Þis getæl hæfþ dēor þe efencyndhǣmende gebǣre habbaþ, ēac hǣmede, wōgung, mōdlufe, geocbende, and acennendhāde swā Bruce Bagemihl cȳðde in his 1999. gēares bēc Biological Exuberance: Animal Homosexuality and Natural Diversity.