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Canada's constitution consists of written text and unwritten traditions and conventions. The Constitution Act, 1867 (formerly the British North America Act) established governance based on parliamentary precedent "similar in principle to that of the United Kingdom" and divided powers between the federal and provincial governments. The Constitution Act, 1982 added a Canadian Charter of Rights and Freedoms, which guarantees basic rights and freedoms for Canadians that generally cannot be overridden by legislation of any level of government in Canada. However, a notwithstanding clause, allows the federal parliament and the provincial legislatures to override certain sections of the Charter temporarily, for a period of five years.

The Chamber of the House of Commons.

The monarch is represented by a viceroy, the Governor General, who is empowered to exercise almost all of the constitutional duties of the sovereign, though wielding these powers almost always on the advice of the appointed Queen's Privy Council for Canada. In practice, the only body to direct the use of the executive powers is the Cabinet a committee of the Privy Council made up of Ministers of the Crown, all of whom are responsible to the elected House of Commons. The Cabinet is headed by the Prime Minister, who holds the conventional position of head of government; to ensure the stability of government, the Governor General will usually appoint the person who is the current leader of the political party that can obtain the confidence of a plurality in the House of Commons. The Prime Minister chooses the Cabinet, and by convention, the Governor General respects the Prime Minister's choices. Michaëlle Jean has served as Governor General since September 27, 2005, and Stephen Harper, leader of the Conservative Party, has been her Prime Minister since February 6, 2006.

The federal parliament is made up of the Queen and two houses: an elected House of Commons and an appointed Senate. Each member in the House of Commons is elected by simple plurality in a riding or electoral district; general elections are called by the Governor General when the Prime Minister so advises. While there is no minimum term for a Parliament, a new election must be called within five years of the last general election. Members of the Senate, whose seats are apportioned on a regional basis, are chosen by the Prime Minister and formally appointed by the Governor General, and serve until age 75.

Four parties have substantial representation in the federal parliament: the Conservative Party of Canada, the Liberal Party of Canada, the New Democratic Party (NDP), and the Bloc Québécois. The current government is formed by the Conservative Party of Canada. While the Green Party of Canada and other smaller parties do not have current representation in Parliament, the list of historical parties with elected representation is substantial.

The Supreme Court of Canada in Ottawa, west of Parliament Hill.

Canada's judiciary plays an important role in interpreting laws and has the power to strike down laws that violate the Constitution. The Supreme Court of Canada is the highest court and final arbiter and is led by the Right Honourable Madam Chief Justice Beverley McLachlin, P.C. Its nine members are appointed by the Governor General on the advice of the Prime Minister. All judges at the superior and appellate levels are appointed by the Governor General on the advice of the prime minister and minister of justice, after consultation with non-governmental legal bodies. The federal cabinet appoints justices to superior courts at the provincial and territorial levels. Judicial posts at the lower provincial and territorial levels are filled by their respective governments (see Court system of Canada for more detail).

Common law prevails everywhere except in Quebec, where civil law predominates. Criminal law is solely a federal responsibility and is uniform throughout Canada. Law enforcement, including criminal courts, is a provincial responsibility, but in rural areas of all provinces except Ontario and Quebec, policing is contracted to the federal Royal Canadian Mounted Police (RCMP).

The Peacekeeping Monument in Ottawa.

Canada and the United States share the world's longest undefended border, co-operate on military campaigns and exercises, and are each other's largest trading partners. Canada has nevertheless maintained an independent foreign policy, most notably maintaining full relations with Cuba and declining participation in the Iraq War. Canada also maintains historic ties to the United Kingdom and France and to other former British and French colonies through Canada's membership in the Commonwealth of Nations and La Francophonie (French-Speaking Countries).

Canada currently employs a professional, volunteer military force of about 64,000 regular and 26,000 reserve personnel. The unified Canadian Forces (CF) comprise the army, navy, and air force. Major CF equipment deployed includes 1,400 armoured fighting vehicles, 34 combat vessels, and 861 aircraft.

Lester B. Pearson with 1957 Nobel Peace Prize.

Strong attachment to the British Empire and Commonwealth in English Canada led to major participation in British military efforts in the Second Boer War, the First World War, and the Second World War. Since then, Canada has been an advocate for multilateralism, making efforts to resolve global issues in collaboration with other nations.

Canada joined the United Nations in 1945 and became a founding member of NATO in 1949. During the Cold War, Canada was a major contributor to UN forces in the Korean War, and founded the North American Aerospace Defense Command (NORAD) in cooperation with the United States to defend against aerial attacks from the Soviet Union.

Canada has played a leading role in UN peacekeeping efforts. During the Suez Crisis of 1956, Lester B. Pearson eased tensions by proposing the inception of the United Nations Peacekeeping Force. Canada has since served in 50 peacekeeping missions, including every UN peacekeeping effort until 1989

and has since maintained forces in international missions in the former Yugoslavia and elsewhere.

Canada joined the Organization of American States (OAS) in 1990; Canada hosted the OAS General Assembly in Windsor in June 2000 and the third Summit of the Americas in Quebec City in April 2001. Canada seeks to expand its ties to Pacific Rim economies through membership in the Asia-Pacific Economic Cooperation forum (APEC).

Canadian soldiers in Afghanistan.

Since 2001, Canada has had troops deployed in Afghanistan as part of the US stabilization force and the UN-authorized, NATO-commanded International Security Assistance Force. Canada's Disaster Assistance Response Team (DART) has participated in three major relief efforts in the past two years; the two-hundred member team has been deployed in relief operations after the December 2004 tsunami in South Asia, Hurricane Katrina in September 2005 and the Kashmir earthquake in October 2005.

In February 2007, Canada, Italy, Britain, Norway, and Russia announced their funding commitments to launch a $1.5 billion project to help develop vaccines they said could save millions of lives in poor nations, and called on others to join them. " Rich Nations Launch Vaccine Pact". Reuters. February 10, 2007. In August 2007, Canadian sovereignty in Arctic waters was challenged following a Russian expedition which planted a Russian flag at the seabed at the North Pole. Canada has considered that area to be sovereign territory since 1925.

A geopolitical map of Canada, exhibiting its ten provinces and three territories.

Canada is a federation composed of ten provinces and three territories; in turn, these may be grouped into numerous regions. Western Canada consists of British Columbia and three Prairie provinces (Alberta, Saskatchewan, Manitoba). Eastern Canada consists of Central Canada (Quebec and Ontario) and Atlantic Canada (comprised of the three Maritime provinces of New Brunswick, Prince Edward Island, and Nova Scotia; and Newfoundland and Labrador). Three territories (Yukon, Northwest Territories, and Nunavut) comprise Northern Canada. Provinces have a large degree of autonomy from the federal government, territories somewhat less. Each has its own provincial or territorial symbols.

The provinces are responsible for most of Canada's social programs (such as health care, education, and welfare) and together collect more revenue than the federal government, an almost unique structure among federations in the world. Using its spending powers, the federal government can initiate national policies in provincial areas, such as the Canada Health Act; the provinces can opt out of these, but rarely do so in practice. Equalization payments are made by the federal government to ensure that reasonably uniform standards of services and taxation are kept between the richer and poorer provinces.

All provinces have unicameral, elected legislatures headed by a Premier selected in the same way as the Prime Minister of Canada. Each province also has a Lieutenant-Governor representing the Queen, analogous to the Governor General of Canada, appointed on the recommendation of the Prime Minister of Canada, though with increasing levels of consultation with provincial governments in recent years.

A satellite composite image of Canada. Boreal forests prevail on the rocky Canadian Shield. Ice and tundra are prominent in the Arctic. Glaciers are visible in the Canadian Rockies and Coast Mountains. Flat and fertile Prairies facilitate agriculture. The Great Lakes feed the St. Lawrence River (in the southeast) where lowlands host much of Canada's population.

Canada occupies a major northern portion of North America, sharing land borders with the contiguous United States to the south and with the US state of Alaska to the northwest, stretching from the Atlantic Ocean in the east to the Pacific Ocean in the west; to the north lies the Arctic Ocean. By total area (including its waters), Canada is the second largest country in the world, after Russia, and largest on the continent. By land area it ranks fourth, after Russia, China, and the United States. World Factbook: Area Country Comparison Table Since 1925, Canada has claimed the portion of the Arctic between 60°W and 141°W longitude, but this claim is not universally recognized. The northernmost settlement in Canada and in the world is Canadian Forces Station (CFS) Alert on the northern tip of Ellesmere Island—latitude 82.5°N—just 817 kilometres (450 nautical miles) from the North Pole. Canada has the longest coastline in the world: 243,000 kilometres.

The population density, , is among the lowest in the world. The most densely populated part of the country is the Quebec City-Windsor Corridor along the Great Lakes and Saint Lawrence River in the southeast.

To the north of this region is the broad Canadian Shield, an area of rock scoured clean by the last ice age, thinly soiled, rich in minerals, and dotted with lakes and rivers. Canada by far has more lakes than any other country and has a large amount of the world's freshwater.

The Horseshoe Falls in Ontario is the largest component of Niagara Falls, one of the world's most voluminous waterfalls, a major source of hydroelectric power, and a tourist destination.

In eastern Canada, the Saint Lawrence River widens into the Gulf of Saint Lawrence, the world's largest estuary, that contains the island of Newfoundland. South of the Gulf, the Canadian Maritimes protrude eastward along the Appalachian Mountain range from northern New England and the Gaspé Peninsula of Quebec. New Brunswick and Nova Scotia are divided by the Bay of Fundy, which experiences the world's largest tidal variations. Ontario and Hudson Bay dominate central Canada. West of Ontario, the broad, flat Canadian Prairies spread toward the Rocky Mountains, which separate them from British Columbia.

In western Canada, the Mackenzie River flows from the Great Slave Lake to the Arctic Ocean. A tributary of a tributary of the Mackenzie is the South Nahanni River, which is home to Virginia Falls, a waterfall about twice as high as Niagara Falls.

Northern Canadian vegetation tapers from coniferous forests to tundra and finally to Arctic barrens in the far north. The northern Canadian mainland is ringed with a vast archipelago containing some of the world's largest islands.

Average winter and summer high temperatures across Canada vary depending on the location. Winters can be harsh in many regions of the country, particularly in the interior and Prairie provinces which experience a continental climate, where daily average temperatures are near −15 °C (5 °F) but can drop below −40 °C (−40 °F) with severe wind chills. In non-coastal regions, snow can cover the ground almost six months of the year, (more in the north). Coastal British Columbia is an exception and enjoys a temperate climate with a mild and rainy winter.

On the east and west coast average high temperatures are generally in the low 20s °C (70s °F), while between the coasts the average summer high temperature ranges from 25 to 30 °C (75 to 85 °F) with occasional extreme heat in some interior locations exceeding 40 °C (104 °F). For a more complete description of climate across Canada see Environment Canada's Website.

Canadian banknotes depicting, top to bottom, Wilfrid Laurier, John A. Macdonald, Queen Elizabeth II, William Lyon Mackenzie King, and Robert Borden.

Canada is one of the world's wealthiest nations with a high per capita income, a member of the Organisation for Economic Co-operation and Development (OECD) and Group of Eight (G8). Canada is a free market economy with slightly more government intervention than the United States, but much less than most European nations. Canada has traditionally had a lower per capita gross domestic product (GDP) than its southern neighbour (whereas wealth has been more equally divided), but higher than the large western European economies . Since the early 1990's, the Canadian economy has been growing rapidly with low unemployment and large government surpluses on the federal level. Today Canada closely resembles the US in its market-oriented economic system, pattern of production, and high living standards. While as of October 2007, Canada's national unemployment rate of 5.9% is its lowest in 33 years. Provincial unemployment rates vary from a low of 3.6% in Alberta to a high of 14.6% in Newfoundland and Labrador.

In the past century, the growth of the manufacturing, mining, and service sectors has transformed the nation from a largely rural economy into one primarily industrial and urban. As with other first world nations, the Canadian economy is dominated by the service industry, which employs about three quarters of Canadians. However, Canada is unusual among developed countries in the importance of the primary sector, with the logging and oil industries being two of Canada's most important.

Canada is one of the few developed nations that is a net exporter of energy. Atlantic Canada has vast offshore deposits of natural gas and large oil and gas resources are centred in Alberta. The vast Athabasca Tar Sands give Canada the world's second largest reserves of oil behind Saudi Arabia. In Quebec, British Columbia, Newfoundland & Labrador, Ontario and Manitoba, hydroelectric power is a cheap and clean source of renewable energy.

Canada is one of the world's most important suppliers of agricultural products, with the Canadian Prairies one of the most important suppliers of wheat, canola and other grains. Canada is the world's largest producer of zinc and uranium and a world leader in many other natural resources such as gold, nickel, aluminum, and lead; many, if not most, towns in the northern part of the country, where agriculture is difficult, exist because of a nearby mine or source of timber. Canada also has a sizeable manufacturing sector centred in southern Ontario and Quebec, with automobiles and aeronautics representing particularly important industries.

Canada is highly dependent on international trade, especially trade with the United States. The 1989 Canada-US Free Trade Agreement (FTA) and 1994 North American Free Trade Agreement (NAFTA) (which included Mexico) touched off a dramatic increase in trade and economic integration with the US Since 2001, Canada has successfully avoided economic recession and has maintained the best overall economic performance in the G8. Since the mid 1990s, Canada's federal government has posted annual budgetary surpluses and has steadily paid down the national debt.

Toronto, Ontario skyline with the CN tower. Toronto is Canada's most populous metropolitan area with 5,113,149 people.

Canada's 2006 census counted 31,612,897, an increase of 5.4% since 2001. Population growth is from immigration and, to a lesser extent, natural growth. About three-quarters of Canada's population lives within 150 kilometres (90 mi) of the US border. A similar proportion live in urban areas concentrated in the Quebec City-Windsor Corridor (notably: the Greater Golden Horseshoe anchored around Toronto, Montreal, Ottawa, and their environs), the BC Lower Mainland (Vancouver and environs), and the Calgary-Edmonton Corridor in Alberta.

According to the 2001 census, it has 34 ethnic groups with at least one hundred thousand members each, with 83% of the total population claiming they are white. Ethnic diversity of Canada

The largest ethnic group is English (20.2%), followed by French (15.8%), Scottish (14.0%), Irish (12.9%), German (9.3%), Italian (4.3%), Chinese (3.7%), Ukrainian (3.6%), and First Nations (3.4%); 40% of respondents identified their ethnicity as "Canadian." Canada's aboriginal population is growing almost twice as fast as the Canadian average. In 2001, 13.4% of the population belonged to non-aboriginal visible minorities.

In 2001, 49% of the Vancouver population and 42.8% of Toronto's population were visible minorities. In March 2005, Statistics Canada projected that people of non-European origins will constitute a majority in both Toronto and Vancouver by 2012. Canadian People - Learn About Canada's People According to Statistics Canada's forecasts, the number of visible minorities in Canada is expected to double by 2017. Roughly one out of every five people in Canada could be a member of a visible minority by 2017. Visible majority by 2017

Canada has the highest per capita immigration rate in the world, driven by economic policy and family reunification; Canada also accepts large numbers of refugees. Newcomers settle mostly in the major urban areas of Toronto, Vancouver and Montreal. By the 1990s and 2000s, almost all of Canada’s immigrants came from Asia. Inflow of foreign-born population by country of birth, by year

Support for religious pluralism is an important part of Canada's political culture. According to 2001 census, 77.1% of Canadians identified as being Christians; of this, Catholics make up the largest group (43.6% of Canadians). The largest Protestant denomination is the United Church of Canada; about 16.5% of Canadians declare no religious affiliation, and the remaining 6.3% were affiliated with religions other than Christianity, of which the largest is Islam numbering 1.9%, followed by Judaism: 1.1%.

Canadian provinces and territories are responsible for education. Each system is similar while reflecting regional history, culture and geography. The mandatory school age ranges between 5–7 to 16–18 years, contributing to an adult literacy rate that is 99%. Postsecondary education is also administered by provincial and territorial governments, who provide most of the funding; the federal government administers additional research grants, student loans and scholarships. In 2002, 43% of Canadians aged between 25 and 64 had post-secondary education; for those aged 25 to 34 the post-secondary attainment reaches 51%.

A Kwakwaka'wakw totem pole and traditional "big house" in Victoria, BC.

Canadian culture has historically been influenced by British, French, and Aboriginal cultures and traditions. It has also been influenced by American culture because of its proximity and migration between the two countries. American media and entertainment are popular if not dominant in Canada; conversely, many Canadian cultural products and entertainers are successful in the US and worldwide. Many cultural products are marketed toward a unified "North American" or global market.

The creation and preservation of distinctly Canadian culture are supported by federal government programs, laws and institutions such as the Canadian Broadcasting Corporation (CBC), the National Film Board of Canada (NFB), and the Canadian Radio-television and Telecommunications Commission (CRTC).

Canada is a geographically vast and ethnically diverse country. There are cultural variations and distinctions from province to province and region to region. Canadian culture has also been greatly influenced by immigration from all over the world. Many Canadians value multiculturalism, and see Canadian culture as being inherently multicultural. Multicultural heritage is enshrined in Section 27 of the Canadian Charter of Rights and Freedoms.

The Royal Canadian Mounted Police, seen here at Expo 67, are the federal and national police force of Canada and an international icon.

National symbols are influenced by natural, historical, and First Nations sources. Particularly, the use of the maple leaf as a Canadian symbol dates back to the early 18th century and is depicted on its current and previous flags, the penny, and on the coat of arms. Other prominent symbols include the beaver, Canada goose, common loon, the Crown, and the RCMP.

Canada's official national sports are ice hockey (winter) and lacrosse (summer). Hockey is a national pastime and the most popular spectator sport in the country. It is the most popular sport Canadians play, with 1.65 million active participants in 2004. Canada's six largest metropolitan areas - Toronto, Montreal, Vancouver, Ottawa, Calgary, and Edmonton - have franchises in the National Hockey League (NHL), and there are more Canadian players in the league than from all other countries combined. After hockey, other popular spectator sports include curling and football; the latter is played professionally in the Canadian Football League (CFL). Golf, baseball, skiing, soccer, volleyball, and basketball are widely played at youth and amateur levels, but professional leagues and franchises are not as widespread.

Canada has hosted several high-profile international sporting events, including the 1976 Summer Olympics, the 1988 Winter Olympics, and the 2007 FIFA U-20 World Cup. Canada will be the host country for the 2010 Winter Olympics in Vancouver and Whistler, British Columbia.

The population of Montreal, Quebec is mainly Francophone, with a significant Anglophone community.

Canada's two official languages are English and French. Official Bilingualism in Canada is law, defined in the Canadian Charter of Rights and Freedoms, the Official Languages Act, and Official Language Regulations; it is applied by the Commissioner of Official Languages. English and French have equal status in federal courts, Parliament, and in all federal institutions. The public has the right, where there is sufficient demand, to receive federal government services in either English or French, and official language minorities are guaranteed their own schools in all provinces and territories.

English and French are the mother tongues of 59.7% and 23.2% of the population respectively, and the languages most spoken at home by 68.3% and 22.3% of the population respectively. 98.5% of Canadians speak English or French (English only: 67.5%, French only: 13.3%, both: 17.7%). English and French Official Language Communities, defined by First Official Language Spoken, constitute 73.0% and 23.6% of the population.

Although 85% of French-speaking Canadians live in Quebec, there are substantial Francophone populations in Ontario, Alberta and southern Manitoba, with an Acadian population in the northern and southeastern parts of New Brunswick constituting 35% of that province's population as well as concentrations in Southwestern Nova Scotia and on Cape Breton Island. Ontario has the largest French-speaking population outside Quebec. The Charter of the French Language in Quebec makes French the official language in Quebec, and New Brunswick is the only province to have a statement of official bilingualism in the constitution. Other provinces have no official language(s) as such, but French is used as a language of instruction, in courts, and other government services in addition to English. Manitoba, Ontario and Quebec allow for both English and French to be spoken in the provincial legislatures, and laws are enacted in both languages. In Ontario, French has some legal status but is not fully co-official. Several aboriginal languages have official status in Northwest Territories. Inuktitut is the majority language in Nunavut, and one of three official languages in the territory.

Non-official languages are important in Canada, with 5,202,245 people listing one as a first language. Some significant non-official first languages include Chinese (853,745 first-language speakers), Italian (469,485), German (438,080), and Punjabi (271,220).

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Beetles are a group of insects which have the largest number of species. They are placed in the order Coleoptera,which means "sheathed wing" and contains more described species than in any other order in the animal kingdom, constituting about twenty-five percent of all known life-forms. James K. Liebherr and Joseph V. McHugh in Resh, V. H. & R. T. Cardé (Editors) 2003. Encyclopedia of Insects. Academic Press. Forty percent of all described insect species are beetles (about 350,000 species ), and new species are frequently discovered. Estimates put the total number of species, described and undescribed, at between 5 and 8 million.

Beetles can be found in almost all habitats, but are not known to occur in the sea or in the polar regions. They interact with their ecosystems in several ways. They often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are prey of various animals including birds and mammals. Certain species are agricultural pests, such as the Colorado potato beetle Leptinotarsa decemlineata, the boll weevil Anthonomus grandis, the red flour beetle Tribolium castaneum, and the mungbean or cowpea beetle Callosobruchus maculatus, while other species of beetles are important controls of agricultural pests. For example, coccinellidae ("ladybirds" or "ladybugs") consume aphids, scale insects, thrips, and other plant-sucking insects that damage crops.

The name "Coleoptera" was given by Aristotle for the hardened shield like forewings (coleo = shield + ptera = wing).

A cockchafer with its elytra raised, exposing the membranous flight wings, where the veins are visible

Trogodendron fasciculatum, a clerid beetle with bright yellow antennae

Other characters of this group which are believed to be monophyletic include a holometabolous life cycle; having a prothorax that is distinct from and freely articulating with the mesothorax; the meso- and meta-thoracic segments fusing to form a pterothorax; a depressed body shape with the legs on the ventral surface; the coxae of legs recessed into cavities formed by heavily sclerotized thoracic sclerites; the abdominal sternites more sclerotized than the tergites; antennae with 11 or fewer segments; and terminal genitalic appendages retracted into the abdomen and invisible at rest.

The general anatomy of beetles is quite uniform, although specific organs and appendages may vary greatly in appearance and function between the many families in the order. Like all insects, beetles' bodies are divided into three sections: the head, the thorax, and the abdomen. When viewed from below, the thorax is that part from which all three pairs of legs and both pairs of wings arise. The abdomen is everything posterior to the thorax. When viewed from above, most beetles appear to have three clear sections, but this is deceptive: on the beetle's upper surface, the middle "section" is a hard plate called the pronotum, which is only the front part of the thorax; the back part of the thorax is concealed by the beetle's wings. Like all arthropods, beetles are segmented organisms, and all three of the major sections of the body are themselves composed of several further segments, although these are not always readily discernible. This further segmentation is usually best seen on the abdomen.

Beetles are generally characterised by a particularly hard exoskeleton and hard forewings (elytra). The beetle's exoskeleton is made up of numerous plates called sclerites, separated by thin sutures. This design creates the armoured defences of the beetle while maintaining flexibility. The elytra are not used for flight, but tend to cover the hind part of the body and protect the second pair of wings (alae). The elytra must be raised in order to move the hind flight wings. A beetle's flight wings are crossed with veins and are folded after landing, often along these veins, and are stored below the elytra.

In some beetles, the ability to fly has been lost. These include the ground beetles (family Carabidae) and some "true weevils" (family Curculionidae), but also some desert and cave-dwelling species of other families. Many of these species have the two elytra fused together, forming a solid shield over the abdomen. In a few families, both the ability to fly and the elytra have been lost, with the best known example being the glow-worms of the family Phengodidae, in which the females are larviform throughout their lives.

Beetles have mouthparts similar to those of grasshoppers. Of these parts, the most commonly known are probably the mandibles, which appear as large pincers on the front of some beetles. The mandibles are a pair of hard, often tooth-like structures that move horizontally to grasp, crush, or cut food or enemies (see defence, below). Two pairs of finger-like appendages are found around the mouth in most beetles, serving to move food into the mouth. These are the maxillary and labial palpi.

The eyes are compound and may display remarkable adaptability, as in the case of whirligig beetles (family Gyrinidae), in which the eyes are split to allow a view both above and below the waterline. Other species also have divided eyes — some longhorn beetles (family Cerambycidae) and weevils — while many beetles have eyes that are notched to some degree. A few beetle genera also possess ocelli, which are small, simple eyes usually situated farther back on the head (on the vertex).

Beetles' antennae are primarily organs of smell, but may also be used to feel out a beetle's environment physically. They may also be used in some families during mating, or among a few beetles for defence. Antennae vary greatly in form within the Coleoptera, but are often similar within any given family. In some cases, males and females of the same species will have different antennal forms. Antennae may be clavate (flabellate and lamellate are sub-forms of clavate, or clubbed antennae), filiform, geniculate, moniliform, pectinate, or serrate. For images of these antennal forms see antenna (biology).

Acilius sulcatus, a diving beetle showing hind legs adapted for life in water

The legs, which are multi-segmented, end in two to five small segments called tarsi. Like many other insect orders beetles bear claws, usually one pair, on the end of the last tarsal segment of each leg. While most beetles use their legs for walking, legs may be variously modified and adapted for other uses. Among aquatic families — Dytiscidae, Haliplidae, many species of Hydrophilidae and others — the legs, most notably the last pair, are modified for swimming and often bear rows of long hairs to aid this purpose. Other beetles have fossorial legs that are widened and often spined for digging. Species with such adaptations are found among the scarabs, ground beetles, and clown beetles (family Histeridae). The hind legs of some beetles, such as flea beetles (within Chrysomelidae) and flea weevils (within Curculionidae), are enlarged and designed for jumping.

Oxygen is obtained via a tracheal system. Air enters a series of tubes along the body through openings called spiracles, and is then taken into increasingly finer fibres. Pumping movements of the body force the air through the system.

Beetles have haemolymph instead of blood, and the open circulatory system of the beetle is powered by a tube-like heart attached to the top inside of the thorax.

Scarabaeiform larva of the cockchafer, Melolontha melolontha

Beetles are endopterygotes with complete metamorphosis.

A single female may lay from several dozen to several thousand eggs during her lifetime. Eggs are usually laid according to the substrate the larva will feed on upon hatching. Among others, they can be laid loose in the substrate (e.g. flour beetle), laid in clumps on leaves (e.g. Colorado potato beetle), or individually attached (e.g. mungbean beetle and other seed borers) or buried in the medium (e.g. carrot weevil).

The larva is usually the principal feeding stage of the beetle life cycle. Larvae tend to feed voraciously once they emerge from their eggs. Some feed externally on plants, such as those of certain leaf beetles, while others feed within their food sources. Examples of internal feeders are most Buprestidae and longhorn beetles. The larvae of many beetle families are predatory like the adults (ground beetles, ladybirds, rove beetles). The larval period varies between species but can be as long as several years.

Beetle larvae can be differentiated from other insect larvae by their hardened, often darkened head, the presence of chewing mouthparts, and spiracles along the sides of the body. Like adult beetles, the larvae are varied in appearance, particularly between beetle families. Beetles whose larvae are somewhat flattened and are highly mobile are the ground beetles, some rove beetles, and others; their larvae are described as campodeiform. Some beetle larvae resemble hardened worms with dark head capsules and minute legs. These are elateriform larvae, and are found in the click beetle (Elateridae) and darkling beetle (Tenebrionidae) families. Some elateriform larvae of click beetles are known as wireworms. Beetles in the families of the Scarabaeoidea have short, thick larvae described as scarabaeiform, but more commonly known as grubs.

All beetle larvae go through several instars, which are the developmental stages between each moult. In many species the larvae simply increase in size with each successive instar as more food is consumed. In some cases, however, more dramatic changes occur. Among certain beetle families or genera, particularly those that exhibit parasitic lifestyles, the first instar (the planidium) is highly mobile in order to search out a host, while the following instars are more sedentary and remain on or within their host. This is known as hypermetamorphosis; examples include the blister beetles (family Meloidae) and some rove beetles, particularly those of the genus Aleochara.

As with all endopterygotes, beetle larvae pupate, and from this pupa emerges a fully formed, sexually mature adult beetle, or imago. Adults have an extremely variable lifespan, from weeks to years, depending on the species.

Beetles mating in San Francisco