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Flora
The south Florida pine rockland is unique geographically. Southern Florida is located at the southern end of the temperate North American Floristic Region and at the northern end of the tropical Caribbean Floristic Region.[13] This location allows for the mixture of vegetation from two distinct areas at the extremes of their ranges.[14] Around 9% of all pine rockland plants found in Miami-Dade County alone are endemic to Florida, and around 14 taxa are endemic to the Miami-Dade pine rockland itself.[15][16] Overall, there are over 537 plant species found throughout the south Florida rocklands.[17]
Pine rocklands are defined by an open canopy of South Florida slash pine with heights ranging from 20–24 meters, but can be less due to past disturbance.[4] Seedlings are fire adapted and spend 2 to 5 years in the "grass stage" building nutrient reserves to facilitate growth above normal fire heights.[18] Many sites were logged extensively in the early 1990s reducing slash pine coverage and killing understory vegetation. Subsequent plantings in the late 1980s led to many even-aged stands with different stand densities.[19]
The subcanopy is dominated by a variety of palms and tropical hardwoods, depending on the location, substrate, and fire regime. Pine rocklands in the lower Keys often have high amounts of Thrinax and Coccothrinax, while rocklands in Miami-Dade County have high amounts of Sabal palmetto, Ficus aurea, and Serenoa repens. Sites that share borders with hardwood hammocks or that are infrequently burned will have high amounts of hardwood species, including Metopium toxiferum, Quercus elliotti, Quercus virginiana, Sideroxylon salicifolium, and Lysiloma latisiliquum. The shrub layer near wet prairies and marshes is composed of wetland species such as Acacia pinetorum, Sambucus canadensis, and Taxodium ascendens.
The herbaceous layer is extremely diverse and home to several species considered rare, endangered, threatened or critically imperiled by one or more agencies.[8] Many species found in rocklands throughout south Florida are restricted to individual plots or to specific regions due to changes in soil type, extreme fragmentation, and fire suppression. Most plants are fire-adapted and depend on frequent burning to limit shading and increased humidity from encroaching hardwood species. Grasses and sedges including Andropogon spp., Schizachyrium spp., Muhlenbergia capillaris, Arsitida purpurascens, and Rhynchospora spp. dominate the landscape. Flowering species include Croton linearis, Pinguicula pumilla, Angadenia berteroi, Amorpha herbacea var. crenulate, and a number of different Euphorbia spp.
Fire
Fire plays a critical role in maintaining the vegetative community.[7] It is estimated that around 70% of the state's terrestrial plant species are fired-adapted, fire-dependent, or pyrogenic, probably resulting from the state's high incidence of lightning strikes.[20][21] In fact, the Florida landscape is estimated to have been dominated by dry season lightning-induced and human-ignited fires.[22] Most of the native plant species found in pine rocklands are adapted to periodic fires, with increased abundance and flowering of native plants found in plots post-burn.[23] These fires help curb hardwood encroachment, spur pine regeneration, and allow light to reach the herbaceous layer.[24][25] In the absence of fire, hardwood species from nearby hardwood hammocks invade and shade out natural vegetation. As the density of hardwood species increases, fire effectiveness decreases due to the increase in humidity and accumulation of poor fire fuels. This ecotone between pine rockland and hardwood hammock is clear when natural or frequent, low-intensity prescribed fires occur. In the absence of frequent fire, this distinction becomes less apparent.
Presently, many isolated pine rockland sites throughout Miami-Dade County and the lower Keys suffer from fire suppression, leading to drastically altered plant compositions.[26] It is estimated that around 5,000,000 ha of pine rocklands burned in 1926, whereas only 76,486 ha burned in 1995 and 2003.[27] Most sites located in Miami-Dade County and the lower Keys are located near residential or commercial lots and are often unable to implement prescribed burns.[28] The longer fire is suppressed, the harder and more dangerous it is to prescribe effective burns. Fuel loads increase between fires and result in hotter and higher flames.[26] Fires that are too high or intense can result in crown fire or mortality at all levels of the canopy, including below ground biomass.[10] Frequent fire is used to great effect in the Long Pine Key in Everglades National Park.
Rockland hammock
Main article: Tropical hardwood hammock
Rockland hammocks form on regions of rockland where a lack of fire has allowed hardwood trees to become dominant, nearly all of which are tropical in origin.[29] Natural firebreaks include exposed limestone cliffs and solution sinkholes.[30] Canopy species include gumbo-limbo (Bursera simaruba), paradise tree (Simarouba glauca), pigeonplum (Coccoloba diversifolia), Florida strangler fig (Ficus aurea), false mastic (Sideroxylon foetidissimum), willow bustic (Dipholis salicifolia), short-leaf fig (Ficus citrifolia), false tamarind (Lysiloma latisiliquum), West Indian mahogany (Swietenia mahagoni), and pepperleaf sweetwood (Licaria triandra). Epiphytes that grow in the canopy include Spanish moss (Tillandsia usneoides) and ball moss (T. recurvata). Plants such as black ironwood (Krugiodendron ferreum), inkwood (Exothea paniculata), lancewood (Damburneya coriacea), marlberry (Ardisia escallonoides), poisonwood (Metopium toxiferum), satinleaf (Chrysophyllum oliviforme), white stopper (Eugenia axillaris), shiny oysterwood (Gymnanthes lucida), wild coffee (Psychotria nervosa), shortleaf wild coffee (Psychotria tenuifolia), and pale lidflower (Calyptranthes pallens) grow in the understory. Southern live oak (Quercus virginiana), a temperate species, can be found on hammock margins.[29]
Conservation
Because of its high elevation, the Miami Rock Ridge was the first area to be impacted by development. The clearing of large tracts for development has now reduced the pine rocklands to about 20,000 acres (81 km2), most of which are now protected inside the Everglades National Park. Camp Everglades is a 253-acre (102 ha) campground owned by the Boy Scouts and located within the park. The pine forest is fire dependent, and the flora and fauna have adapted to the frequent fires ignited by summer lightning storms. The camp has prescribed fires that help maintain the forest.[31]
MIAMI-DADE COUNTY
ENVIRONMENTALLY ENDANGERED LANDS PROGRAM
MANAGEMENT PLAN
PART II: MANAGEMENT OF SPECIFIC HABITAT TYPES
CHAPTER 1: THE PINE ROCKLAND HABITAT
Prepared by:
URS Corporation Southern
With the support of
The Institute for Regional Conservation and
Muller and Associates, Inc.
August 2007
EEL Program, Management Plan, Part II – Pine Rockland (DRAFT) _____________________________________________________________________________________________________________________
Page ii
PART II: Management of Specific Habitat Types
CHAPTER 1: The Pine Rockland Habitat
Table of Contents
1.0 INTRODUCTION .................................................................................................................................................1
1.1 OVERVIEW OF THE PINE ROCKLAND ECOSYSTEM................................................................................................1
1.2 PURPOSE ..............................................................................................................................................................2
2.0 HISTORICAL REFERENCE CONDITIONS FOR THE PINE ROCKLAND ECOSYSTEM ....................3
2.1 ORIGINAL PINE ROCKLAND DISTRIBUTION IN MIAMI-DADE COUNTY.................................................................3
2.2 PHYSIOGRAPHY ...................................................................................................................................................3
2.2.1 Geology.......................................................................................................................................................3
2.2.2 Soils.............................................................................................................................................................5
2.2.3 Hydrology....................................................................................................................................................5
2.3 CLIMATE..............................................................................................................................................................6
2.4 VEGETATION STRUCTURE AND COMPOSITION .....................................................................................................6
2.4.1 Canopy ........................................................................................................................................................6
2.4.2 Subcanopy ...................................................................................................................................................7
2.4.3 Herb Layer ..................................................................................................................................................7
2.5 ASSOCIATION WITH OTHER HABITAT TYPES .......................................................................................................8
2.6 HISTORICAL SUCCESSIONAL PROCESSES .............................................................................................................9
2.6.1 Fire............................................................................................................................................................10
2.6.2 Tropical Cyclones .....................................................................................................................................10
2.6.3 Freezes ......................................................................................................................................................11
2.7 RARE ORGANISMS .............................................................................................................................................11
3.0 CURRENT CONDITIONS, THREATS, AND TRENDS ................................................................................12
3.1 CURRENT DISTRIBUTION OF PINE ROCKLANDS..................................................................................................12
3.2 PHYSIOGRAPHY .................................................................................................................................................12
3.2.1 Soils...........................................................................................................................................................12
3.2.2 Hydrology..................................................................................................................................................14
3.2.3 Sea Level Rise ...........................................................................................................................................14
3.3 CLIMATE............................................................................................................................................................14
3.4 VEGETATION STRUCTURE AND COMPOSITION ...................................................................................................14
3.4.1 Canopy ......................................................................................................................................................15
3.4.2 Subcanopy .................................................................................................................................................15
3.4.3 Herb Layer ................................................................................................................................................16
3.4.4 Edges.........................................................................................................................................................16
3.5 ASSOCIATION WITH OTHER HABITAT TYPES .....................................................................................................16
3.6 SUCCESSIONAL PROCESSES UNDER CURRENT CONDITIONS...............................................................................16
3.6.1 Fire............................................................................................................................................................16
3.6.2 Tropical Cyclones .....................................................................................................................................17
3.6.3 Human Controlled Processes....................................................................................................................17
3.7 RARE ORGANISMS .............................................................................................................................................18
3.7.1 Plants ........................................................................................................................................................18
3.7.2 Animals .....................................................................................................................................................21
3.8 FRAGMENTATION, OWNERSHIP, AND PRESERVATION STATUS OF REMAINING FRAGMENTS ..............................23
3.8.1 Fragmentation...........................................................................................................................................23
3.8.2 EEL Program ............................................................................................................................................23
3.8.3 Parks & Recreation Department...............................................................................................................23
3.8.4 Other Public Lands ...................................................................................................................................23
3.8.5 Private Preserves ......................................................................................................................................24
EEL Program, Management Plan, Part II – Pine Rockland (DRAFT) _____________________________________________________________________________________________________________________
Page iii
3.8.6 Natural Forest Community System............................................................................................................24
3.8.7 Other .........................................................................................................................................................24
3.9 FUTURE PRESERVATION ESTIMATES..................................................................................................................25
3.10 EXOTIC ORGANISMS ........................................................................................................................................25
3.10.1 Plants ......................................................................................................................................................25
3.10.2 Animals ...................................................................................................................................................25
3.11 OTHER PROBLEM SPECIES ...............................................................................................................................27
3.12 POLLUTANTS....................................................................................................................................................28
3.13 CULTURAL RESOURCES ...................................................................................................................................28
4.0 MANAGEMENT ISSUES ..................................................................................................................................29
4.1 ACQUISITION NEEDS FOR REMAINING PINE ROCKLAND FRAGMENTS................................................................29
4.2 MITIGATION/MANAGEMENT FOR FRAGMENTATION EFFECTS............................................................................30
4.2.1 Acquisition and Restoration of Vacant Land between Parcels .................................................................30
4.2.2 Zoning Around and Between Parcels........................................................................................................30
4.3 TARGETS FOR VEGETATION STRUCTURE ...........................................................................................................31