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1#0 | Asam deoksiribonukleat | Asam deoksiribonukleat, lebih dikenal dengan singkatan DNA (bahasa Inggris: "deoxyribonucleic acid"), adalah sejenis biomolekul yang menyimpan dan menyandi instruksi-instruksi genetika setiap organisme dan banyak jenis virus. Instruksi-instruksi genetika ini berperan penting dalam pertumbuhan, perkembangan, dan fungsi organisme dan virus. DNA merupakan asam nukleat; bersamaan dengan protein dan karbohidrat, asam nukleat adalah makromolekul esensial bagi seluruh makhluk hidup yang diketahui. Kebanyakan molekul DNA terdiri dari dua unting biopolimer yang berpilin satu sama lainnya membentuk heliks ganda. Dua unting DNA ini dikenal sebagai polinukleotida karena keduanya terdiri dari satuan-satuan molekul yang disebut nukleotida. Tiap-tiap nukleotida terdiri atas salah satu jenis basa nitrogen (guanina (G), adenina (A), timina (T), atau sitosina (C)), gula monosakarida yang disebut deoksiribosa, dan gugus fosfat. Nukleotida-nukelotida ini kemudian tersambung dalam satu rantai ikatan kovalen antara gula satu nukleotida dengan fosfat nukelotida lainnya. Hasilnya adalah rantai punggung gula-fosfat yang berselang-seling. Menurut kaidah pasangan basa (A dengan T dan C dengan G), ikatan hidrogen mengikat basa-basa dari kedua unting polinukleotida membentuk DNA unting ganda | [
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1#1 | Asam deoksiribonukleat | Dua unting DNA bersifat anti-paralel, yang berarti bahwa keduanya berpasangan secara berlawanan. Pada setiap gugus gula, terikat salah satu dari empat jenis nukleobasa. Urutan-urutan empat nukleobasa di sepanjang rantai punggung DNA inilah yang menyimpan kode informasi biologis. Melalui proses biokimia yang disebut transkripsi, unting DNA digunakan sebagai templat untuk membuat unting RNA. Unting RNA ini kemudian ditranslasikan untuk menentukan urutan asam amino protein yang dibangun. | [
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1#2 | Asam deoksiribonukleat | Struktur kimia DNA yang ada membuatnya sangat cocok untuk menyimpan informasi biologis setiap makhluk hidup. Rantai punggung DNA resisten terhadap pembelahan kimia, dan kedua-dua unting dalam struktur unting ganda DNA menyimpan informasi biologis yang sama. Karenanya, informasi biologis ini akan direplikasi ketika dua unting DNA dipisahkan. Sebagian besar DNA (lebih dari 98% pada manusia) bersifat non-kode, yang berarti bagian ini tidak berfungsi menyandikan protein. | [
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1#3 | Asam deoksiribonukleat | Dalam sel, DNA tersusun dalam kromosom. Semasa pembelahan sel, kromosom-kromosom ini diduplikasi dalam proses yang disebut replikasi DNA. Organisme eukariotik (hewan, tumbuhan, fungi, dan protista) menyimpan kebanyakan DNA-nya dalam inti sel dan sebagian kecil sisanya dalam organel seperti mitokondria ataupun kloroplas. Sebaliknya organisme prokariotik (bakteri dan arkaea) menyimpan DNA-nya hanya dalam sitoplasma. Dalam kromosom, protein kromatin seperti histon berperan dalam penyusunan DNA menjadi struktur kompak. Struktur kompak inilah yang kemudian berinteraksi antara DNA dengan protein lainnya, sehingga membantu kontrol bagian-bagian DNA mana sajakah yang dapat ditranskripsikan. | [
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1#4 | Asam deoksiribonukleat | Para ilmuwan menggunakan DNA sebagai alat molekuler untuk menyingkap teori-teori dan hukum-hukum fisika, seperti misalnya teorema ergodik dan teori elastisitas. Sifat-sifat materi DNA yang khas membuatnya sangat menarik untuk diteliti bagi ilmuwan dan insinyur yang bekerja di bidang mikrofabrikasi dan nanofabrikasi material. Beberapa kemajuan di bidang material ini misalnya origami DNA dan material hibrida berbasis DNA. | [
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1#5 | Asam deoksiribonukleat | DNA merupakan sebuah polimer yang terdiri dari satuan-satuan berulang yang disebut nukleotida. Tiap-tiap nukleotida terdiri dari tiga komponen utama, yakni gugus fosfat, gula deoksiribosa, dan basa nitrogen (nukleobasa). Pada DNA, nukleobasa yang ditemukan adalah Adenina (A), Guanina (G), Sitosina (C) dan Timina (T). Nukleobasa yang terhubung dengan sebuah gugus gula disebut sebagai nukleosida, dan nukleosida yang terhubung dengan satu atau lebih gugus fosfat disebut sebagai nukleotida. Polimer yang terdiri dari nukleotida yang saling terhubung menjadi satu rantai disebut sebagai polinukleotida. Sehingga DNA termasuk pula ke dalam polinukleotida. | [
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1#6 | Asam deoksiribonukleat | Rantai punggung unting DNA terdiri dari gugus fosfat dan gula yang berselang-seling. Gula pada DNA adalah gula pentosa (berkarbon lima), yaitu 2-deoksiribosa. Dua gugus gula terhubung dengan fosfat melalui ikatan fosfodiester antara atom karbon ketiga pada cincin satu gula dan atom karbon kelima pada gula lainnya. Ikatan yang tidak simetris ini membuat DNA memiliki arah atau orientasi tertentu. Pada struktur heliks ganda, orientasi rantai nukleotida pada satu unting berlawanan dengan orientasi nukleotida unting lainnya. Hal ini disebut sebagai "antiparalel". Kedua ujung asimetris DNA disebut sebagai 5' (lima prima) dan 3' (tiga prima). Ujung 5' memiliki gugus fosfat terminus, sedangkan ujung 3' memiliki gugus hidroksi terminus. Salah satu perbedaan utama DNA dan RNA adalah gula penyusunnya, yakni gula 2-deoksiribosa pada DNA digantikan gula ribosa pada RNA. | [
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1#7 | Asam deoksiribonukleat | Dalam organisme hidup, DNA biasanya ditemukan dalam bentuk berpasangan dan terikat kuat. Dua unting DNA saling berpilin membentuk heliks ganda. Heliks ganda ini distabilisasi oleh dua gaya utama: ikatan hidrogen antar nukleotida dan interaksi tumpukan antar nukleobasa aromatik. Dalam lingkungan sel yang berair, ikatan π konjugasi antar basa nukleotida tersusun tegak lurus terhadap sumbu pilinan DNA. Hal ini meminimalisasi interaksi dengan cangkang solvasi, dan sehingganya menurunkan energi bebas Gibbs. | [
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1#8 | Asam deoksiribonukleat | Struktur DNA semua jenis spesies terdiri dari dua rantai heliks yang berpilin dengan jarak antar putaran heliks 34 Å (3,4 nanometer) dan jari-jari 10 Å (1.0 nanometer). Menurut kajian lainnya, ketika diukur menggunakan larutan tertentu, rantai DNA memiliki lebar 22-26 Å (2,2-2,6 nanometer) sedangkan satu satuan nukleotida memiliki panjang 33 Å (0,33 nm). Walaupun satuan nukleotida ini sangatlah kecil, polimer DNA dapat memiliki jutaan nukleotida yang terangkai seperti rantai. Misalnya, kromosom 1 yang merupakan kromosom terbesar pada manusia mengandung sekitar 220 juta pasangan basa. | [
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1#9 | Asam deoksiribonukleat | Nukleobasa diklasifikasikan ke dalam dua jenis: purina (A dan G) yang berupa fusi senyawa heterolingkar beranggota lima dengan senyawa heterolingkar beranggota enam, dan pirimidina (C dan T) yang berupa cincin beranggota enam. Pirimidina lainnya, urasil (U), biasanya menggantikan timina pada DNA. Perbedaan urasil dengan timina terletak pada ketiadaan gugus metil pada cincin urasil. Selain kelima nukleobasa tersebut, terdapat pula sejumlah besar analog asam nukleat buatan yang telah disintesis untuk mengkaji sifat-sifat asam nukleat dan digunakan dalam bioteknologi. | [
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1#10 | Asam deoksiribonukleat | Urasil biasanya tidak ditemukan dalam DNA (ditemukan dalam sel hanya sebagai produk uraian sitosina). Namun pada sejumlah bakteriofag– bakteriofag PBS1 dan PBS2 "Bacillus subtilis" dan bakteriofag piR1-37 "Yersinia"– timina telah digantikan oleh urasil. Fag lainnya - fag S6 Staphylococcus - juga telah diidentifikasi mempunyai urasil pada genomnya. | [
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1#11 | Asam deoksiribonukleat | Basa J (beta-d-glukopiranosiloksimetilurasil) yang merupakan bentuk modifikasi dari urasil juga dapat ditemukan pada sejumlah organisme: flagellata "Diplonema" dan "Euglena", dan seluruh organisme marga kinetoplastid Biosintesis basa J terjadi dalam dua tahap: pada tahap pertama, basa timina spesifik pada DNA diubah menjadi hidroksimetildeoksiuridina (HOMedU); pada tahap kedua HOMedU diglikosilasi menjadi basa J. Protein-protein yang mengikat basa J ini juga telah berhasil diidentifikasi. Protein-protein ini tampaknya merupakan kerabat jauh dari onkogen Tet1 yang terlibat dalam patogenesis leukimia myeloid akut. Basa J tampaknya bekerja sebagai sinyal terminasi untuk RNA polimerase II. | [
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1#12 | Asam deoksiribonukleat | Pada struktur heliks ganda DNA, terdapat ruang antar unting DNA yang juga berbentuk alur heliks. Ruang kosong ini bersebelahan dengan pasangan basa dan merupakan tapak ikatan yang potensial. Dikarenakan kedua unting DNA tidak berposisi secara simetris satu sama lainnya, alur yang dihasilkan jugalah tidak berukuran sama. Satu alur yang disebut alur mayor, memiliki lebar 22 Å, sedangkan alur lainnya yang disebut alur minor, memiliki lebar 12 Å. Lebarnya alur mayor berarti bahwa tepi-tepi basa nukleotida dapat lebih mudah diakses melalui alur mayor daripada melalui alur minor. Akibatnya, protein-protein seperti faktor-faktor transkripsi yang mengikat pada urutan basa tertentu biasanya melakukan kontak dengan basa melalui alur mayor. Situasi ini dapat bervariasi pada konformasi DNA yang tak lazim dalam sel, walaupun alur mayor dan minor selalu dinamai demikian untuk menrefleksikan perbedaan ukuran yang terlihat apabila DNA dipuntir balik menjadi bentuk lazim B. | [
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1#13 | Asam deoksiribonukleat | Pada heliks ganda DNA, tiap jenis nukleobasa pada satu unting DNA berikatan hanya dengan satu jenis nukleobasa dari unting DNA lainnya. Hal ini disebut sebagai pemasangan basa komplementer. Purina akan membentuk ikatan hidrogen dengan pirimidina; adenina berikatan dengan timina dalam dua ikatan hidrogen, dan sitosina berikatan dengan guanina dalam tiga ikatan hidrogen. Susunan dua nukleotida ini disebut sebagai satu pasangan basa. Karena ikatan hidrogen tidak bersifat kovalen, ia dapat putuskan dan digabung kembali relatif mudah. Kedua unting DNA dalam heliks ganda oleh karenanya dapat ditarik terbuka seperti zipper, baik melalui gaya mekanika maupun temperatur tinggi. Karena pasangan basa ini bersifat komplementer, semua informasi pada urutan unting ganda heliks DNA terduplikasi pada tiap unting. Hal ini sangat penting dalam replikasi DNA. Interaksi reversible dan spesifik antara pasangan basa komplementer sangat kritikal terhadap keseluruhan fungsi DNA dalam makhluk hidup. | [
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1#14 | Asam deoksiribonukleat | Sebagaimana telah disebutkan di atas, kebanyakan molekul DNA ditemukan dalam keadaan unting ganda yang berikatan secara non-kovalen dan berbentuk heliks. Struktur unting ganda ini (dsDNA, "double-stranded DNA") utamanya distabilkan oleh interaksi tumpukan basa intra-unting. Interaksi yang terkuat ada pada tumpukan G dengan C. Kedua unting tersebut dapat dipisahkan menjadi dua molekul DNA unting tunggal (ssDNA, "single-stranded DNA") melalui proses yang dinamakan "peleburan" DNA. Peleburan terjadi pada temperatur tinggi, kadar garam yang reandah, dan nilai pH yang tinggi (DNA juga melebur pada nilai pH rendah, tetapi dikarenakan DNA tidak stabil akibat depurinasi asam, peleburan pH rendah jarang digunakan). | [
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1#15 | Asam deoksiribonukleat | Stabilitas dsDNA tidak hanya bergantung pada kandungan GC (% pasangan basa G,C) DNA, namun juga tergantung pada urutan basa (tumpukan basa) dan panjang molekul DNA tersebut (molekul yang lebih panjang lebih stabil). Oleh sebab itu, kekuatan ikatan antar dua unting DNA ditentukan oleh persentase pasangan basa GC dan keseluruhan panjang heliks ganda DNA. Heliks DNA yang panjang dengan kandungan GC yang tinggi memiliki interaksi antar-unting yang lebih kuat; sebaliknya heliks DNA yang pendek dengan kandungan AT yang tinggi memiliki interaksi antar-unting yang lebih lemah. Dalam proses biologis, bagian heliks ganda DNA yang perlu dipisahkan dengan mudah seperti kotak Pribnow TATAAT pada beberapa promotor cenderung memiliki kandungan AT yang tinggi. | [
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1#16 | Asam deoksiribonukleat | Stabilitas DNA dapat diukur melalui berbagai cara; umumnya stabilitas DNA diukur berdasarkan temperatur lebur DNA (disebut juga nilai "T"), yakni temperatur di mana 50% molekul DNA unting ganda melebur menjadi molekul DNA unting tunggal. Temperatur lebur ini bergantung pada kekuatan ionik dan konsentrasi DNA. Ketika seluruh pasangan basa dalam heliks ganda DNA melebur, kedua unting DNA akan terpisah sebagai dua molekul yang independen. Unting-unting tunggal DNA ini tidak memiliki bentuk tunggal yang sama, walaupun beberapa konformasi lebih stabil daripada konformasi lainnya. | [
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1#17 | Asam deoksiribonukleat | Sebuah urutan sekuens DNA disebut sebagai "sense" apabila urutan basa DNA-nya sama dengan urutan kopi RNA duta yang ditranslasikan menjadi protein. Urutan pada unting komplementernya disebut sebagai urutan "antisense". Baik urutan sense dan antisense dapat ditemukan pada berbagai bagian unting DNA yang sama (kedua unting DNA dapat mengandung baik urutan sense maupun antisense). Pada prokariota dan eukariota, urutan RNA antisense juga diproduksi, namun fungsi RNA antisense ini tidaklah diketahui dengan jelas. RNA antisense diajukan terlibat dalam regulasi ekspresi gen melalui pemasangan basa RNA-RNA. | [
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1#18 | Asam deoksiribonukleat | Pada sebagian kecil urutan DNA prokariota dan eukariota, dan sebagian besar urutan DNA plasmid dan virus, perbedaan antara unting sense dan antisense menjadi kabur dikarenakan terdapatnya gen yang tumpang tindih. Dalam hal ini, beberapa urutan DNA memiliki tugas ganda, yakni menyandikan protein pertama ketika dibaca melalui salah satu unting, dan menyandikan protein kedua ketika dibaca dengan arah berlawanan melalui unting komplementernya. Pada bakteri, ketumpangtindihan ini kemungkinan terlibat dalam regulasi transkripsi gen. Sedangkan pada virus, gen yang tumpang tindih ini meningkatkan jumlah informasi yang dapat disandikan dalam genom virus yang berukuran kecil. | [
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1#19 | Asam deoksiribonukleat | DNA dapat dipuntir menjadi seperti tali melalui proses yang disebut pemilinan kumparan DNA. Pada kondisi "relaksasi", unting DNA biasanya akan mengitari sumbu heliks ganda setiap 10,4 pasangan basa. Namun jika DNA dipuntir, unting-untingnya dapat tergulung menjadi lebih rapat ataupun tergulung menjadi lebih longgar. Jika DNA dipuntir searah putaran heliks, basa-basa dalam unting DNA akan terikat lebih rapat. Hal ini dinamakan pemilinan kumparan positif. Sebaliknya jika DNA dipuntir berlawanan putaran heliks, basa-basa dalam unting DNA akan terlepas lebih mudah. Hal ini dinamakan pemilinan kumparan negatif. Secara alamiah, kebanyakan DNA memiliki bentuk pemilinan kumparan negatif yang disebabkan oleh enzim topoisomerase. Enzim ini juga diperlukan untuk melepaskan tegangan puntiran yang dialami DNA semasa proses transkripsi dan replikasi.
Terdapat banyak kemungkinan konformasi-konformasi DNA yang dapat kita temukan, di antaranya A-DNA, B-DNA, dan Z-DNA, walaupun hanya B-DNA dan Z-DNA saja yang telah diamati secara langsung pada organisme fungsional. Konformasi-konformasi yang diadopsi oleh DNA bergantung pada tingkat hidrasi DNA, urutan DNA, tingkat dan arah pilinan kumparan DNA, modifikasi kimiawi pada basa DNA, jenis dan konsentrasi ion-ion logam, maupun keberadaan poliamina dalam larutan. | [
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1#20 | Asam deoksiribonukleat | Replikasi merupakan proses pelipatgandaan DNA. Proses replikasi ini diperlukan ketika sel akan membelah diri. Pada setiap sel, kecuali sel gamet, pembelahan diri harus disertai dengan replikasi DNA supaya semua sel turunan memiliki informasi genetik yang sama. Pada dasarnya, proses replikasi memanfaatkan fakta bahwa DNA terdiri dari dua rantai dan rantai yang satu merupakan "konjugat" dari rantai pasangannya. Dengan kata lain, dengan mengetahui susunan satu rantai, maka susunan rantai pasangan dapat dengan mudah dibentuk. | [
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1#21 | Asam deoksiribonukleat | Ada beberapa teori yang mencoba menjelaskan bagaimana proses replikasi DNA ini terjadi. Salah satu teori yang paling populer menyatakan bahwa pada masing-masing DNA baru yang diperoleh pada akhir proses replikasi; satu rantai tunggal merupakan rantai DNA dari rantai DNA sebelumnya, sedangkan rantai pasangannya merupakan rantai yang baru disintesis. Rantai tunggal yang diperoleh dari DNA sebelumnya tersebut bertindak sebagai "cetakan" untuk membuat rantai pasangannya. | [
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1#22 | Asam deoksiribonukleat | Proses replikasi memerlukan protein atau enzim pembantu; salah satu yang terpenting dikenal dengan nama DNA polimerase, yang merupakan enzim pembantu pembentukan rantai DNA baru yang merupakan suatu polimer. Proses replikasi diawali dengan pembukaan untaian ganda DNA pada titik-titik tertentu di sepanjang rantai DNA. Proses pembukaan rantai DNA ini dibantu oleh enzim helikase yang dapat mengenali titik-titik tersebut, dan enzim girase yang mampu membuka pilinan rantai DNA. | [
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1#23 | Asam deoksiribonukleat | Setelah cukup ruang terbentuk akibat pembukaan untaian ganda ini, DNA polimerase masuk dan mengikat diri pada kedua rantai DNA yang sudah terbuka secara lokal tersebut. Proses pembukaan rantai ganda tersebut berlangsung disertai dengan pergeseran DNA polimerase mengikuti arah membukanya rantai ganda. Monomer DNA ditambahkan di kedua sisi rantai yang membuka setiap kali DNA polimerase bergeser. Hal ini berlanjut sampai seluruh rantai telah benar-benar terpisah. | [
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1#24 | Asam deoksiribonukleat | Proses replikasi DNA ini merupakan proses yang rumit namun teliti. Proses sintesis rantai DNA baru memiliki suatu mekanisme yang mencegah terjadinya kesalahan pemasukan monomer yang dapat berakibat fatal. Karena mekanisme inilah kemungkinan terjadinya kesalahan sintesis amatlah kecil. | [
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1#25 | Asam deoksiribonukleat | Ilmuwan forensik dapat menggunakan DNA yang terletak dalam darah, sperma, kulit, liur atau rambut yang tersisa di tempat kejadian kejahatan untuk mengidentifikasi kemungkinan tersangka, sebuah proses yang disebut fingerprinting genetika atau pemrofilan DNA ("DNA profiling"). Dalam pemrofilan DNA panjang relatif dari bagian DNA yang berulang seperti short tandem repeats dan minisatelit, dibandingkan. Pemrofilan DNA dikembangkan pada 1984 oleh genetikawan Inggris Alec Jeffreys dari Universitas Leicester, dan pertama kali digunakan untuk mendakwa Colin Pitchfork pada 1988 dalam kasus pembunuhan Enderby di Leicestershire, Inggris. | [
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1#26 | Asam deoksiribonukleat | Banyak yurisdiksi membutuhkan terdakwa dari kejahatan tertentu untuk menyediakan sebuah contoh DNA untuk dimasukkan ke dalam "database" komputer. Hal ini telah membantu investigator menyelesaikan kasus lama di mana pelanggar tidak diketahui dan hanya contoh DNA yang diperoleh dari tempat kejadian (terutama dalam kasus perkosaan antar orang tak dikenal). Metode ini adalah salah satu teknik paling tepercaya untuk mengidentifikasi seorang pelaku kejahatan, tetapi tidak selalu sempurna, misalnya bila tidak ada DNA yang dapat diperoleh, atau bila tempat kejadian terkontaminasi oleh DNA dari banyak orang. | [
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1#27 | Asam deoksiribonukleat | DNA memainkan peran penting dalam ilmu komputer, baik sebagai masalah riset dan sebagai sebuah cara komputasi. | [
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1#28 | Asam deoksiribonukleat | Riset dalam algoritme pencarian string, yang menemukan kejadian dari urutan huruf di dalam urutan huruf yang lebih besar, dimotivasi sebagian oleh riset DNA, dimana algoritme ini digunakan untuk mencari urutan tertentu dari nukleotida dalam sebuah urutan yang besar. Dalam aplikasi lainnya seperti editor text, bahkan algoritme sederhana untuk masalah ini biasanya mencukupi, tetapi urutan DNA menyebabkan algoritme-algoritme ini untuk menunjukkan sifat kasus-mendekati-terburuk dikarenakan jumlah kecil dari karakter yang berbeda. | [
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1#29 | Asam deoksiribonukleat | Teori database juga telah dipengaruhi oleh riset DNA, yang memiliki masalah khusus untuk menaruh dan memanipulasi urutan DNA. Database yang dikhususkan untuk riset DNA disebut database genomik, dam harus menangani sejumlah tantangan teknis yang unik yang dihubungkan dengan operasi pembandingan kira-kira, pembandingan urutan, mencari pola yang berulang, dan pencarian homologi. | [
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1#30 | Asam deoksiribonukleat | DNA pertama kali berhasil dimurnikan pada tahun 1868 oleh ilmuwan Swiss Friedrich Miescher di Tubingen, Jerman, yang menamainya "nuclein" berdasarkan lokasinya di dalam inti sel. Namun, penelitian terhadap peranan DNA di dalam sel baru dimulai pada awal abad 20, bersamaan dengan ditemukannya postulat genetika Mendel. DNA dan protein dianggap dua molekul yang paling memungkinkan sebagai pembawa sifat genetis berdasarkan teori tersebut. | [
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1#31 | Asam deoksiribonukleat | Dua eksperimen pada dekade 40-an membuktikan fungsi DNA sebagai materi genetik. Dalam penelitian oleh Avery dan rekan-rekannya, ekstrak dari sel bakteri yang satu gagal men-transform sel bakteri lainnya kecuali jika DNA dalam ekstrak dibiarkan utuh. Eksperimen yang dilakukan Hershey dan Chase membuktikan hal yang sama dengan menggunakan pencari jejak radioaktif (). | [
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1#32 | Asam deoksiribonukleat | Misteri yang belum terpecahkan ketika itu adalah: ""bagaimanakah struktur DNA sehingga ia mampu bertugas sebagai materi genetik"". Persoalan ini dijawab oleh Francis Crick dan koleganya James Watson berdasarkan hasil difraksi sinar X pada DNA oleh Maurice Wilkins dan Rosalind Franklin. | [
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1#33 | Asam deoksiribonukleat | Pada tahun 1953, James Watson dan Francis Crick mendefinisikan DNA sebagai polimer yang terdiri dari 4 basa dari asam nukleat, dua dari kelompok purina:adenina dan guanina; dan dua lainnya dari kelompok pirimidina:sitosina dan timina. Keempat nukleobasa tersebut terhubung dengan glukosa fosfat. | [
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1#34 | Asam deoksiribonukleat | Maurice Wilkins dan Rosalind Franklin menemukan bahwa molekul DNA berbentuk heliks yang berputar setiap 3,4 nm, sedangkan jarak antar molekul nukleobasa adalah 0,34 nm, hingga dapat ditentukan bahwa terdapat 10 molekul nukleobasa pada setiap putaran DNA. Setelah diketahui bahwa diameter heliks DNA sekitar 2 nm, baru diketahui bahwa DNA terdiri bukan dari 1 rantai, melainkan 2 rantai heliks. | [
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1#35 | Asam deoksiribonukleat | Crick, Watson, dan Wilkins mendapatkan hadiah Nobel Kedokteran pada 1962 atas penemuan ini. Franklin, karena sudah wafat pada waktu itu, tidak dapat dianugerahi hadiah ini. | [
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1#36 | Asam deoksiribonukleat | Konfirmasi akhir mekanisme replikasi DNA dilakukan lewat percobaan Meselson-Stahl yang dilakukan tahun 1958. | [
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3#0 | Anwar Sadat | Jenderal Besar Mohammed Anwar Al Sadat (; ) adalah seorang tentara dan politikus Mesir. Ia menjabat sebagai Presiden ketiga Mesir pada periode 15 Oktober 1970 hingga terbunuhnya pada 6 Oktober 1981. Oleh dunia Barat ia dianggap sebagai orang yang sangat berpengaruh di Mesir dan di Timur Tengah dalam sejarah modern. | [
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3#1 | Anwar Sadat | Sadat dilahirkan di Mit Abu Al-Kum, Al-Minufiyah, Mesir, dalam sebuah keluarga Mesir-sudan yang miskin, dengan 12 saudara laki-laki dan perempuan. Ayahnya adalah seorang Mesir, sementara ibunya orang Sudan. Ia lulus dari Akademi Militer Kerajaan di Kairo pada 1938 dan ditempatkan di Korps Isyarat. Ia bergabung dengan Gerakan Perwira Bebas, yang bertekad untuk membebaskan Mesir dari kekuasaan Britania Raya. | [
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3#2 | Anwar Sadat | Pada Perang Dunia II ia dipenjarakan oleh Britania atas usaha-usahanya untuk mendapatkan bantuan dari Kekuatan Poros dalam mengusir pasukan-pasukan pendudukan Britania. Ia ikut serta dalam kudeta 1952 yang menggulingkan Raja Farouk II. Ketika revolusi meletus, ia diperintahkan mengambil alih jaringan radio dan mengumumkan pecahnya revolusi kepada rakyat Mesir. | [
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] |
3#3 | Anwar Sadat | Pada 1964, setelah memegang berbagai jabatan dalam pemerintahan Mesir, ia dipilih oleh Presiden Gamal Abdel Nasser untuk menjabat sebagai Wakil Presiden. Ia menduduki jabatan itu hingga 1966, dan sekali lagi dari 1969 hingga 1970. | [
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3#4 | Anwar Sadat | Setelah Nasser meninggal, Anwar Sadat dilantik menjadi Presiden. | [
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3#5 | Anwar Sadat | Pada tahun 1973, Anwar Sadat, bersama-sama dengan Hafez Al Assad, Syria, memimpin Mesir dalam Perang Yom Kippur melawan Israel, untuk merebut kembali semenanjung Sinai, yang dicaplok oleh Israel ketika Krisis Terusan Suez 1956 dan Perang Enam Hari.Meskipun dalam pertempuran ini masih dipertentangkan pihak menang ataupun kalah, serta hasil Perjanjian Camp David yang menetapkan Sinai kembali ketangan Mesir, keberhasilan Anwar Sadat menaikan moral rakyat Mesir dan Dunia Arab serta mengadakan untuk perjanjian damai beberapa tahun berikutnya. | [
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3#6 | Anwar Sadat | Perjanjian damai Camp David yang diprakarsai Jimmy Carter dan Henry Kissinger memang mengembalikan wilayah Mesir yang sebelumnya direbut oleh Israel pada perang 1967. Namun tidak mengembalikan Dataran Tinggi Golan yang direbut Israel kepada Syria pada perang 1967. Meski secara politik, perang Yom Kippur atau Perang Ramadhan 1973 itu menguntungkan dunia Arab, masalah Palestina dan Jerusalem terutama Jerusalem Timur yang direbut Israel pada perang 1967 masih mengganjal bahkan beberapa kalangan mengatakan dilupakan. Hal ini membuat kemarahan dari kalangan PLO, kaum fundamentalis dan pergerakan Islam dan kalangan Palestina serta dunia Arab, terutama setelah kunjungannya ke Jerussalem atas undangan Manachem Begin. | [
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3#7 | Anwar Sadat | Pada tahun 1977, Anwar Sadat mengadakan kunjungan ke Jerusalem atas undangan Perdana Menteri Israel, Menachem Begin yang merupakan awal dari perundingan perdamaian antara Israel dan Mesir. Pada tahun 1978, terciptalah Perjanjian Damai Camp David, yang mana Anwar Sadat dan Menachem Begin menerima Hadiah Nobel Perdamaian. Bagaimanapun tindakan ini ditentang hebat oleh dunia Arab. Banyak yang percaya bahwa hanya dengan ancaman militer dapat memaksa Israel berunding mengenai Palestina, dan Perjanjian Damai Camp David menepikan Mesir yang dianggap kekuatan militer di dunia Arab yang signifikan disamping Syria dan Irak pada saat itu. | [
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3#8 | Anwar Sadat | Pada September 1981, Anwar Sadat mengenakan tindakan represif kepada organisasi pergerakan Islam yang diaggapnya fundamentalis, termasuk kumpulan pelajar, dan organisasi Koptik, yang dianggapnya dapat mengganggu stabilitas nasional Mesir, dengan mengadakan tindakan penangkapan dan penahanan menyebabkan dia dikecam diseluruh dunia di atas pelanggaran HAM dalam tindakannya itu. | [
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3#9 | Anwar Sadat | Pada 6 Oktober 1981, Presiden Anwar Sadat tewas ditembak dalam sebuah parade militer oleh anggota tentara anggota Jihad Islam. Ini merupakan organisasi muslim Mesir berhaluan keras yang menentang perjanjian damai Mesir dengan Israel. Tindakan represif anggota Jihad Islam terlihat dalam peristiwa September. Anwar Sadat kemudian digantikan oleh Wakil Presiden Hosni Mubarak. | [
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4#0 | Azhar Mansor | Datuk Azhar Mansor adalah orang Malaysia pertama yang berhasil mengelilingi dunia dengan berlayar. Mulanya ia merencanakan untuk melakukan pelayaran tersebut dalam waktu kurang dari 100 hari, tetapi ia menghadapi masalah dengan tiang layar yang patah ketika berada di Cape Horn, Amerika Selatan. Namun Datuk Azhar Mansor akhirnya berhasil menempuh seluruh pelayarannya dalam waktu 190 hari. Kegigihannya adalah contoh keunggulan Semangat Malaysia Baru. Datuk Azhar Mansor yang berusia 41 tahun ketika berangkat pada 2 Februari 1999 itu belayar dengan kapal layarnya bernama Jalur Gemilang yang berharga RM2 juta. | [
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4#1 | Azhar Mansor | Pelayaran Datuk Azhar meliputi 21,600 mil laut. Pada 11 Agustus 1999, Datuk Azhar Mansor tiba dengan selamat di Awana Porto Malai Resort, Langkawi dan disambut sebagai seorang pahlawan oleh rakyat Malaysia dipimpin oleh Perdana Menteri Malaysia. Kepahlawanan Datuk Azhar Mansor adalah sesuatu yang dapat dibanggakan oleh setiap rakyat Malaysia. Usaha dan kegigihannya dalam mengarungi badai dan ombak adalah sumbangannya sebagai anak merdeka yang tidak gentar menanggung penderitaan rakyat yang hidup pada masa sebelum kemerdekaan. | [
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5#0 | Arkeologi | Arkeologi atau ilmu kepurbakalaan berasal dari bahasa Indonesia , "archaeo" yang berarti "kuno" dan "logos", "ilmu". Nama alternatif arkeologi adalah ilmu sejarah kebudayaan material. Arkeologi adalah ilmu yang mempelajari kebudayaan (manusia) masa lalu melalui kajian sistematis atas data bendawi yang ditinggalkan. Kajian sistematis meliputi penemuan, dokumentasi, analisis, dan interpretasi data berupa artefak (budaya bendawi, seperti kapak batu dan bangunan candi) dan ekofak (benda lingkungan, seperti batuan, rupa muka bumi, dan fosil) maupun fitur (artefaktual yang tidak dapat dilepaskan dari tempatnya (situs arkeologi). Teknik penelitian yang khas adalah penggalian (ekskavasi) arkeologis, meskipun survei juga mendapatkan porsi yang cukup besar. Arkeolog adalah sebutan untuk para sarjana, praktisi, atau ahli di bidang arkeologi. | [
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5#1 | Arkeologi | Tujuan arkeologi beragam dan menjadi perdebatan yang panjang. Di antaranya adalah yang disebut dengan paradigma arkeologi, yaitu menyusun sejarah kebudayaan, memahami perilaku manusia, serta mengerti proses perubahan budaya. Karena bertujuan untuk memahami budaya manusia, maka ilmu ini termasuk ke dalam kelompok ilmu humaniora. Meskipun demikian, terdapat berbagai ilmu bantu yang digunakan, antara lain sejarah, antropologi, geologi (dengan ilmu tentang lapisan pembentuk bumi yang menjadi acuan relatif umur suatu temuan arkeologis), geografi, arsitektur, paleoantropologi dan bioantropologi, fisika (antara lain dengan karbon c-14 untuk mendapatkan pertanggalan mutlak), ilmu metalurgi (untuk mendapatkan unsur-unsur suatu benda logam), serta filologi (mempelajari naskah lama). | [
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5#2 | Arkeologi | Arkeologi pada masa sekarang merangkumi berbagai bidang yang berkait. Sebagai contoh, penemuan mayat yang dikubur akan menarik minat pakar dari berbagai bidang untuk mengkaji tentang pakaian dan jenis bahan digunakan, bentuk keramik dan cara penyebaran, kepercayaan melalui apa yang dikebumikan bersama mayat tersebut, pakar kimia yang mampu menentukan usia galian melalui cara seperti metode pengukuran karbon 14. Sedangkan pakar genetik yang ingin mengetahui pergerakan perpindahan manusia purba, meneliti DNAnya. | [
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5#3 | Arkeologi | Secara khusus, arkeologi mempelajari budaya masa silam, yang sudah berusia tua, baik pada masa prasejarah (sebelum dikenal tulisan), maupun pada masa sejarah (ketika terdapat bukti-bukti tertulis). Pada perkembangannya, arkeologi juga dapat mempelajari budaya masa kini, sebagaimana dipopulerkan dalam kajian budaya bendawi modern ("modern material culture"). | [
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5#4 | Arkeologi | Karena bergantung pada benda-benda peninggalan masa lalu, maka arkeologi sangat membutuhkan kelestarian benda-benda tersebut sebagai sumber data. Oleh karena itu, kemudian dikembangkan disiplin lain, yaitu pengelolaan sumberdaya arkeologi ("Archaeological Resources Management"), atau lebih luas lagi adalah pengelolaan sumberdaya budaya ("CRM, Culture Resources Management"). | [
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5#5 | Arkeologi | Di Indonesia, perkembangan arkeologi dimulai dari lembaga-lembaga yang bergerak di bidang kebudayaan, seperti Koninklijk Bataviaasch Genootschap van Kunsten en Wetenschappen yang kemudian di Jakarta mendirikan museum tertua, sekarang menjadi Museum Nasional Indonesia. Lembaga pemerintah pada masa Kolonial yang bergerak di bidang arkeologi adalah Oudheidkundige Dienst yang banyak membuat survei dan pemugaran atas bangunan-bangunan purbakala terutama candi. Pada masa Kemerdekaan, lembaga tersebut menjadi Dinas Purbakala hingga berkembang sekarang menjadi berbagai lembaga seperti Balai Pelestarian Peninggalan Purbakala dan Balai Arkeologi yang tersebar di daerah-daerah dan Direktorat Purbakala serta Pusat Penelitian dan Pengembangan Arkeologi Nasional di Jakarta. Di samping itu, terdapat 6 (enam) perguruan tinggi yang membuka jurusan arkeologi untuk mendidik tenaga sarjana di bidang arkeologi. Perguruan-perguruan tinggi tersebut adalah Universitas Indonesia (Departemen Arkeologi, Fakultas Ilmu Pengetahuan Budaya), Universitas Gadjah Mada (Departemen Arkeologi, Fakultas Ilmu Budaya), Universitas Hasanuddin (Departemen Arkeologi, Fakultas Ilmu Budaya), Universitas Udayana (Jurusan Arkeologi, Fakultas Sastra dan Budaya), Universitas Jambi (Jurusan Arkeologi, Fakultas Ilmu Budaya), dan Universitas Haluoleo (Jurusan Arkeologi, Fakultas Ilmu Budaya). | [
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5#6 | Arkeologi | Ahli arkeologi Indonesia terhimpun dalam Ikatan Ahli Arkeologi Indonesia atau IAAI. Tokoh-tokoh arkeologi Indonesia yang terkenal antara lain adalah R. Soekmono yang mengepalai pemugaran Candi Borobudur, dan R.P. Soejono, yang merupakan pendiri dan ketua Ikatan Ahli Arkeologi Indonesia pertama dan mantan kepala Pusat Penelitian Arkeologi Nasional. | [
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5#7 | Arkeologi | Disiplin Arkeologi Indonesia masih secara kuat diwarnai dengan pembagian kronologis, yaitu periode Prasejarah, periode Klasik (zaman Hindu-Buddha), periode Islam, serta periode Kolonial. Oleh karena itu, dalam arkeologi Indonesia dikenal spesialisasi menurut periode, yaitu Arkeologi Prasejarah, Arkeologi Klasik, Arkeologi Islam, serta Arkeologi Kolonial. Satu keistimewaan dari arkeologi Indonesia adalah masuknya disiplin Epigrafi, yang menekuni pembacaan prasasti kuno. Pada perkembangan sekarang telah berkembang minat-minat khusus seperti etnoarkeologi, arkeologi bawah air, dan arkeometri. Terdapat pula sub-disiplin yang berkembang karena persinggungan dengan ilmu lain, seperti Arkeologi Lingkungan atau Arkeologi Ekologi, Arkeologi Ekonomi, Arkeologi Seni, Arkeologi Demografi, dan Arkeologi Arsitektur. | [
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6#0 | Antropologi | Antropologi adalah ilmu tentang manusia. Antropologi berasal dari kata Yunani άνθρωπος (baca: "anthropos") yang berarti "manusia" atau "orang", dan "logos" yang berarti "wacana" (dalam pengertian "bernalar", "berakal") atau secara etimologis antropologi berarti ilmu yang mempelajari manusia. Dalam melakukan kajian terhadap manusia, antropologi mengedepankan dua konsep penting yaitu: holistik dan komparatif. Karena itu kajian antropologi sangat memperhatikan aspek sejarah dan penjelasan menyeluruh untuk menggambarkan manusia melalui pengetahuan ilmu sosial ilmu hayati (alam), dan juga humaniora. | [
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6#1 | Antropologi | Antropologi bertujuan untuk lebih memahami dan mengapresiasi manusia sebagai entitas biologis homo sapiens dan makhluk sosial dalam kerangka kerja yang interdisipliner dan komprehensif. Oleh karena itu, antropologi menggunakan teori evolusi biologi dalam memberikan arti dan fakta sejarah dalam menjelaskan perjalanan umat manusia di bumi sejak awal kemunculannya. Antropologi juga menggunakan kajian lintas-budaya (Inggris "cross-cultural") dalam menekankan dan menjelaskan perbedaan antara kelompok-kelompok manusia dalam perspektif material budaya, perilaku sosial, bahasa, dan pandangan hidup ("worldview"). | [
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] |
6#2 | Antropologi | Dengan orientasinya yang holistik, antropologi dibagi menjadi empat cabang ilmu yang saling berkaitan, yaitu: antropologi biologi, antropologi sosial budaya, arkeologi, dan linguistik. Keempat cabang tersebut memiliki kajian-kajian konsentrasi tersendiri dalam kekhususan akademik dan penelitian ilmiah, dengan topik yang unik dan metode penelitian yang berbeda-beda. | [
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6#3 | Antropologi | Antropologi lahir atau berawal dari ketertarikan orang-orang Eropa pada ciri-ciri fisik, adat istiadat, dan budaya etnis-etnis lain yang berbeda dari masyarakat yang dikenal di Eropa. Pada saat itu kajian antropologi lebih memusatkan pada penduduk yang merupakan masyarakat tunggal, tunggal dalam arti kesatuan masyarakat yang tinggal di suatu kawasan geografis yang sama, memiliki ciri fisik dan bahasa yang digunakan serupa, serta cara hidup yang sama. Namun demikian dalam perkembangannya, ilmu antropologi kemudian tidak lagi hanya mempelajari kelompok manusia tunggal yang mendiami suatu wilayah geografis yang sama. Kajian-kajian antropologi mengenai isu-isu migrasi misalnya kemudian melahirkan penelitian-penelitian etnografis multi-situs. Hal ini terjadi karena dalam perkembangannya, pergerakan manusia baik dalam satu kawasan regional tertentu hingga dalam cakupan global adalah fenomena yang semakin umum terjadi.Antropologi merupakan disiplin ilmu yang luas di mana humaniora, sosial, dan ilmu pengetahuan alam digabung dalam menjelaskan apa itu manusia dan artinya menjadi manusia. Antropologi dibangun berdasarkan pengetahuan dari ilmu alam, termasuk penemuan tentang asal usul dan evolusi Homo sapiens, ciri-ciri fisik manusia, perilaku manusia, variasi di antara berbagai kelompok manusia, bagaimana masa lalu evolusi Homo sapiens telah memengaruhi organisasi dan budaya sosial. Serta dari ilmu-ilmu sosial, antropologi memelajari organisasi hubungan manusia sosial dan budaya, sistem keturunan dan hubungan kekerabatan, spiritualitas dan religi, lembaga, konflik sosial, dan lain-lain. Antropologi awal berasal dari Yunani klasik dan Persia yang memelajari dan mencoba untuk memahami keragaman budaya yang dapat diamati. Pada saat ini, antropologi (akhir abad ke-20) telah menjadi sentral dalam pengembangan beberapa bidang interdisipliner baru seperti ilmu kognitif, studi globalisasi, genetik, dan berbagai penelitian etnis. | [
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6#4 | Antropologi | Secara garis besar antropologi terdiri dari: | [
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6#5 | Antropologi | Antropologi Biologi atau juga disebut Antropologi Fisik merupakan cabang ilmu antropologi yang memelajari manusia dan primata bukan manusia (non-human primates) dalam arti biologis, evolusi, dan demografi. Antropologi Biologi/Fisik memfokuskan pada faktor biologis dan sosial yang memengaruhi (atau yang menentukan) evolusi manusia dan primata lainnya, yang menghasilkan, mempertahankan, atau mengubah variasi genetik dan fisiologisnya pada saat ini. | [
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6#6 | Antropologi | Antropologi Biologi dibagi lagi menjadi beberapa cabang ilmu, diantaranya yaitu: | [
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6#7 | Antropologi | Antropologi sosial merupakan studi yang mempelajari hubungan antara orang-orang dan kelompok. Sementara Antropologi Budaya merupakan studi komparasi bagaimana orang-orang memahami dunia di sekitar mereka dengan cara yang berbeda-beda. Antropologi Sosial berkaitan erat dengan sosiologi dan sejarah yang bertujuan mencari pemahaman struktur sosial dari suatu kelompok sosial yang berbeda seperti subkultur, etnik, dan kelompok minoritas. Antropologi Budaya lebih berhubungan dengan filsafat, literatur atau sastra, dan seni tentang bagaimana suatu kebudayaan memengaruhi pengalaman seseorang (diri sendiri) dan kelompok, memberikan kontribusi untuk pemahaman yang lebih lengkap terhadap pengetahuan, adat istiadat, dan pranata masyarakat. Dalam praktiknya tidak ada perbedaan yang sangat mencolok antara Antropologi Sosial dan Antropologi Budaya, dan bahkan sering saling tumpang tindih di antara keduanya.Seperti halnya sosiologi, antropologi sebagai sebuah ilmu juga mengalami tahapan-tahapan dalam perkembangannya. | [
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6#8 | Antropologi | Koentjaraninggrat menyusun perkembangan ilmu Antropologi menjadi empat fase sebagai berikut: | [
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6#9 | Antropologi | Sekitar abad ke-15-16, bangsa-bangsa di Eropa mulai berlomba-lomba untuk menjelajahi dunia. Mulai dari Afrika, Amerika, Asia, hingga ke Australia. Dalam penjelajahannya mereka banyak menemukan hal-hal baru. Mereka juga banyak menjumpai suku-suku yang asing bagi mereka. Kisah-kisah petualangan dan penemuan mereka kemudian mereka catat di buku harian ataupun jurnal perjalanan. Mereka mencatat segala sesuatu yang berhubungan dengan suku-suku asing tersebut. Mulai dari ciri-ciri fisik, kebudayaan, susunan masyarakat, atau bahasa dari suku tersebut. Bahan-bahan yang berisi tentang deskripsi suku asing tersebut kemudian dikenal dengan bahan etnografi atau deskripsi tentang bangsa-bangsa. | [
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6#10 | Antropologi | Bahan etnografi itu menarik perhatian pelajar-pelajar di Eropa. Kemudian, pada permulaan abad ke-19 perhatian bangsa Eropa terhadap bahan-bahan etnografi suku luar Eropa dari sudut pandang ilmiah, menjadi sangat besar. Karena itu, timbul usaha-usaha untuk mengintegrasikan seluruh himpunan bahan etnografi. | [
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6#11 | Antropologi | Pada fase ini, bahan-bahan etnografi tersebut telah disusun menjadi karangan-karangan berdasarkan cara berpikir evolusi masyarakat pada saat itu. masyarakat dan kebudayaan berevolusi secara perlahan-lahan dan dalam jangka waktu yang lama. Mereka menganggap bangsa-bangsa selain Eropa sebagai bangsa-bangsa primitif yang tertinggal, dan menganggap Eropa sebagai bangsa yang tinggi kebudayaannya | [
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6#12 | Antropologi | Pada fase ini, Antopologi bertujuan akademis, mereka mempelajari masyarakat dan kebudayaan primitif dengan maksud untuk memperoleh pemahaman tentang tingkat-tingkat sejarah penyebaran kebudayaan manusia. | [
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6#13 | Antropologi | Pada fase ini, negara-negara di Eropa berlomba-lomba membangun koloni di benua lain seperti Asia, Amerika, Australia dan Afrika. Dalam rangka membangun koloni-koloni tersebut, muncul berbagai kendala seperti serangan dari bangsa asli, pemberontakan-pemberontakan, cuaca yang kurang cocok bagi bangsa Eropa serta hambatan-hambatan lain. Dalam menghadapinya, pemerintahan kolonial negara Eropa berusaha mencari-cari kelemahan suku asli untuk kemudian menaklukannya. Untuk itulah mereka mulai mempelajari bahan-bahan etnografi tentang suku-suku bangsa di luar Eropa, mempelajari kebudayaan dan kebiasaannya, untuk kepentingan pemerintah kolonial. | [
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6#14 | Antropologi | Pada fase ini, Antropologi berkembang secara pesat. Kebudayaan-kebudayaan suku bangsa asli yang di jajah bangsa Eropa, mulai hilang akibat terpengaruh kebudayaan bangsa Eropa. | [
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6#15 | Antropologi | Pada masa ini pula terjadi sebuah perang besar di Eropa, Perang Dunia II. Perang ini membawa banyak perubahan dalam kehidupan manusia dan membawa sebagian besar negara-negara di dunia kepada kehancuran total. Kehancuran itu menghasilkan kemiskinan, kesenjangan sosial, dan kesengsaraan yang tak berujung. | [
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6#16 | Antropologi | Namun pada saat itu juga, muncul semangat nasionalisme bangsa-bangsa yang dijajah Eropa untuk keluar dari belenggu penjajahan. Sebagian dari bangsa-bangsa tersebut berhasil mereka. Namun banyak masyarakatnya yang masih memendam dendam terhadap bangsa Eropa yang telah menjajah mereka selama bertahun-tahun. | [
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6#17 | Antropologi | Proses-proses perubahan tersebut menyebabkan perhatian ilmu antropologi tidak lagi ditujukan kepada penduduk pedesaan di luar Eropa, tetapi juga kepada suku bangsa di daerah pedalaman Eropa seperti suku bangsa Soami, Flam dan Lapp. | [
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7#0 | Analisis leksikal | Analisis leksikal (bahasa Inggris: "lexical analysis") adalah sebuah proses yang mendahului parsing sebuah rangkaian karakter. Ia menerima masukan serangkaian karakter (seperti dalam dokumen plain-text atau source code) dan menghasilkan deretan simbol yang masing-masing dinamakan token; proses parsing akan lebih mudah dilakukan bila inputnya sudah berupa token. | [
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7#1 | Analisis leksikal | Analisis leksikal terdiri dari dua tahap. Tahap pertama adalah pemindaian (scanning); scanner biasanya dibuat berdasarkan prinsip Finite State Machine ("mesin dengan jumlah keadaan terbatas"). Pada tahap ini, scanner akan membaca input karakter-ke-karakter, mengubah keadaannya sendiri berdasarkan karakter yang tengah dibaca. Setiap kondisi final (input dianggap valid) akan dicatat, bersama dengan lokasi input. Pada akhirnya scanner akan menemui keadaan penolakan, yang tidak akan berubah dengan input karakter apapun. Deteksi rekursi semacam ini akan mengakhiri proses pemindaian dan memindahkan keadaan scanner ke keadaan final terakhir, dan karenanya menyimpan informasi jenis dan besar lexeme valid yang terpanjang di dalam input. | [
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7#2 | Analisis leksikal | Namun lexeme tersebut belum punya nilai semantik apapun; pemberian nilai semantik pada setiap unit leksikal adalah tugas dari evaluator yang memeriksa semua karakter setiap lexeme dan memberinya nilai tertentu. Saat sebuah lexeme telah memiliki informasi mengenai tipe dan nilainya, ia dapat secara valid disebut sebagai token. | [
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7#3 | Analisis leksikal | Analisis leksikal membuat pekerjaan parser jadi lebih mudah; daripada membangun nama setiap fungsi dan variabel dari karakter-karakter yang menyusunnya, dengan analisis leksikal, parser cukup hanya berurusan dengan sekumpulan token dan nilai sintaksis masing-masing. Terlepas dari efisiensi pemrograman yang dapat dicapai dengan penggunaannya, proses kerja analisis leksikal yang membaca lebih dari sekali setiap karakter dari input yang diberikan menjadikan penganalisa leksikal sebagai sub-sistem yang paling intensif melakukan komputasi, terutama bila digunakan dalam sebuah kompilator. | [
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8#0 | Bahasa Indonesia | Bahasa Indonesia adalah bentuk standar bahasa Melayu yang dijadikan sebagai bahasa resmi Republik Indonesia dan bahasa persatuan bangsa Indonesia. Bahasa Indonesia diresmikan penggunaannya setelah Proklamasi Kemerdekaan Indonesia, tepatnya sehari sesudahnya, bersamaan dengan mulai berlakunya konstitusi. Di Timor Leste, bahasa Indonesia berstatus sebagai bahasa kerja. | [
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8#1 | Bahasa Indonesia | Dari sudut pandang linguistik, bahasa Indonesia adalah salah satu dari banyak ragam bahasa Melayu. Dasar yang dipakai adalah bahasa Melayu Riau (wilayah Kepulauan Riau sekarang) dari abad ke-19. Dalam perkembangannya ia mengalami perubahan akibat penggunaanya sebagai bahasa kerja di lingkungan administrasi kolonial dan berbagai proses pembakuan sejak awal abad ke-20. Penamaan "bahasa Indonesia" diawali sejak dicanangkannya Sumpah Pemuda, 28 Oktober 1928, untuk menghindari kesan "imperialisme bahasa" apabila nama bahasa Melayu tetap digunakan. Proses ini menyebabkan berbedanya bahasa Indonesia saat ini dari varian bahasa Melayu yang digunakan di Riau maupun Semenanjung Malaya. Hingga saat ini, bahasa Indonesia merupakan bahasa yang hidup, yang terus menghasilkan kata-kata baru, baik melalui penciptaan maupun penyerapan dari bahasa daerah dan bahasa asing. | [
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8#2 | Bahasa Indonesia | Meskipun dipahami dan dituturkan oleh lebih dari 90% warga Indonesia, bahasa Indonesia bukanlah bahasa ibu bagi kebanyakan penuturnya. Sebagian besar warga Indonesia menggunakan salah satu dari 748 bahasa yang ada di Indonesia sebagai bahasa ibu. Penutur bahasa Indonesia kerap kali menggunakan versi sehari-hari (kolokial) dan/atau mencampuradukkan dengan dialek Melayu lainnya atau bahasa ibunya. Meskipun demikian, bahasa Indonesia digunakan sangat luas di perguruan-perguruan, di media massa, sastra, perangkat lunak, surat-menyurat resmi, dan berbagai forum publik lainnya, sehingga dapatlah dikatakan bahwa bahasa Indonesia digunakan oleh semua warga Indonesia. | [
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8#3 | Bahasa Indonesia | Fonologi dan tata bahasa bahasa Indonesia dianggap relatif mudah. Menurut sebagian orang, dasar-dasar yang penting untuk komunikasi dasar dapat dipelajari hanya dalam kurun waktu beberapa minggu. | [
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8#4 | Bahasa Indonesia | Bahasa Indonesia adalah varian bahasa Melayu, sebuah bahasa Austronesia dari cabang bahasa-bahasa Sunda-Sulawesi, yang digunakan sebagai "lingua franca" di Nusantara kemungkinan sejak abad-abad awal penanggalan modern. | [
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8#5 | Bahasa Indonesia | Aksara pertama dalam bahasa Melayu atau Jawi ditemukan di pesisir tenggara Pulau Sumatera, mengindikasikan bahwa bahasa ini menyebar ke berbagai tempat di Nusantara dari wilayah ini, berkat penggunaannya oleh Kerajaan Sriwijaya yang menguasai jalur perdagangan. Istilah Melayu atau sebutan bagi wilayahnya sebagai Malaya sendiri berasal dari Kerajaan Malayu yang bertempat di Batang Hari, Jambi. | [
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8#6 | Bahasa Indonesia | Istilah Melayu atau Malayu berasal dari Kerajaan Malayu, sebuah kerajaan Hindu-Budha pada abad ke-7 di hulu sungai Batanghari, Jambi di pulau Sumatera, jadi secara geografis semula hanya mengacu kepada wilayah kerajaan tersebut yang merupakan sebagian dari wilayah pulau Sumatera. Dalam perkembangannya pemakaian istilah Melayu mencakup wilayah geografis yang lebih luas dari wilayah Kerajaan Malayu tersebut, mencakup negeri-negeri di pulau Sumatera sehingga pulau tersebut disebut juga Bumi Melayu seperti disebutkan dalam Kakawin Nagarakretagama. | [
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8#7 | Bahasa Indonesia | Ibukota Kerajaan Melayu semakin mundur ke pedalaman karena serangan Sriwijaya dan masyarakatnya diaspora keluar Bumi Melayu, belakangan masyarakat pendukungnya yang mundur ke pedalaman berasimilasi ke dalam masyarakat Minangkabau menjadi klan Malayu (suku Melayu Minangkabau) yang merupakan salah satu marga di Sumatera Barat. Sriwijaya berpengaruh luas hingga ke Filipina membawa penyebaran Bahasa Melayu semakin meluas, tampak dalam prasasti Keping Tembaga Laguna. | [
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8#8 | Bahasa Indonesia | Bahasa Melayu kuno yang berkembang di Bumi Melayu tersebut berlogat "o" seperti Melayu Jambi, Minangkabau, Kerinci, Palembang dan Bengkulu. Semenanjung Malaka dalam Nagarakretagama disebut Hujung Medini artinya Semenanjung Medini. | [
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8#9 | Bahasa Indonesia | Dalam perkembangannya orang Melayu migrasi ke Semenanjung Malaysia (= Hujung Medini) dan lebih banyak lagi pada masa perkembangan kerajaan-kerajaan Islam yang pusat mandalanya adalah Kesultanan Malaka, istilah Melayu bergeser kepada Semenanjung Malaka (= Semenanjung Malaysia) yang akhirnya disebut Semenanjung Melayu atau Tanah Melayu. Tetapi nyatalah bahwa istilah Melayu itui berasal dari Indonesia. Bahasa Melayu yang berkembang di sekitar daerah Semenanjung Malaka berlogat "e". | [
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8#10 | Bahasa Indonesia | Kesultanan Malaka dimusnahkan oleh Portugis tahun 1512 sehingga penduduknya diaspora sampai ke kawasan timur kepulauan Nusantara. Bahasa Melayu Purba sendiri diduga berasal dari pulau Kalimantan, jadi diduga pemakai bahasa Melayu ini bukan penduduk asli Sumatera tetapi dari pulau Kalimantan. Suku Dayak yang diduga memiliki hubungan dengan suku Melayu kuno di Sumatera misalnya Dayak Salako, Dayak Kanayatn (Kendayan), dan Dayak Iban yang semuanya berlogat "a" seperti bahasa Melayu Baku. Penduduk asli Sumatera sebelumnya kedatangan pemakai bahasa Melayu tersebut adalah nenek moyang suku Nias dan suku Mentawai. | [
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8#11 | Bahasa Indonesia | Dalam perkembangannya istilah Melayu kemudian mengalami perluasan makna, sehingga muncul istilah Kepulauan Melayu untuk menamakan kepulauan Nusantara. | [
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8#12 | Bahasa Indonesia | Secara sudut pandang historis juga dipakai sebagai nama bangsa yang menjadi nenek moyang penduduk kepulauan Nusantara, yang dikenal sebagai rumpun Indo-Melayu terdiri Proto Melayu (Melayu Tua/Melayu Polinesia) dan Deutero Melayu (Melayu Muda). Setelah mengalami kurun masa yang panjang sampai dengan kedatangan dan perkembangannya agama Islam, suku Melayu sebagai etnik mengalami penyempitan makna menjadi sebuah etnoreligius (Muslim) yang sebenarnya di dalamnya juga telah mengalami amalgamasi dari beberapa unsur etnis. | [
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8#13 | Bahasa Indonesia | M. Muhar Omtatok, seorang Seniman, Budayawan dan Sejarahwan menjelaskan sebagai berikut: "Melayu secara puak (etnis, suku), bukan dilihat dari faktor genekologi seperti kebanyakan puak-puak lain. Di Malaysia, tetap mengaku berpuak Melayu walau moyang mereka berpuak Jawa, Mandailing, Bugis, Keling dan lainnya. Beberapa tempat di Sumatera Utara, ada beberapa Komunitas keturunan Batak yang mengaku Orang Kampong - Puak Melayu | [
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8#14 | Bahasa Indonesia | Kerajaan Sriwijaya dari abad ke-7 Masehi diketahui memakai bahasa Melayu (sebagai bahasa Melayu Kuno) sebagai bahasa kenegaraan. Lima prasasti kuno yang ditemukan di Sumatera bagian selatan peninggalan kerajaan itu menggunakan bahasa Melayu yang bertaburan kata-kata pinjaman dari bahasa Sanskerta, suatu bahasa Indo-Eropa dari cabang Indo-Iran. Jangkauan penggunaan bahasa ini diketahui cukup luas, karena ditemukan pula dokumen-dokumen dari abad berikutnya di Pulau Jawa dan Pulau Luzon. Kata-kata seperti "samudra, istri, raja, putra, kepala, kawin", dan "kaca" masuk pada periode hingga abad ke-15 Masehi. | [
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8#15 | Bahasa Indonesia | Pada abad ke-15 berkembang bentuk yang dianggap sebagai bahasa Melayu Klasik ("classical Malay" atau "medieval Malay"). Bentuk ini dipakai oleh Kesultanan Melaka, yang perkembangannya kelak disebut sebagai "bahasa Melayu Tinggi". Penggunaannya terbatas di kalangan keluarga kerajaan di sekitar Sumatera, Jawa, dan Semenanjung Malaya. Laporan Portugis, misalnya oleh Tome Pires, menyebutkan adanya bahasa yang dipahami oleh semua pedagang di wilayah Sumatera dan Jawa. Magellan dilaporkan memiliki budak dari Nusantara yang menjadi juru bahasa di wilayah itu. Ciri paling menonjol dalam ragam sejarah ini adalah mulai masuknya kata-kata pinjaman dari bahasa Arab dan bahasa Parsi, sebagai akibat dari penyebaran agama Islam yang mulai masuk sejak abad ke-12. Kata-kata bahasa Arab seperti masjid, kalbu, kitab, kursi, selamat, dan kertas, serta kata-kata Parsi seperti anggur, cambuk, dewan, saudagar, tamasya, dan tembakau masuk pada periode ini. Proses penyerapan dari bahasa Arab terus berlangsung hingga sekarang. | [
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8#16 | Bahasa Indonesia | Kedatangan pedagang Portugis, diikuti oleh Belanda, Spanyol, dan Inggris meningkatkan informasi dan mengubah kebiasaan masyarakat pengguna bahasa Melayu. Bahasa Portugis banyak memperkaya kata-kata untuk kebiasaan Eropa dalam kehidupan sehari-hari, seperti gereja, sepatu, sabun, meja, bola, bolu, dan jendela. Bahasa Belanda terutama banyak memberi pengayaan di bidang administrasi, kegiatan resmi (misalnya dalam upacara dan kemiliteran), dan teknologi hingga awal abad ke-20. Kata-kata seperti asbak, polisi, kulkas, knalpot, dan stempel adalah pinjaman dari bahasa ini. | [
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8#17 | Bahasa Indonesia | Bahasa yang dipakai pendatang dari Cina juga lambat laun dipakai oleh penutur bahasa Melayu, akibat kontak di antara mereka yang mulai intensif di bawah penjajahan Belanda. Sudah dapat diduga, kata-kata Tionghoa yang masuk biasanya berkaitan dengan perniagaan dan keperluan sehari-hari, seperti pisau, tauge, tahu, loteng, teko, tauke, dan cukong. | [
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8#18 | Bahasa Indonesia | Jan Huyghen van Linschoten pada abad ke-17 dan Alfred Russel Wallace pada abad ke-19 menyatakan bahwa bahasa orang Melayu/Melaka dianggap sebagai bahasa yang paling penting di "dunia timur".
Luasnya penggunaan bahasa Melayu ini melahirkan berbagai varian lokal dan temporal. Bahasa perdagangan menggunakan bahasa Melayu di berbagai pelabuhan Nusantara bercampur dengan bahasa Portugis, bahasa Tionghoa, maupun bahasa setempat. Terjadi proses pidginisasi di beberapa kota pelabuhan di kawasan timur Nusantara, misalnya di Manado, Ambon, dan Kupang. Orang-orang Tionghoa di Semarang dan Surabaya juga menggunakan varian bahasa Melayu pidgin. Terdapat pula bahasa Melayu Tionghoa di Batavia. Varian yang terakhir ini malah dipakai sebagai bahasa pengantar bagi beberapa surat kabar pertama berbahasa Melayu (sejak akhir abad ke-19). Varian-varian lokal ini secara umum dinamakan "bahasa Melayu Pasar" oleh para peneliti bahasa. | [
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8#19 | Bahasa Indonesia | Terobosan penting terjadi ketika pada pertengahan abad ke-19 Raja Ali Haji dari istana Riau-Johor (pecahan Kesultanan Melaka) menulis kamus ekabahasa untuk bahasa Melayu. Sejak saat itu dapat dikatakan bahwa bahasa ini adalah bahasa yang "full-fledged", sama tinggi dengan bahasa-bahasa internasional pada masa itu, karena memiliki kaidah dan dokumentasi kata yang terdefinisi dengan jelas. | [
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8#20 | Bahasa Indonesia | Hingga akhir abad ke-19 dapat dikatakan terdapat paling sedikit dua kelompok bahasa Melayu yang dikenal masyarakat Nusantara: bahasa Melayu Pasar yang kolokial dan tidak baku serta bahasa Melayu Tinggi yang terbatas pemakaiannya tetapi memiliki standar. Bahasa ini dapat dikatakan sebagai "lingua franca", tetapi kebanyakan berstatus sebagai bahasa kedua atau ketiga. | [
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