Plantarum (Endlicher) 1: 577. 1838.</ref>
* ''Gaertnera'' <small>Lam.</small>
* ''Pagamea'' <small>Aubl.</small>

; [[계요등족]](Paederieae <small>[[오귀스탱 피람 드 캉돌|DC.]]</small>)<ref>Candolle, Augustin Pyramus de. Prodromus Systematis Naturalis Regni Vegetabilis 4: 342, 470. 1830.</ref>
* [[계요등속]](''Paederia'' <small>L.</small>)
* ''Leptodermis'' <small>Wall.</small>
* ''Pseudopyxis'' <small>Miq.</small>
* ''Saprosma'' <small>Blume</small>
* ''Serissa'' <small>Comm. ex A.Juss.</small>
* ''Spermadictyon'' <small>Roxb.</small>

; [[꼭두서니족]](Rubieae <small>[[앙리 에르네스트 바용|Baill.]]</small>)<ref>Baillon, Henri Ernest. Histoire des Plantes 7: 365, 390. 1880</ref>
* [[갈퀴덩굴속]](''Galium'' <small>L.</small>)
* [[갈퀴아재비속]](''Asperula'' <small>L.</small>)
* [[꼭두서니속]](''Rubia'' <small>L.</small>)
* [[꽃갈퀴덩굴속]](''Sherardia'' <small>L.</small>)
* ''Callipeltis'' <small>Steven</small>
* ''Crucianella'' <small>L.</small>
* ''Cruciata'' <small>Mill.</small>
* ''Didymaea'' <small>Hook.f.</small>
* × ''Galiasperula'' <small>Ronniger</small>
* ''Kelloggia'' <small>Torr. ex Benth. & Hook.f.</small>
* ''Mericarpaea'' <small>Boiss.</small>
* ''Microphysa'' <small>Schrenk</small>
* ''Phuopsis'' <small>(Griseb.) Hook.f.</small>
* ''Valantia'' <small>L.</small>

; [[노니족]](Morindeae <small>[[빈첸츠 프란츠 코스텔레츠키|Kostel.]]</small>)<ref>Kosteletzky, Vincenz Franz. Allgemeine Medizinisch-Pharmazeutische Flora 2: 565. 1833.</ref>
* [[노니속]](''Morinda'' <small>L.</small>)
* ''Appunia'' <small>Hook.f.</small>
* ''Coelopyrena'' <small>Valeton</small>
* ''Coelospermum'' <small>Blume</small>
* ''Gynochthodes'' <small>Blume</small>

; [[다나이스족]](Danaideae <small>[[비르기타 브레메르|B.Bremer]] & [[장프랑수아 마넨|Manen]]</small>)<ref>Bremer, Birgitta & Manen, Jean-François. Plant Systematics and Evolution 225: 60. 2000.</ref>
* ''Danais'' <small>Comm. ex Vent.</small>
* ''Payera'' <small>Baill.</small>
* ''Schismatoclada'' <small>Baker</small>

; [[둔니아족]](Dunnieae <small>[[카타리나 뤼딘|Rydin]] & [[비르기타 브레메르|B.Bremer]]</small>)<ref>Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 121. 2009.</ref>
* ''Dunnia'' <small>Tutcher</small>

; [[무주나무족]](Lasiantheae <small>[[비르기타 브레메르|B.Bremer]] & [[장프랑수아 마넨|Manen]]</small>)<ref>Bremer, Birgitta & Manen, Jean-François. Plant Systematics and Evolution 225: 57. 2000.</ref>
* [[무주나무속]](''Lasianthus'' <small>Jack</small>)
* ''Ronabea'' <small>Aubl.</small>
* ''Saldinia'' <small>A.Rich. ex DC.</small>
* ''Trichostachys'' <small>Hook.f.</small>

; [[스크라데라족]](Schradereae <small>[[코르넬리스 엘리자 베르튀스 브레메캄프|Bremek.]]</small>)<ref>Bremekamp, Cornelis Eliza Bertus. Recueil des Travaux Botaniques Néerlandais 31: 253. 1934.</ref>
* ''Lecananthus'' <small>Jack</small>
* ''Leucocodon'' <small>Gardner</small>
* ''Schradera'' <small>Vahl</small>

; [[스키조콜레아족]](Schizocoleeae <small>[[카타리나 뤼딘|Rydin]] & [[비르기타 브레메르|B.Bremer]]</small>)<ref>Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 120. 2009.</ref>
* ''Schizocolea'' <small>Bremek.</small>

; [[스페르마코케족]](Spermacoceae <small>[[프리드리히 프셰미르 폰 베르히톨트|Bercht.]] & [[얀 스바토플루크 프레슬|J.Presl]]</small>)<ref>Berchtold, Bedřich Všemír von & Presl, Jan Svatopluk. O Prirozenosti Rostlin 256. 1820.</ref>
* [[낚시돌풀속]](''Leptopetalum'' <small>Hook. & Arn.</small>)
* [[백령풀속]](''Hexasepalum'' <small>Bartl. ex DC.</small>)
* [[백운풀속]](''Scleromitrion'' <small>(Wight & Arn.) Meisn.</small>)
* [[제주백운풀속]](''Oldenlandia'' <small>L.</small>)
* [[큰백령풀속]](''Diodia'' <small>L.</small>)
* [[탐라풀속]](''Neanotis'' <small>W.H.Lewis</small>)
* ''Agathisanthemum'' <small>Klotzsch</small>
* ''Amphiasma'' <small>Bremek.</small>
* ''Amphistemon'' <small>Groeninckx</small>
* ''Anthospermopsis'' <small>(K.Schum.) J.H.Kirkbr.</small>
* ''Arcytophyllum'' <small>Willd.</small>
* ''Astiella'' <small>Jovet</small>
* ''Bouvardia'' <small>Salisb.</small>
* ''Carajasia'' <small>R.M.Salas, E.L.Cabral & Dessein</small>
* ''Carterella'' <small>Terrell</small>
* ''Conostomium'' <small>(Stapf) Cufod.</small>
* ''Cordylostigma'' <small>Groeninckx & Dessein</small>
* ''Crusea'' <small>Schltdl. & Cham.</small>
* ''Debia'' <small>Neupane & N.Wikstr.</small>
* ''Denscantia'' <small>E.L.Cabral & Bacigalupo</small>
* ''Dentella'' <small>J.R.Forst. & G.Forst.</small>
* ''Diacrodon'' <small>Sprague</small>
* ''Dibrachionostylus'' <small>Bremek.</small>
* ''Dimetia'' <small>(Wight & Arn.) Meisn.</small>
* ''Edrastima'' <small>Raf.</small>
* ''Eleuthranthes'' <small>F.Muell. ex Benth.</small>
* ''Emmeorhiza'' <small>Pohl ex Endl.</small>
* ''Ernodea'' <small>Sw.</small>
* ''Exallage'' <small>Bremek.</small>
* ''Galianthe'' <small>Griseb.</small>
* ''Gomphocalyx'' <small>Baker</small>
* ''Hedyotis'' <small>L.</small>
* ''Hedythyrsus'' <small>Bremek.</small>
* ''Houstonia'' <small>L.</small>
* ''Hydrophylax'' <small>L.f.</small>
* ''Involucrella'' <small>(Benth. & Hook.f.) Neupane & N.Wikstr.</small>
* ''Kadua'' <small>Cham. & Schltdl.</small>
* ''Kohautia'' <small>Cham. & Schltdl.</small>
* ''Lelya'' <small>Bremek.</small>
* ''Leptomischus'' <small>Drake</small>
* ''Manettia'' <small>Mutis ex L.</small>
* ''Manostachya'' <small>Bremek.</small>
* ''Mexotis'' <small>Terrell & H.Rob.</small>
* ''Mitracarpus'' <small>Zucc.</small>
* ''Mitrasacmopsis'' <small>Jovet</small>
* ''Nesohedyotis'' <small>(Hook.f.) Bremek.</small>
* ''Oldenlandiopsis'' <small>Terrell & W.H.Lewis</small>
* ''Opercularia'' <small>Gaertn.</small>
* ''Pentanopsis'' <small>Rendle</small>
* ''Pentodon'' <small>Hochst.</small>
* ''Phialiphora'' <small>Groeninckx</small>
* ''Phyllocrater'' <small>Wernham</small>
* ''Phylohydrax'' <small>Puff</small>
* ''Planaltina'' <small>R.M.Salas & E.L.Cabral</small>
* ''Pseudonesohedyotis'' <small>Tennant</small>
* ''Psyllocarpus'' <small>Mart. & Zucc.</small>
* ''Richardia'' <small>L.</small>
* ''Sacosperma'' <small>G.Taylor</small>
* ''Schwendenera'' <small>K.Schum.</small>
* ''Spermacoce'' <small>L.</small>
* ''Staelia'' <small>Cham. & Schltdl.</small>
* ''Stenaria'' <small>(Raf.) Terrell</small>
* ''Stenotis'' <small>Terrell</small>
* ''Synaptantha'' <small>Hook.f.</small>
* ''Tessiera'' <small>DC.</small>
* ''Thamnoldenlandia'' <small>Groeninckx</small>

; [[아르고스템마족]](Argostemmateae <small>[[코르넬리스 엘리자 베르튀스 브레메캄프|Bremek.]] ex [[버나드 버드코트|Verdc.]]</small>)<ref>Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.</ref>
* ''Argostemma'' <small>Wall.</small>
* ''Mouretia'' <small>Pit.</small>
* ''Mycetia'' <small>Reinw.</small>
* ''Neohymenopogon'' <small>Bennet</small>

; [[안토스페르뭄족]](Anthospermeae <small>[[아델베르트 폰 샤미소|Cham.]] & [[디데리히 프란츠 레온하르트 폰 슐레히텐달|Schltdl.]]</small>)<ref>Chamisso, Ludolf Karl Adelbert von & Schlechtendal, Diederich Franz Leonhard von. Linnaea 3: 309. 1828.</ref>
* ''Anthospermum'' <small>L.</small>
* ''Carpacoce'' <small>Sond.</small>
* ''Coprosma'' <small>J.R.Forst. & G.Forst.</small>
* ''Durringtonia'' <small>R.J.F.Hend. & Guymer</small>
* ''Galopina'' <small>Thunb.</small>
* ''Leptostigma'' <small>Arn.</small>
* ''Nenax'' <small>Gaertn.</small>
* ''Nertera'' <small>Banks ex Gaertn.</small>
* ''Normandia'' <small>Hook.f.</small>
* ''Phyllis'' <small>L.</small>
* ''Pomax'' <small>Sol. ex DC.</small>

; [[오피오리자족]](Ophiorrhizeae <small>[[코르넬리스 엘리자 베르튀스 브레메캄프|Bremek.]] ex [[버나드 버드코트|Verdc.]]</small>)<ref>Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.</ref>
* ''Coptophyllum'' <small>Korth.</small>
* ''Keenania'' <small>Hook.f.</small>
* ''Lerchea'' <small>L.</small>
* ''Neurocalyx'' <small>Hook.</small>
* ''Ophiorrhiza'' <small>L.</small>
* ''Spiradiclis'' <small>Blume</small>
* ''Xanthophytum'' <small>Reinw. ex Blume</small>

; [[우로필룸족]](Urophylleae <small>[[코르넬리스 엘리자 베르튀스 브레메캄프|Bremek.]] ex [[버나드 버드코트|Verdc.]]</small>)<ref>Bremekamp, Cornelis Eliza Bertus. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.</ref>
* ''Amphidasya'' <small>Standl.</small>
* ''Antherostele'' <small>Bremek.</small>
* ''Pauridiantha'' <small>Hook.f.</small>
* ''Pentaloncha'' <small>Hook.f.</small>
* ''Raritebe'' <small>Wernham</small>
* ''Temnopteryx'' <small>Hook.f.</small>
* ''Urophyllum'' <small>Jack ex Wall.</small>

; [[코우사레아족]](Coussareeae <small>[[조지프 돌턴 후커|Hook.f.]]</small>)<ref>Hooker, Joseph Dalton. Genera Plantarum 2: 9. 1873.</ref>
* ''Bradea'' <small>Standl.</small>
* ''Coccocypselum'' <small>P.Browne</small>
* ''Coussarea'' <small>Aubl.</small>
* ''Cruckshanksia'' <small>Hook. & Arn.</small>
* ''Declieuxia'' <small>Kunth</small>
* ''Faramea'' <small>Aubl.</small>
* ''Heterophyllaea'' <small>Hook.f.</small>
* ''Hindsia'' <small>Benth. ex Lindl.</small>
* ''Oreopolus'' <small>Schltdl.</small>

; [[콜레토이케마족]](Colletoecemateae <small>[[카타리나 뤼딘|Rydin]] & [[비르기타 브레메르|B.Bremer]]</small>)<ref>Rydin, Catarina & Bremer, Birgitta. Plant Systematics and Evolution 278: 120. 2009.</ref>
* ''Colletoecema'' <small>E.M.A.Petit</small>

; [[크녹시아족]](Knoxieae <small>[[조지 벤섬|Benth.]] & [[조지프 돌턴 후커|Hook.f.]]</small>)<ref>Bentham, George & Hooker, Joseph Dalton. Genera Plantarum 2: 9, 21. 1873.</ref>
* ''Batopedina'' <small>Verdc.</small>
* ''Carphalea'' <small>Juss.</small>
* ''Chamaepentas'' <small>Bremek.</small>
* ''Dirichletia'' <small>Klotzsch</small>
* ''Dolichopentas'' <small>Kårehed & B.Bremer</small>
* ''Knoxia'' <small>L.</small>
* ''Lathraeocarpa'' <small>Bremek.</small>
* ''Otiophora'' <small>Zucc.</small>
* ''Otomeria'' <small>Benth.</small>
* ''Paracarphalea'' <small>Razafim., Ferm, B.Bremer & Kårehed</small>
* ''Paraknoxia'' <small>Bremek.</small>
* ''Parapentas'' <small>Bremek.</small>
* ''Pentanisia'' <small>Harv.</small>
* ''Pentas'' <small>Benth.</small>
* ''Phyllopentas'' <small>(Verdc.) Kårehed & B.Bremer</small>
* ''Rhodopentas'' <small>Kårehed & B.Bremer</small>
* ''Triainolepis'' <small>Hook.f.</small>

; [[크라테리스페르뭄족]](Craterispermeae <small>[[버나드 버드코트|Verdc.]]</small>)<ref>Verdcourt, Bernard. Bulletin du Jardin Botanique de l'État à Bruxelles 28: 281. 1958.</ref>
* ''Craterispermum'' <small>Benth.</small>

; [[키아노네우론족]](Cyanoneuroneae <small>[[실뱅 조르주 라자피만딤비손|Razafim.]] & [[비르기타 브레메르|B.Bremer]]</small>)<ref>Razafimandimbison, Sylvain Georges & Bremer, Birgitta. Taxon 64(2): 294. 2015.</ref>
* ''Cyanoneuron'' <small>Tange</small>

; [[텔리고눔족]](Theligoneae <small>[[앙리 에르네스트 바용|Baill.]]</small>)<ref>Baillon, Henri Ernest. Histoire des Plantes 4: 46, 55. 1872.</ref>
* ''Theligonum'' <small>L.</small>

; [[팔리코우레아족]](Palicoureeae <small>[[엘마르 로브레흐트|Robbr.]] & [[장프랑수아 마넨|Manen]]</small>)<ref>Robbrecht, Elmar & Manen, Jean-François. Systematics and Geography of Plants 76: 125, 136. 2006.</ref>
* ''Carapichea'' <small>Aubl.</small>
* ''Chassalia'' <small>Comm. ex Poir.</small>
* ''Ditrichanthus'' <small>Borhidi, E.Martínez & Ramos</small>
* ''Eumachia'' <small>DC.</small>
* ''Geophila'' <small>D.Don</small>
* ''Hymenocoleus'' <small>Robbr.</small>
* ''Notopleura'' <small>(Hook.f.) Bremek.</small>
* ''Palicourea'' <small>Aubl.</small>
* ''Puffia'' <small>Razafim. & B.Bremer</small>
* ''Rudgea'' <small>Salisb.</small>
* ''Tromlyca'' <small>Borhidi</small>

; [[페라마족]](Perameae <small>[[코르넬리스 엘리자 베르튀스 브레메캄프|Bremek.]] ex [[스티븐 P. 다윈|S.P.Darwin]]</small>)<ref>Bremekamp, Cornelis Eliza Bertus. Taxon 25(5-6): 605. 1976.</ref>
* ''Perama'' <small>Aubl.</small>

; [[퐁케비아족]](Foonchewieae <small>[[왕루이장|R.J.Wang]]</small>)<ref>Wang, Rui-jiang. Journal of Systematics and Evolution 50(5): 472–473. 2012.</ref>
* ''Foonchewia'' <small>R.J.Wang</small>

; [[푸토리아족]](Putorieae <small>[[로버트 스위트|Sweet]]</small>)<ref>Sweet, Robert. Hortus Britannicus, ed. 3 325. 1839.</ref>
* ''Plocama'' <small>Aiton</small>

; [[프리스마토메리스족]](Prismatomerideae <small>[[롼윈전|Y.Z.Ruan]]</small>)<ref>Ruan, Yun Zhen. Acta Phytotaxonomica Sinica 26(6): 446. 1988.</ref>
* ''Gentingia'' <small>J.T.Johanss. & K.M.Wong</small>
* ''Motleyia'' <small>J.T.Johanss.</small>
* ''Prismatomeris'' <small>Thwaites</small>
* ''Rennellia'' <small>Korth.</small>

; [[프시코트리아족]](Psychotrieae <small>[[아델베르트 폰 샤미소|Cham.]] & [[디데리히 프란츠 레온하르트 폰 슐레히텐달|Schltdl.]]</small>)<ref>Chamisso, Ludolf Karl Adelbert von & Schlechtendal, Diederich Franz Leonhard von. Linnaea 4(1): 4. 1829.</ref>
* ''Amaracarpus'' <small>Blume</small>
* ''Anthorrhiza'' <small>C.R.Huxley & Jebb</small>
* ''Calycosia'' <small>A.Gray</small>
* ''Dolianthus'' <small>C.H.Wright</small>
* ''Hedstromia'' <small>A.C.Sm.</small>
* ''Hydnophytum'' <small>Jack</small>
* ''Myrmecodia'' <small>Jack</small>
* ''Myrmephytum'' <small>Becc.</small>
* ''Psychotria'' <small>L.</small>
* ''Squamellaria'' <small>Becc.</small>
* ''Streblosa'' <small>Korth.</small>

; [[호자덩굴족]](Mitchelleae <small>[[실뱅 조르주 라자피만딤비손|Razafim.]] & [[비르기타 브레메르|B.Bremer]]</small>)<ref>Razafimandimbison, Sylvain Georges & Bremer, Birgitta. Molecular Phylogenetics and Evolution 48(1): 221. 2008.</ref>
* [[호자나무속]](''Damnacanthus'' <small>C.F.Gaertn.</small>)
* [[호자덩굴속]](''Mitchella'' <small>L.</small>)

; 분류 불확실(''[[incertae sedis]]'')
* ''Aphanocarpus'' <small>Steyerm.</small>
* ''Clarkella'' <small>Hook.f.</small>
* ''Coryphothamnus'' <small>Steyerm.</small>
* ''Pagameopsis'' <small>Steyerm.</small>
}}

== 계통 분류 ==
{{분지군|style=line-height:100%
|분류1=[[꼭두서니과]]<ref name="Bremer et al. 2009">{{저널 인용 |성1=Bremer |이름1=Birgitta |성2=Eriksson |이름2=Torsten |제목=Time Tree of Rubiaceae: Phylogeny and Dating the Family, Subfamilies, and Tribes |저널=International Journal of Plant Sciences |날짜=2009 |권=170 |호=6 |쪽=766–793 |doi=10.1086/599077 |url=http://www.bergianska.se/polopoly_fs/1.68504.1325516084!/menu/standard/file/Bremer_Eriksson_2009.pdf |언어=en}}</ref>
|1={{분지군
 |1='''꼭두서니아과'''
 |2={{분지군
  |1=[[치자나무아과]]
  |2=[[구슬꽃나무아과]]
  }}
 |3={{분지군
  |1=[[콥토사펠타족]]
  |2=[[루쿨리아족]]
  }}
 }}
}}

== 각주 ==
{{각주}}

{{분류군 식별자|from=Q6113134}}

{{전거 통제}}

[[분류:꼭두서니아과| ]]
[[분류:1958년 기재된 식물]]