colpate pollen folding in the $\gamma$-$f$ plane, showing equatorial cross-sections of the pollen shapes. The relative change of volume upon desiccation is $\Delta V/V_0=0.00$ throughout, and the colpi (shown in red in the cross-sections) span $A_{ap}/A_0=0/0$ of the total area of the pollen surface. Numbers next to the pollen shapes denote the elongation of the bounding box of the shapes, $0u_z/(u_x+u_y)$, indicating the prolateness of the grain. The shaded region of the phase diagram corresponds to the elastic parameters where the apertures close completely and nearly symmetrically, without asymmetric deformation of the exine.[]{data-label="fig:S0"}](FigS0-small.pdf){width="00.00000%"}