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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | ormal structure of the hemoglobin molecule) may cause both. In any case, hemoglobin deficiency decreases blood oxygen-carrying capacity. Hemoglobin deficiency is, in general, strictly distinguished from hypoxemia, defined as decreased partial pressure of oxygen in blood, although both are causes of hypoxia (insufficient oxygen supply to tissues).
Other common causes of low hemoglobin include loss of blood, nutritional deficiency, bone marrow problems, chemotherapy, kidney failure, or abnormal hemoglobin (s | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | uch as that of sickle-cell disease).
The ability of each hemoglobin molecule to carry oxygen is normally modified by altered blood pH or CO2, causing an altered oxygen–hemoglobin dissociation curve. However, it can also be pathologically altered in, e.g., carbon monoxide poisoning.
Decrease of hemoglobin, with or without an absolute decrease of red blood cells, leads to symptoms of anemia. Anemia has many different causes, although iron deficiency and its resultant iron deficiency anemia are the most comm | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | on causes in the Western world. As absence of iron decreases heme synthesis, red blood cells in iron deficiency anemia are hypochromic (lacking the red hemoglobin pigment) and microcytic (smaller than normal). Other anemias are rarer. In hemolysis (accelerated breakdown of red blood cells), associated jaundice is caused by the hemoglobin metabolite bilirubin, and the circulating hemoglobin can cause kidney failure.
Some mutations in the globin chain are associated with the hemoglobinopathies, such as sickl | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | e-cell disease and thalassemia. Other mutations, as discussed at the beginning of the article, are benign and are referred to merely as hemoglobin variants.
There is a group of genetic disorders, known as the porphyrias that are characterized by errors in metabolic pathways of heme synthesis. King George III of the United Kingdom was probably the most famous porphyria sufferer.
To a small extent, hemoglobin A slowly combines with glucose at the terminal valine (an alpha aminoacid) of each β chain. The res | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | ulting molecule is often referred to as Hb A1c, a glycated hemoglobin. The binding of glucose to amino acids in the hemoglobin takes place spontaneously (without the help of an enzyme) in many proteins, and is not known to serve a useful purpose. However, as the concentration of glucose in the blood increases, the percentage of Hb A that turns into Hb A1c increases. In diabetics whose glucose usually runs high, the percent Hb A1c also runs high. Because of the slow rate of Hb A combination with glucose, the | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | Hb A1c percentage reflects a weighted average of blood glucose levels over the lifetime of red cells, which is approximately 120 days. The levels of glycated hemoglobin are therefore measured in order to monitor the long-term control of the chronic disease of type 2 diabetes mellitus (T2DM). Poor control of T2DM results in high levels of glycated hemoglobin in the red blood cells. The normal reference range is approximately 4.0–5.9%. Though difficult to obtain, values less than 7% are recommended for peopl | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | e with T2DM. Levels greater than 9% are associated with poor control of the glycated hemoglobin, and levels greater than 12% are associated with very poor control. Diabetics who keep their glycated hemoglobin levels close to 7% have a much better chance of avoiding the complications that may accompany diabetes (than those whose levels are 8% or higher). In addition, increased glycated of hemoglobin increases its affinity for oxygen, therefore preventing its release at the tissue and inducing a level of hypo | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | xia in extreme cases.
Elevated levels of hemoglobin are associated with increased numbers or sizes of red blood cells, called polycythemia. This elevation may be caused by congenital heart disease, cor pulmonale, pulmonary fibrosis, too much erythropoietin, or polycythemia vera. High hemoglobin levels may also be caused by exposure to high altitudes, smoking, dehydration (artificially by concentrating Hb), advanced lung disease and certain tumors.
A recent study done in Pondicherry, India, shows its impor | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | tance in coronary artery disease.
Diagnostic uses
Hemoglobin concentration measurement is among the most commonly performed blood tests, usually as part of a complete blood count. For example, it is typically tested before or after blood donation. Results are reported in g/L, g/dL or mol/L. 1 g/dL equals about 0.6206 mmol/L, although the latter units are not used as often due to uncertainty regarding the polymeric state of the molecule. This conversion factor, using the single globin unit molecular weight | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | of 16,000 Da, is more common for hemoglobin concentration in blood. For MCHC (mean corpuscular hemoglobin concentration) the conversion factor 0.155, which uses the tetramer weight of 64,500 Da, is more common. Normal levels are:
Men: 13.8 to 18.0 g/dL (138 to 180 g/L, or 8.56 to 11.17 mmol/L)
Women: 12.1 to 15.1 g/dL (121 to 151 g/L, or 7.51 to 9.37 mmol/L)
Children: 11 to 16 g/dL (110 to 160 g/L, or 6.83 to 9.93 mmol/L)
Pregnant women: 11 to 14 g/dL (110 to 140 g/L, or 6.83 to 8.69 mmol/L) (9.5 to 15 | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | usual value during pregnancy)
Normal values of hemoglobin in the 1st and 3rd trimesters of pregnant women must be at least 11 g/dL and at least 10.5 g/dL during the 2nd trimester.
Dehydration or hyperhydration can greatly influence measured hemoglobin levels. Albumin can indicate hydration status.
If the concentration is below normal, this is called anemia. Anemias are classified by the size of red blood cells, the cells that contain hemoglobin in vertebrates. The anemia is called "microcytic" if red ce | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | lls are small, "macrocytic" if they are large, and "normocytic" otherwise.
Hematocrit, the proportion of blood volume occupied by red blood cells, is typically about three times the hemoglobin concentration measured in g/dL. For example, if the hemoglobin is measured at 17 g/dL, that compares with a hematocrit of 51%.
Laboratory hemoglobin test methods require a blood sample (arterial, venous, or capillary) and analysis on hematology analyzer and CO-oximeter. Additionally, a new noninvasive hemoglobin (Sp | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | Hb) test method called Pulse CO-Oximetry is also available with comparable accuracy to invasive methods.
Concentrations of oxy- and deoxyhemoglobin can be measured continuously, regionally and noninvasively using NIRS. NIRS can be used both on the head and on muscles. This technique is often used for research in e.g. elite sports training, ergonomics, rehabilitation, patient monitoring, neonatal research, functional brain monitoring, brain–computer interface, urology (bladder contraction), neurology (Neuro | [
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] |
13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | vascular coupling) and more.
Long-term control of blood sugar concentration can be measured by the concentration of Hb A1c. Measuring it directly would require many samples because blood sugar levels vary widely through the day. Hb A1c is the product of the irreversible reaction of hemoglobin A with glucose. A higher glucose concentration results in more Hb A1c. Because the reaction is slow, the Hb A1c proportion represents glucose level in blood averaged over the half-life of red blood cells, is typically | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | ~120 days. An Hb A1c proportion of 6.0% or less show good long-term glucose control, while values above 7.0% are elevated. This test is especially useful for diabetics.
The functional magnetic resonance imaging (fMRI) machine uses the signal from deoxyhemoglobin, which is sensitive to magnetic fields since it is paramagnetic. Combined measurement with NIRS shows good correlation with both the oxy- and deoxyhemoglobin signal compared to the BOLD signal.
Athletic tracking and self tracking uses
Hemoglobin | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | can be tracked noninvasively, to build an individual data set tracking the hemoconcentration and hemodilution effects of daily activities for better understanding of sports performance and training. Athletes are often concerned about endurance and intensity of exercise. The sensor uses light-emitting diodes that emit red and infrared light through the tissue to a light detector, which then sends a signal to a processor to calculate the absorption of light by the hemoglobin protein.
This sensor is similar | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | to a pulse oximeter, which consists of a small sensing device that clips to the finger.
Analogues in non-vertebrate organisms
A variety of oxygen-transport and -binding proteins exist in organisms throughout the animal and plant kingdoms. Organisms including bacteria, protozoans, and fungi all have hemoglobin-like proteins whose known and predicted roles include the reversible binding of gaseous ligands. Since many of these proteins contain globins and the heme moiety (iron in a flat porphyrin support), t | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | hey are often called hemoglobins, even if their overall tertiary structure is very different from that of vertebrate hemoglobin. In particular, the distinction of "myoglobin" and hemoglobin in lower animals is often impossible, because some of these organisms do not contain muscles. Or, they may have a recognizable separate circulatory system but not one that deals with oxygen transport (for example, many insects and other arthropods). In all these groups, heme/globin-containing molecules (even monomeric gl | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | obin ones) that deal with gas-binding are referred to as oxyhemoglobins. In addition to dealing with transport and sensing of oxygen, they may also deal with NO, CO2, sulfide compounds, and even O2 scavenging in environments that must be anaerobic. They may even deal with detoxification of chlorinated materials in a way analogous to heme-containing P450 enzymes and peroxidases.
The structure of hemoglobins varies across species. Hemoglobin occurs in all kingdoms of organisms, but not in all organisms. Prim | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | itive species such as bacteria, protozoa, algae, and plants often have single-globin hemoglobins. Many nematode worms, molluscs, and crustaceans contain very large multisubunit molecules, much larger than those in vertebrates. In particular, chimeric hemoglobins found in fungi and giant annelids may contain both globin and other types of proteins.
One of the most striking occurrences and uses of hemoglobin in organisms is in the giant tube worm (Riftia pachyptila, also called Vestimentifera), which can rea | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | ch 2.4 meters length and populates ocean volcanic vents. Instead of a digestive tract, these worms contain a population of bacteria constituting half the organism's weight. The bacteria oxidize H2S from the vent with O2 from the water to produce energy to make food from H2O and CO2. The worms' upper end is a deep-red fan-like structure ("plume"), which extends into the water and absorbs H2S and O2 for the bacteria, and CO2 for use as synthetic raw material similar to photosynthetic plants. The structures ar | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | e bright red due to their content of several extraordinarily complex hemoglobins that have up to 144 globin chains, each including associated heme structures. These hemoglobins are remarkable for being able to carry oxygen in the presence of sulfide, and even to carry sulfide, without being completely "poisoned" or inhibited by it as hemoglobins in most other species are.
Other oxygen-binding proteins
Myoglobin Found in the muscle tissue of many vertebrates, including humans, it gives muscle tissue a dist | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | inct red or dark gray color. It is very similar to hemoglobin in structure and sequence, but is not a tetramer; instead, it is a monomer that lacks cooperative binding. It is used to store oxygen rather than transport it.
Hemocyanin The second most common oxygen-transporting protein found in nature, it is found in the blood of many arthropods and molluscs. Uses copper prosthetic groups instead of iron heme groups and is blue in color when oxygenated.
Hemerythrin Some marine invertebrates and a few species | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | of annelid use this iron-containing non-heme protein to carry oxygen in their blood. Appears pink/violet when oxygenated, clear when not.
Chlorocruorin Found in many annelids, it is very similar to erythrocruorin, but the heme group is significantly different in structure. Appears green when deoxygenated and red when oxygenated.
Vanabins Also known as vanadium chromagens, they are found in the blood of sea squirts. They were once hypothesized to use the metal vanadium as an oxygen binding prosthetic grou | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | p. However, although they do contain vanadium by preference, they apparently bind little oxygen, and thus have some other function, which has not been elucidated (sea squirts also contain some hemoglobin). They may act as toxins.
Erythrocruorin Found in many annelids, including earthworms, it is a giant free-floating blood protein containing many dozens—possibly hundreds—of iron- and heme-bearing protein subunits bound together into a single protein complex with a molecular mass greater than 3.5 million da | [
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] |
13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | ltons.
Pinnaglobin Only seen in the mollusc Pinna nobilis. Brown manganese-based porphyrin protein.
Leghemoglobin In leguminous plants, such as alfalfa or soybeans, the nitrogen fixing bacteria in the roots are protected from oxygen by this iron heme containing oxygen-binding protein. The specific enzyme protected is nitrogenase, which is unable to reduce nitrogen gas in the presence of free oxygen.
Coboglobin A synthetic cobalt-based porphyrin. Coboprotein would appear colorless when oxygenated, but yel | [
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] |
13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | low when in veins.
Presence in nonerythroid cells
Some nonerythroid cells (i.e., cells other than the red blood cell line) contain hemoglobin. In the brain, these include the A9 dopaminergic neurons in the substantia nigra, astrocytes in the cerebral cortex and hippocampus, and in all mature oligodendrocytes. It has been suggested that brain hemoglobin in these cells may enable the "storage of oxygen to provide a homeostatic mechanism in anoxic conditions, which is especially important for A9 DA neurons th | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | at have an elevated metabolism with a high requirement for energy production". It has been noted further that "A9 dopaminergic neurons may be at particular risk since in addition to their high mitochondrial activity they are under intense oxidative stress caused by the production of hydrogen peroxide via autoxidation and/or monoamine oxidase (MAO)-mediated deamination of dopamine and the subsequent reaction of accessible ferrous iron to generate highly toxic hydroxyl radicals". This may explain the risk of | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | these cells for degeneration in Parkinson's disease. The hemoglobin-derived iron in these cells is not the cause of the post-mortem darkness of these cells (origin of the Latin name, substantia nigra), but rather is due to neuromelanin.
Outside the brain, hemoglobin has non-oxygen-carrying functions as an antioxidant and a regulator of iron metabolism in macrophages, alveolar cells, and mesangial cells in the kidney.
In history, art and music
Historically, an association between the color of blood and ru | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | st occurs in the association of the planet Mars, with the Roman god of war, since the planet is an orange-red, which reminded the ancients of blood. Although the color of the planet is due to iron compounds in combination with oxygen in the Martian soil, it is a common misconception that the iron in hemoglobin and its oxides gives blood its red color. The color is actually due to the porphyrin moiety of hemoglobin to which the iron is bound, not the iron itself, although the ligation and redox state of the | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | iron can influence the pi to pi* or n to pi* electronic transitions of the porphyrin and hence its optical characteristics.
Artist Julian Voss-Andreae created a sculpture called Heart of Steel (Hemoglobin) in 2005, based on the protein's backbone. The sculpture was made from glass and weathering steel. The intentional rusting of the initially shiny work of art mirrors hemoglobin's fundamental chemical reaction of oxygen binding to iron.
Montreal artist Nicolas Baier created Lustre (Hémoglobine), a sculptu | [
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13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | re in stainless steel that shows the structure of the hemoglobin molecule. It is displayed in the atrium of McGill University Health Centre's research centre in Montreal. The sculpture measures about 10 metres × 10 metres × 10 metres.
See also
Carbaminohemoglobin (Hb associated with )
Carboxyhemoglobin (Hb associated with CO)
Chlorophyll (Mg heme)
Complete Blood Count
Delta globin
Hemoglobinometer
Hemoprotein
Methemoglobin (ferric Hb, or ferrihemoglobin)
Oxyhemoglobin (with diatomic oxygen, color | [
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] |
13483 | https://en.wikipedia.org/wiki/Hemoglobin | Hemoglobin | ed blood-red)
Vaska's complex – iridium organometallic complex notable for its ability to bind to O2 reversibly
Tegillarca granosa
References
Further reading
External links
National Anemia Action Council – anemia.org
New hemoglobin type causes mock diagnosis with pulse oxymeters
Animation of hemoglobin: from deoxy to oxy form
Related Questions:
Ideal Hemoglobin Level in Pregnancy
How low can hemoglobin go before death
Hemoglobin Level
Hemoglobins
Equilibrium chemistry
Respiratory physiology | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | England became inhabited more than 800,000 years ago, as the discovery of stone tools and footprints at Happisburgh in Norfolk has indicated. The earliest evidence for early modern humans in Northwestern Europe, a jawbone discovered in Devon at Kents Cavern in 1927, was re-dated in 2011 to between 41,000 and 44,000 years old. Continuous human habitation in England dates to around 13,000 years ago (see Creswellian), at the end of the Last Glacial Period. The region has numerous remains from the Mesolithic, N | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | eolithic and Bronze Age, such as Stonehenge and Avebury. In the Iron Age, all of Britain south of the Firth of Forth, was inhabited by the Celtic people known as the Britons, including some Belgic tribes (e.g. the Atrebates, the Catuvellauni, the Trinovantes, etc.) in the south east. In AD 43 the Roman conquest of Britain began; the Romans maintained control of their province of Britannia until the early 5th century.
The end of Roman rule in Britain facilitated the Anglo-Saxon settlement of Britain, which | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | historians often regard as the origin of England and of the English people. The Anglo-Saxons, a collection of various Germanic peoples, established several kingdoms that became the primary powers in present-day England and parts of southern Scotland. They introduced the Old English language, which largely displaced the previous British language. The Anglo-Saxons warred with British successor states in western Britain and the Hen Ogledd (Old North; the Brittonic-speaking parts of northern Britain), as well a | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | s with each other. Raids by Vikings became frequent after about AD 800, and the Norsemen settled in large parts of what is now England. During this period, several rulers attempted to unite the various Anglo-Saxon kingdoms, an effort that led to the emergence of the Kingdom of England by the 10th century.
In 1066, a Norman expedition invaded and conquered England. The Norman dynasty, established by William the Conqueror, ruled England for over half a century before the period of succession crisis known as | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | the Anarchy (1135–1154). Following the Anarchy, England came under the rule of the House of Plantagenet, a dynasty which later inherited claims to the Kingdom of France. During this period, Magna Carta was signed. A succession crisis in France led to the Hundred Years' War (1337–1453), a series of conflicts involving the peoples of both nations. Following the Hundred Years' Wars, England became embroiled in its own succession wars. The Wars of the Roses pitted two branches of the House of Plantagenet agains | [
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] |
13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | t one another, the House of York and the House of Lancaster. The Lancastrian Henry Tudor ended the War of the Roses and established the Tudor dynasty in 1485.
Under the Tudors and the later Stuart dynasty, England became a colonial power. During the rule of the Stuarts, the English Civil War took place between the Parliamentarians and the Royalists, which resulted in the execution of King Charles I (1649) and the establishment of a series of republican governments—first, a Parliamentary republic known as t | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | he Commonwealth of England (1649–1653), then a military dictatorship under Oliver Cromwell known as the Protectorate (1653–1659). The Stuarts returned to the restored throne in 1660, though continued questions over religion and power resulted in the deposition of another Stuart king, James II, in the Glorious Revolution (1688). England, which had subsumed Wales in the 16th century under Henry VIII, united with Scotland in 1707 to form a new sovereign state called Great Britain. Following the Industrial Revo | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | lution, which started in England, Great Britain ruled a colonial Empire, the largest in recorded history. Following a process of decolonisation in the 20th century, mainly caused by the weakening of Great Britain's power in the two World Wars, almost all of the empire's overseas territories became independent countries. However, , its cultural impact remains widespread and deep in many of them.
Prehistory
Stone Age
The time from Britain's first inhabitation until the Last Glacial Maximum is known as the | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | Old Stone Age, or Palaeolithic era. Archaeological evidence indicates that what was to become England was colonised by humans long before the rest of the British Isles because of its more hospitable climate between and during the various glacial periods of the distant past. This earliest evidence, from Happisburgh in Norfolk, includes the oldest hominid footprints found outside Africa, and points to dates of more than 800,000 RCYBP. These earliest inhabitants were hunter-gatherers. Low sea-levels meant that | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | Britain was attached to the continent for much of this earliest period of history, and varying temperatures over tens of thousands of years meant that it was not always inhabited.
England has been continually inhabited since the last Ice Age ended around 9000 BC, the beginning of the Middle Stone Age, or Mesolithic era. Rising sea-levels cut off Britain from the continent for the last time around 6500 BC. The population by then was exclusively anatomically modern humans, and the evidence suggests that the | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ir societies were increasingly complex and they were manipulating their environment and prey in new ways, possibly selective burning of then omnipresent woodland to create clearings for herds to gather and then hunt them. Hunting was mainly done with simple projectile weapons such as javelin and possibly sling. Bow and arrow was known in Western Europe since at least 9000 BC. The climate continued to warm and the population probably rose.
The New Stone Age, or Neolithic era, began with the introduction of | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | farming, ultimately from the Middle East, around 4000 BC. It is not known whether this was caused by a substantial folk movement or native adoption of foreign practices or both. People began to lead a more settled lifestyle. Monumental collective tombs were built for the dead in the form of chambered cairns and long barrows. Towards the end of the period, other kinds of monumental stone alignments begin to appear, such as Stonehenge; their cosmic alignments show a preoccupation with the sky and planets. Fli | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | nt technology produced a number of highly artistic pieces as well as purely pragmatic. More extensive woodland clearance was done for fields and pastures. The Sweet Track in the Somerset Levels is one of the oldest timber trackways known in Northern Europe and among the oldest roads in the world, dated by dendrochronology to the winter of 3807–3806 BC; it too is thought to have been a primarily religious structure. Archaeological evidence from North Yorkshire indicates that salt was being manufactured there | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | in the Neolithic.
Later Prehistory
The Bronze Age began around 2500 BC with the appearance of bronze objects. This coincides with the appearance of the characteristic Beaker culture, which occurred primarily by folk movement. The Bronze Age saw a shift of emphasis from the communal to the individual, and the rise of increasingly powerful elites whose power came from their prowess as hunters and warriors and their controlling the flow of precious resources to manipulate tin and copper into high-status bro | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | nze objects such as swords and axes. Settlement became increasingly permanent and intensive. Towards the end of the Bronze Age, many examples of very fine metalwork began to be deposited in rivers, presumably for ritual reasons and perhaps reflecting a progressive change in emphasis from the sky to the earth, as a rising population put increasing pressure on the land. England largely became bound up with the Atlantic trade system, which created a cultural continuum over a large part of Western Europe. It is | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | possible that the Celtic languages developed or spread to England as part of this system; by the end of the Iron Age there is much evidence that they were spoken across all England and western parts of Britain.
The Iron Age is conventionally said to begin around 800 BC. The Atlantic system had by this time effectively collapsed, although England maintained contacts across the Channel with France, as the Hallstatt culture became widespread across the country. Its continuity suggests it was not accompanied | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | by substantial movement of population; crucially, only a single Hallstatt burial is known from Britain, and even here the evidence is inconclusive. On the whole, burials largely disappear across England, and the dead were disposed of in a way which is archaeologically invisible: excarnation is a widely cited possibility. Hillforts were known since the Late Bronze Age, but a huge number were constructed during 600–400 BC, particularly in the South, while after about 400 BC new forts were rarely built and man | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | y ceased to be regularly inhabited, while a few forts become more and more intensively occupied, suggesting a degree of regional centralisation. Around this time the earliest mentions of Britain appear in the annals of history. The first historical mention of the region is from the Massaliote Periplus, a sailing manual for merchants thought to date to the 6th century BC, and Pytheas of Massilia wrote of his voyage of discovery to the island around 325 BC. Both of these texts are now lost; although quoted by | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | later writers, not enough survives to inform the archaeological interpretation to any significant degree.
Contact with the continent was less than in the Bronze Age but still significant. Goods continued to move to England, with a possible hiatus around 350 to 150 BC. There were a few armed invasions of hordes of migrating Celts. There are two known invasions. Around 300 BC, a group from the Gaulish Parisii tribe apparently took over East Yorkshire, establishing the highly distinctive Arras culture. And f | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | rom around 150–100 BC, groups of Belgae began to control significant parts of the South. These invasions constituted movements of a few people who established themselves as a warrior elite atop existing native systems, rather than replacing them. The Belgic invasion was much larger than the Parisian settlement, but the continuity of pottery style shows that the native population remained in place. Yet, it was accompanied by significant socio-economic change. Proto-urban, or even urban settlements, known as | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | oppida, begin to eclipse the old hillforts, and an elite whose position is based on battle prowess and the ability to manipulate resources re-appears much more distinctly.
In 55 and 54 BC, Julius Caesar, as part of his campaigns in Gaul, invaded Britain and claimed to have scored a number of victories, but he never penetrated further than Hertfordshire and could not establish a province. However, his invasions mark a turning-point in British history. Control of trade, the flow of resources and prestige goo | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ds, became ever more important to the elites of Southern Britain; Rome steadily became the biggest player in all their dealings, as the provider of great wealth and patronage. In retrospect, a full-scale invasion and annexation was inevitable.
Genetic markers of Bell Beaker migration
According to Olalde et al. (2018), around 2500 BC Britain's Neolithic population was largely replaced by a population from North Continental Europe which belonged to the Bell Beaker culture, and was genetically related to the | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | Yamnaya people from the Pontic-Caspian Steppe. This population lacked genetic affinity to the Iberian Bell Beakers, where the Bell Beaker phenomenon originated, but appeared genetically to be an offshoot of the Corded Ware single grave people. While the migration of these Beaker peoples must have been accompanied by a language shift, the Celtic languages were probably introduced by later Celtic migrations.
Roman Britain
After Caesar's expeditions, the Romans began a serious and sustained attempt to conqu | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | er Britain in AD 43, at the behest of Emperor Claudius. They landed in Kent with four legions and defeated two armies led by the kings of the Catuvellauni tribe, Caratacus and Togodumnus, in battles at the Medway and the Thames. Togodumnus was killed, and Caratacus fled to Wales. The Roman force, led by Aulus Plautius, waited for Claudius to come and lead the final march on the Catuvellauni capital at Camulodunum (modern Colchester), before he returned to Rome for his triumph. The Catuvellauni held sway ove | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | r most of the southeastern corner of England; eleven local rulers surrendered, a number of client kingdoms were established, and the rest became a Roman province with Camulodunum as its capital. Over the next four years, the territory was consolidated and the future emperor Vespasian led a campaign into the Southwest where he subjugated two more tribes. By AD 54 the border had been pushed back to the Severn and the Trent, and campaigns were underway to subjugate Northern England and Wales.
But in AD 60, un | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | der the leadership of the warrior-queen Boudicca, the tribes rebelled against the Romans. At first, the rebels had great success. They burned Camulodunum, Londinium and Verulamium (modern-day Colchester, London and St. Albans respectively) to the ground. There is some archaeological evidence that the same happened at Winchester. The Second Legion Augusta, stationed at Exeter, refused to move for fear of revolt among the locals. Londinium governor Suetonius Paulinus evacuated the city before the rebels sacke | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | d and burned it; the fire was so hot that a ten-inch layer of melted red clay remains 15 feet below London's streets. In the end, the rebels were said to have killed 70,000 Romans and Roman sympathisers. Paulinus gathered what was left of the Roman army. In the decisive battle, 10,000 Romans faced nearly 100,000 warriors somewhere along the line of Watling Street, at the end of which Boudicca was utterly defeated. It was said that 80,000 rebels were killed, but only 400 Romans.
Over the next 20 years, the | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | borders expanded slightly, but the governor Agricola incorporated into the province the last pockets of independence in Wales and Northern England. He also led a campaign into Scotland which was recalled by Emperor Domitian. The border gradually formed along the Stanegate road in Northern England, solidified by Hadrian's Wall built in AD 138, despite temporary forays into Scotland.
The Romans and their culture stayed in charge for 350 years. Traces of their presence are ubiquitous throughout England.
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | o-Saxon period
Anglo-Saxon migrations
In the wake of the breakdown of Roman rule in Britain from the middle of the fourth century, present day England was progressively settled by Germanic groups. Collectively known as the Anglo-Saxons, these included Angles, Saxons, Jutes and Frisians. The Battle of Deorham was critical in establishing Anglo-Saxon rule in 577. Saxon mercenaries existed in Britain since before the late Roman period, but the main influx of population probably happened after the fifth centu | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ry. The precise nature of these invasions is not fully known; there are doubts about the legitimacy of historical accounts due to a lack of archaeological finds. Gildas' De Excidio et Conquestu Britanniae, composed in the 6th century, states that when the Roman army departed the Isle of Britannia in the 4th century AD, the indigenous Britons were invaded by Picts, their neighbours to the north (now Scotland) and the Scots (now Ireland). Britons invited the Saxons to the island to repel them but after they v | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | anquished the Scots and Picts, the Saxons turned against the Britons.
Seven kingdoms are traditionally identified as being established by these migrants. Three were clustered in the South east: Sussex, Kent and Essex. The Midlands were dominated by the kingdoms of Mercia and East Anglia. To the north was Northumbria which unified two earlier kingdoms, Bernicia and Deira. Other smaller kingdoms seem to have existed as well, such as Lindsey in what is now Lincolnshire, and the Hwicce in the southwest. Eventu | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ally, the kingdoms were dominated by Northumbria and Mercia in the 7th century, Mercia in the 8th century and then Wessex in the 9th century. Northumbria eventually extended its control north into Scotland and west into Wales. It also subdued Mercia whose first powerful King, Penda, was killed by Oswy in 655. Northumbria's power began to wane after 685 with the defeat and death of its king Aegfrith at the hands of the Picts. Mercian power reached its peak under the rule of Offa, who from 785 had influence o | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ver most of Anglo-Saxon England. Since Offa's death in 796, the supremacy of Wessex was established under Egbert who extended control west into Cornwall before defeating the Mercians at the Battle of Ellendun in 825. Four years later, he received submission and tribute from the Northumbrian king, Eanred.
Since so few contemporary sources exist, the events of the fifth and sixth centuries are difficult to ascertain. As such, the nature of the Anglo-Saxon settlements is debated by historians, archaeologists | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | and linguists. The traditional view, that the Anglo-Saxons drove the Romano-British inhabitants out of what is now England, was subject to reappraisal in the later twentieth century. One suggestion is that the invaders were smaller in number, drawn from an elite class of male warriors that gradually acculturated the natives.
An emerging view is that the scale of the Anglo-Saxon settlement varied across England, and that as such it cannot be described by any one process in particular. Mass migration and pop | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ulation shift seem to be most applicable in the core areas of settlement such as East Anglia and Lincolnshire, while in more peripheral areas to the northwest, much of the native population likely remained in place as the incomers took over as elites. In a study of place names in northeastern England and southern Scotland, Bethany Fox concluded that Anglian migrants settled in large numbers in river valleys, such as those of the Tyne and the Tweed, with the Britons in the less fertile hill country becoming | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | acculturated over a longer period. Fox interprets the process by which English came to dominate this region as "a synthesis of mass-migration and elite-takeover models."
Genetic markers of Anglo-Saxon migrations
Genetic testing has been used to find evidence of large scale immigration of Germanic peoples into England. Weale et al. (2002) found that English Y DNA data showed signs of a mass Anglo-Saxon immigration from the European continent, affecting 50%–100% of the male gene pool in central England. Thi | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | s was based on the similarity of the DNA collected from small English towns to that found in Friesland. A 2003 study by , with samples coming from larger towns, found a large variance in amounts of continental "Germanic" ancestry in different parts of England. In their study, such markers typically ranged from 20% and 45% in southern England, with East Anglia, the east Midlands, and Yorkshire having over 50%. North German and Danish genetic frequencies were indistinguishable, thus precluding any ability to | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | distinguish between the genetic influence of the Anglo-Saxon source populations and the later, and better documented, influx of Danish Vikings. The mean value of continental Germanic genetic input in this study was calculated at 54 percent.
In response to arguments, such as those of Stephen Oppenheimer and Bryan Sykes, that the similarity between English and continental Germanic DNA could have originated from earlier prehistoric migrations, researchers have begun to use data collected from ancient burials | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | to ascertain the level of Anglo-Saxon contribution to the modern English gene pool.
Two studies published in 2016, based on data collected from skeletons found in Iron Age, Roman and Anglo-Saxon era graves in Cambridgeshire and Yorkshire, concluded that the ancestry of the modern English population contains large contributions from both Anglo-Saxon migrants and Romano-British natives.
Heptarchy and Christianisation
Christianisation of Anglo-Saxon England began around 600 AD, influenced by Celtic Christia | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | nity from the northwest and the Roman Catholic Church from the southeast. Augustine, the first Archbishop of Canterbury, took office in 597. In 601, he baptised the first Christian Anglo-Saxon king, Æthelberht of Kent. The last pagan Anglo-Saxon king, Penda of Mercia, died in 655. The last pagan Jutish king, Arwald of the Isle of Wight was killed in 686. The Anglo-Saxon mission on the continent took off in the 8th century, leading to the Christianisation of practically all of the Frankish Empire by 800.
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | roughout the 7th and 8th centuries, power fluctuated between the larger kingdoms. Bede records Æthelberht of Kent as being dominant at the close of the 6th century, but power seems to have shifted northwards to the kingdom of Northumbria, which was formed from the amalgamation of Bernicia and Deira. Edwin of Northumbria probably held dominance over much of Britain, though Bede's Northumbrian bias should be kept in mind. Due to succession crises, Northumbrian hegemony was not constant, and Mercia remained a | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | very powerful kingdom, especially under Penda. Two defeats ended Northumbrian dominance: the Battle of the Trent in 679 against Mercia, and Nechtanesmere in 685 against the Picts.
The so-called "Mercian Supremacy" dominated the 8th century, though it was not constant. Aethelbald and Offa, the two most powerful kings, achieved high status; indeed, Offa was considered the overlord of south Britain by Charlemagne. His power is illustrated by the fact that he summoned the resources to build Offa's Dyke. Howeve | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | r, a rising Wessex, and challenges from smaller kingdoms, kept Mercian power in check, and by the early 9th century the "Mercian Supremacy" was over.
This period has been described as the Heptarchy, though this term has now fallen out of academic use. The term arose because the seven kingdoms of Northumbria, Mercia, Kent, East Anglia, Essex, Sussex and Wessex were the main polities of south Britain. Other small kingdoms were also politically important across this period: Hwicce, Magonsaete, Lindsey and Mid | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | dle Anglia.
Viking challenge and the rise of Wessex
The first recorded landing of Vikings took place in 787 in Dorsetshire, on the south-west coast. The first major attack in Britain was in 793 at Lindisfarne monastery as given by the Anglo-Saxon Chronicle. However, by then the Vikings were almost certainly well-established in Orkney and Shetland, and many other non-recorded raids probably occurred before this. Records do show the first Viking attack on Iona taking place in 794. The arrival of the Vikings | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | (in particular the Danish Great Heathen Army) upset the political and social geography of Britain and Ireland. In 867 Northumbria fell to the Danes; East Anglia fell in 869. Though Wessex managed to contain the Vikings by defeating them at Ashdown in 871, a second invading army landed, leaving the Saxons on a defensive footing. At much the same time, Æthelred, king of Wessex died and was succeeded by his younger brother Alfred. Alfred was immediately confronted with the task of defending Wessex against the | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | Danes. He spent the first five years of his reign paying the invaders off. In 878, Alfred's forces were overwhelmed at Chippenham in a surprise attack.
It was only now, with the independence of Wessex hanging by a thread, that Alfred emerged as a great king. In May 878 he led a force that defeated the Danes at Edington. The victory was so complete that the Danish leader, Guthrum, was forced to accept Christian baptism and withdraw from Mercia. Alfred then set about strengthening the defences of Wessex, bu | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ilding a new navy—60 vessels strong. Alfred's success bought Wessex and Mercia years of peace and sparked economic recovery in previously ravaged areas.
Alfred's success was sustained by his son Edward, whose decisive victories over the Danes in East Anglia in 910 and 911 were followed by a crushing victory at Tempsford in 917. These military gains allowed Edward to fully incorporate Mercia into his kingdom and add East Anglia to his conquests. Edward then set about reinforcing his northern borders against | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | the Danish kingdom of Northumbria. Edward's rapid conquest of the English kingdoms meant Wessex received homage from those that remained, including Gwynedd in Wales and Scotland. His dominance was reinforced by his son Æthelstan, who extended the borders of Wessex northward, in 927 conquering the Kingdom of York and leading a land and naval invasion of Scotland. These conquests led to his adopting the title 'King of the English' for the first time.
The dominance and independence of England was maintained | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | by the kings that followed. It was not until 978 and the accession of Æthelred the Unready that the Danish threat resurfaced. Two powerful Danish kings (Harold Bluetooth and later his son Sweyn) both launched devastating invasions of England. Anglo-Saxon forces were resoundingly defeated at Maldon in 991. More Danish attacks followed, and their victories were frequent. Æthelred's control over his nobles began to falter, and he grew increasingly desperate. His solution was to pay off the Danes: for almost 20 | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | years he paid increasingly large sums to the Danish nobles to keep them from English coasts. These payments, known as Danegelds, crippled the English economy.
Æthelred then made an alliance with Normandy in 1001 through marriage to the Duke's daughter Emma, in the hope of strengthening England. Then he made a great error: in 1002 he ordered the massacre of all the Danes in England. In response, Sweyn began a decade of devastating attacks on England. Northern England, with its sizable Danish population, si | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ded with Sweyn. By 1013, London, Oxford, and Winchester had fallen to the Danes. Æthelred fled to Normandy and Sweyn seized the throne. Sweyn suddenly died in 1014, and Æthelred returned to England, confronted by Sweyn's successor, Cnut. However, in 1016, Æthelred also suddenly died. Cnut swiftly defeated the remaining Saxons, killing Æthelred's son Edmund in the process. Cnut seized the throne, crowning himself King of England.
English unification
Alfred of Wessex died in 899 and was succeeded by his son | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | Edward the Elder. Edward, and his brother-in-law Æthelred of (what was left of) Mercia, began a programme of expansion, building forts and towns on an Alfredian model. On Æthelred's death, his wife (Edward's sister) Æthelflæd ruled as "Lady of the Mercians" and continued expansion. It seems Edward had his son Æthelstan brought up in the Mercian court. On Edward's death, Æthelstan succeeded to the Mercian kingdom, and, after some uncertainty, Wessex.
Æthelstan continued the expansion of his father and aunt | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | and was the first king to achieve direct rulership of what we would now consider England. The titles attributed to him in charters and on coins suggest a still more widespread dominance. His expansion aroused ill-feeling among the other kingdoms of Britain, and he defeated a combined Scottish-Viking army at the Battle of Brunanburh. However, the unification of England was not a certainty. Under Æthelstan's successors Edmund and Eadred the English kings repeatedly lost and regained control of Northumbria. N | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | evertheless, Edgar, who ruled the same expanse as Æthelstan, consolidated the kingdom, which remained united thereafter.
England under the Danes and the Norman conquest
There were renewed Scandinavian attacks on England at the end of the 10th century. Æthelred ruled a long reign but ultimately lost his kingdom to Sweyn of Denmark, though he recovered it following the latter's death. However, Æthelred's son Edmund II Ironside died shortly afterwards, allowing Cnut, Sweyn's son, to become king of England. U | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | nder his rule the kingdom became the centre of government for the North Sea empire which included Denmark and Norway.
Cnut was succeeded by his sons, but in 1042 the native dynasty was restored with the accession of Edward the Confessor. Edward's failure to produce an heir caused a furious conflict over the succession on his death in 1066. His struggles for power against Godwin, Earl of Wessex, the claims of Cnut's Scandinavian successors, and the ambitions of the Normans whom Edward introduced to English | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | politics to bolster his own position caused each to vie for control of Edward's reign.
Harold Godwinson became king, probably appointed by Edward on his deathbed and endorsed by the Witan. But William of Normandy, Harald Hardråde (aided by Harold Godwin's estranged brother Tostig) and Sweyn II of Denmark all asserted claims to the throne. By far the strongest hereditary claim was that of Edgar the Ætheling, but due to his youth and apparent lack of powerful supporters, he did not play a major part in the s | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | truggles of 1066, although he was made king for a short time by the Witan after the death of Harold Godwinson.
In September 1066, Harald III of Norway and Earl Tostig landed in Northern England with a force of around 15,000 men and 300 longships. Harold Godwinson defeated the invaders and killed Harald III of Norway and Tostig at the Battle of Stamford Bridge.
On 28 September 1066, William of Normandy invaded England in a campaign called the Norman Conquest. After marching from Yorkshire, Harold's exhaust | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ed army was defeated and Harold was killed at the Battle of Hastings on 14 October. Further opposition to William in support of Edgar the Ætheling soon collapsed, and William was crowned king on Christmas Day 1066. For five years, he faced a series of rebellions in various parts of England and a half-hearted Danish invasion, but he subdued them and established an enduring regime.
Norman England
The Norman Conquest led to a profound change in the history of the English state. William ordered the compilatio | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | n of the Domesday Book, a survey of the entire population and their lands and property for tax purposes, which reveals that within 20 years of the conquest the English ruling class had been almost entirely dispossessed and replaced by Norman landholders, who monopolised all senior positions in the government and the Church. William and his nobles spoke and conducted court in Norman French, in both Normandy and England. The use of the Anglo-Norman language by the aristocracy endured for centuries and left an | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | indelible mark in the development of modern English.
Upon being crowned, on Christmas Day 1066, William immediately began consolidating his power. By 1067, he faced revolts on all sides and spent four years crushing them. He then imposed his superiority over Scotland and Wales, forcing them to recognise him as overlord.
The English Middle Ages were characterised by civil war, international war, occasional insurrection, and widespread political intrigue among the aristocratic and monarchic elite. England | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | was more than self-sufficient in cereals, dairy products, beef and mutton. Its international economy was based on wool trade, in which wool from the sheepwalks of northern England was exported to the textile cities of Flanders, where it was worked into cloth. Medieval foreign policy was as much shaped by relations with the Flemish textile industry as it was by dynastic adventures in western France. An English textile industry was established in the 15th century, providing the basis for rapid English capital | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | accumulation.
Henry I, the fourth son of William I the Conqueror, succeeded his elder brother William II as King of England in 1100. Henry was also known as "Henry Beauclerc" because he received a formal education, unlike his older brother and heir apparent William who got practical training to be king. Henry worked hard to reform and stabilise the country and smooth the differences between the Anglo-Saxon and Anglo-Norman societies. The loss of his son, William Adelin, in the wreck of the White Ship in N | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | ovember 1120, undermined his reforms. This problem regarding succession cast a long shadow over English history.
Henry I had required the leading barons, ecclesiastics and officials in Normandy and England, to take an oath to accept Matilda (also known as Empress Maud, Henry I's daughter) as his heir. England was far less than enthusiastic to accept an outsider, and a woman, as their ruler.
There is some evidence that Henry was unsure of his own hopes and the oath to make Matilda his heir. Probably Henry | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | hoped Matilda would have a son and step aside as Queen Mother. Upon Henry's death, the Norman and English barons ignored Matilda's claim to the throne, and thus through a series of decisions, Stephen, Henry's favourite nephew, was welcomed by many in England and Normandy as their new king.
On 22 December 1135, Stephen was anointed king with implicit support by the church and nation. Matilda and her own son waited in France until she sparked the civil war from 1139 to 1153 known as the Anarchy. In the autum | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | n of 1139, she invaded England with her illegitimate half-brother Robert of Gloucester. Her husband, Geoffroy V of Anjou, conquered Normandy but did not cross the channel to help his wife. During this breakdown of central authority, nobles built adulterine castles (i.e. castles erected without government permission), which were hated by the peasants, who were forced to build and maintain them.
Stephen was captured, and his government fell. Matilda was proclaimed queen but was soon at odds with her subjects | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | and was expelled from London. The war continued until 1148, when Matilda returned to France. Stephen reigned unopposed until his death in 1154, although his hold on the throne was uneasy. As soon as he regained power, he began to demolish the adulterine castles, but kept a few castles standing, which put him at odds with his heir. His contested reign, civil war, and lawlessness saw a major swing in power towards feudal barons. In trying to appease Scottish and Welsh raiders, he handed over large tracts of | [
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13485 | https://en.wikipedia.org/wiki/History%20of%20England | History of England | land.
England under the Plantagenets
The first Angevins
Empress Matilda and Geoffroy's son, Henry, resumed the invasion; he was already Count of Anjou, Duke of Normandy and Duke of Aquitaine when he landed in England. When Stephen's son and heir apparent Eustace died in 1153, Stephen made an agreement with Henry of Anjou (who became Henry II) to succeed Stephen and guarantee peace between them. The union was retrospectively named the Angevin Empire. Henry II destroyed the remaining adulterine castles and | [
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