 0\) The in vivo data though interesting is difficult to relate to the rest of the data. I do not think that it enhances this paper. e.g. (0) The authors note latency differences between Pbg and LP but there also appear to be rate differences; latency is often longer in weak responses and a fair comparison would need to match the firing rates of the two populations; (0) The authors note the presence of direction selective units in Pbg but not LP (Figure 0A) but as there are only 00 units in LP I am not sure how confident one should be in this. In addition, it is clear from other work that LP organisation depends on e.g. AP location -- were the injection sites and recordings matched in location? As for the other data I could not find confidence intervals on the estimates of functional properties in Pbg and LP, making it difficult to know how well they can be distinguished.  