 barcoded libraries following the manufacturer's instructions. Paired-end reads were aligned using Bowtie0 version 0.0.0 with options:--local--very-sensitive-local--no-unal--no-mixed--no-discordant--phred00 -I 00 -X 000.  Reporting summary {#Sec00} -----------------  Further information on research design is available in the [Nature Research Reporting Summary](#MOESM0){ref-type="media"} linked to this article.  Supplementary information =========================   {#Sec00}  Supplementary Information Peer Review File Reporting Summary  Source Data  **Journal peer review information:** *Nature Communications* thanks 