粪蜣螂属

**糞蜣螂屬**（學名：_Copris_）是金龜子科下的一個屬。

## 下屬物種

本屬包括以下物種：

* _Copris 4dentatus_ Fabricius, 1798
* _Copris (Copris) acutidens_ Motschulsky, 1860
* _Copris (Copris) afgoi_ Cambefort, 1992
* _Copris (Copris) agnus_ Sharp, 1875
* _Copris (Copris) algol_ Nguyen-phung, 1989
* _Copris amabilis_ Kolbe, 1914
* 亞馬孫蜣螂 _Copris amazonicus_ Darling & Génier, 2018
* _Copris amyntor_ Klug, 1855
* _Copris (Copris) anceus_ (Olivier, 1789)
* _Copris andrewesi_ Waterhouse, 1891
* _Copris (Copris) angusticornis_ Arrow, 1933
* _Copris angustus_ Nguyen-phung & Cambefort, 1986
* _Copris anomiopseoides_ Boucomont, 1924
* _Copris antares_ Ferreira, 1958
* _Copris arcturus_ Gillet, 1907
* _Copris ares_ Zelenka, 1993
* _Copris arimotoi_ Ochi & Araya, 1992
* 亞利桑那蜣螂 _Copris arizonensis_ Schaeffer, 1906
* _Copris armatus_ Harold, 1869
* _Copris (Copris) armeniacus_ Faldermann, 1836
* _Copris armiger_ Gillet, 1910
* _Copris arrowi_ Felsche, 1910
* _Copris aspericollis_ Gillet, 1910
* _Copris atropolitus_ Gillet, 1910
* _Copris (Sinocopris) barclayi_ Ochi, Kon & Bai, 2009
* _Copris (Paracopris) bartolozzi_ Ochi, Kon & Pham, 2019
* _Copris basilewskyi_ Ferreira, 1962
* _Copris basipunctatus_ Balthasar, 1942
* _Copris bellator_ Chevrolat, 1844
* _Copris bengalensis_ Gillet, 1911
* _Copris bidens_ Kolbe, 1893
* _Copris bootes_ Klug, 1855
* _Copris boucardi_ Harold, 1869
* _Copris bovinus_ Gillet, 1908
* _Copris brachypterus_ Nomura, 1964
* _Copris brevicornis_ Darling & Génier, 2018
* _Copris brittoni_ Ferreira, 1962
* _Copris (Copris) caelatus_ (Fabricius, 1794)
* _Copris caliginosus_ Kohlmann, Cano & Delgado, 2003
* _Copris (Copris) cambeforti_ Nguyen-phung, 1988
* _Copris cambodiensis_ Ochi, Kon & Kawahara, 2008
* _Copris camerunus_ Felsche, 1904
* _Copris caobangensis_ Bui, Dumack & Bonkowski, 2018
* _Copris capensis_ Waterhouse, 1891
* _Copris cariniceps_ Felsche, 1910
* _Copris carinicus_ Gillet, 1910
* _Copris carmelita_ Fabricius, 1801
* _Copris (Copris) cassius_ Péringuey, 1901
* _Copris (Copris) celebensis_ Lansberge, 1886
* _Copris (Sinocopris) cheni_ Ochi, Kon & Bai, 2007
* _Copris chimalapensis_ Mora-Aguilar & Delgado, 2015
* _Copris (Paracopris) chinmontanus_ Ochi, Kon & Kawahara, 2019
* _Copris colmanti_ Gillet, 1908
* _Copris complexus_ Nguyen-phung, 1989
* _Copris compressipennis_ Gillet, 1910
* _Copris confucius_ Harold, 1877
* _Copris (Sinocopris) continentalis_ Ochi & Kon, 2010
* _Copris coriarius_ Gillet, 1907
* _Copris cornifrons_ Boheman, 1860
* _Copris (Copris) corniger_ Sahlberg, 1823
* _Copris (Copris) corpulentus_ Gillet, 1910
* _Copris (Copris) costaricensis_ Gahan, 1894
* _Copris (Copris) crassus_ Deschodt & Davis, 2015
* _Copris (Paracopris) cribratus_ Gillet, 1927
* _Copris (Copris) cribricollis_ Gillet, 1910
* _Copris (Copris) curlettii_ Moretto, 2011
* _Copris davidi_ Darling & Génier, 2018
* _Copris (Copris) davisi_ Nguyen-phung & Cambefort, 1986
* _Copris (Paracopris) davisoni_ Waterhouse, 1891
* _Copris decellei_ Cambefort, 1992
* _Copris (Copris) delicatus_ Arrow, 1931
* _Copris denticulatus_ Nguyen-phung, 1988
* _Copris diversus_ Waterhouse, 1891
* _Copris (Copris) draco_ Arrow, 1906
* _Copris (Copris) dracunculus_ Ferreira, 1959
* _Copris (Copris) druidum_ Heer, 1862
* _Copris dudleyi_ Cambefort, 1992
* _Copris eburneus_ Cambefort, 1992
* _Copris elpheneroides_ Felsche, 1910
* _Copris elphenor_ Klug, 1855
* _Copris erratus_ Lansberge, 1886
* _Copris evanidus_ Klug, 1855
* _Copris excisus_ Waterhouse, 1891
* _Copris fallaciosus_ Gillet, 1907
* _Copris fallax_ Felsche, 1910
* _Copris (Sinocopris) felschei_ Reitter, 1892
* _Copris fidius_ (Olivier, 1789)
* _Copris frankenbergerianus_ Balthasar, 1958
* _Copris freyi_ Ferreira, 1961
* _Copris fricator_ (Fabricius, 1787)
* _Copris fukiensis_ Balthasar, 1952
* _Copris furciceps_ Felsche, 1910
* _Copris garambae_ Cambefort, 1992
* _Copris gazellarum_ Gillet, 1918
* _Copris gibbulus_ Lansberge, 1886
* _Copris gilleti_ Kolbe, 1907
* _Copris gladiator_ Arrow, 1933
* _Copris gopheri_ Hubbard, 1894
* _Copris gracilis_ Waterhouse, 1891
* _Copris graueri_ Kolbe, 1914
* _Copris (Copris) halffteri_ Matthews, 1959
* _Copris harrisi_ Waterhouse, 1891
* _Copris (Copris) hartli_ Frey, 1975
* _Copris (Copris) hatayamai_ Ochi & Araya, 1992
* _Copris hispanus_ (Linnaeus, 1764)
* _Copris howdeni_ Matthews & Halffter, 1959
* _Copris (Copris) humilis_ Gillet, 1908
* _Copris (Copris) hybridus_ Gillet, 1910
* _Copris (Copris) igualensis_ Warner, 1990
* _Copris (Copris) illotus_ Balthasar, 1942
* _Copris (Paracopris) imitans_ Felsche, 1910
* _Copris (Copris) inaequabilis_ Zhang, 1997
* _Copris incertus_ Say, 1835
* _Copris inemarginatus_ Blatchley, 1918
* _Copris inhalatus_ Quedenfeldt, 1884
* _Copris insidiosus_ Péringuey, 1901
* _Copris integer_ Reiche, 1847
* _Copris interioris_ Kolbe, 1897
* _Copris iris_ Sharp, 1875
* _Copris jacchoides_ Nguyen-phung & Cambefort, 1987
* _Copris (Copris) jacchus_ (Fabricius, 1775)
* _Copris jahi_ Nguyen-phung & Cambefort, 1986
* 爪哇蜣螂 _Copris (Paracopris) javanensis_ Bui, Ziegler & Bonkowski, 2019
* _Copris johkii_ Ochi & Kon, 2014
* _Copris juanmorai_ Mora-Aguilar & Delgado, 2015
* _Copris (Copris) jucundus_ Gillet, 1932
* _Copris (Copris) kachinensis_ Ochi, Kon & Kawahara, 2008
* _Copris (Copris) kartlinus_ Kabakov, 1988
* _Copris kasagii_ Ochi & Kon, 1996
* _Copris (Copris) kasanka_ Josso & Prévost, 2016
* _Copris katangae_ Cambefort & Nguyen-phung, 1996
* _Copris (Copris) keralensis_ Gill, 1986
* _Copris (Sinocopris) kimioi_ Ochi & Kon, 2021
* _Copris (Paracopris) kiriromensis_ Ochi & Kon, 2021
* _Copris kiuchii_ Masumoto, 1989
* _Copris klapperichi_ Balthasar, 1942
* _Copris klugi_ Harold, 1869
* _Copris kusakabei_ Ochi, Kon & Kawahara, 2005
* _Copris laeviceps_ Harold, 1869
* _Copris laevigatus_ Gillet, 1927
* _Copris laiiformis_ Nguyen-phung & Cambefort, 1987
* _Copris laioides_ Boucomont, 1932
* _Copris laius_ Harold, 1868
* _Copris lannathai_ Hanboonsong, Masumoto & Ochi, 2003
* _Copris (Copris) laosensis_ Ochi & Kon, 2021
* _Copris (Copris) leakeyorum_ Paulian, 1976
* _Copris lecontei_ Matthews, 1961
* _Copris (Copris) leruei_ Josso & Prévost, 2017
* _Copris lugubris_ Boheman, 1858
* _Copris lunaris_ (Linnaeus, 1758)
* _Copris macacus_ Lansberge, 1886
* _Copris macclevei_ Warner, 1990
* _Copris (Copris) macer_ Péringuey, 1901
* _Copris maesi_ Ratcliffe, 1998
* _Copris magicus_ Harold, 1881
* _Copris (Copris) manni_ Ochi, Kon & Bai, 2009
* _Copris (Copris) marcoberrai_ Ochi, Kon & Pham, 2019
* _Copris marcus_ Gillet, 1921
* _Copris (Copris) martinae_ Nguyen-phung, 1988
* _Copris martinezi_ Matthews & Halffter, 1968
* _Copris matthewsi_ Delgado & Kohlmann, 2001
* _Copris (Copris) medogensis_ Zhang, 1988
* _Copris (Copris) megaceratoides_ Waterhouse, 1891
* _Copris (Copris) megasoma_ Matthews & Halffter, 1959
* _Copris (Copris) mesacanthus_ Harold, 1878
* _Copris (Copris) mexicanus_ Matthews & Halffter, 1959
* _Copris (Copris) micros_ Ochi & Kon, 2021
* _Copris (Microcopris) mindorensis_ Ochi
* _Copris minutus_ (Drury, 1773)
* _Copris moechus_ LeConte, 1854
* _Copris moffartsi_ Gillet, 1907
* _Copris mongkhoni_ Hanboonsong, Masumoto & Ochi, 2003
* _Copris montivagus_ Gillet, 1908
* _Copris (Paracopris) montreuili_ Ochi, Kon & Pham, 2019
* _Copris (Copris) morettoi_ Josso & Prévost, 2020
* _Copris moroni_ Darling & Génier, 2018
* _Copris morphaeus_ Gillet, 1932
* _Copris (Copris) mourgliai_ Zelenka, 1993
* _Copris (Copris) neglectus_ Moxey, 1963
* _Copris neotridens_ Génier & Prévost, 2017
* _Copris (Copris) nepos_ Gillet, 1908
* _Copris (Copris) nevinsoni_ Waterhouse, 1891
* _Copris (Paracopris) nonarmatus_ Ochi & Kon, 2021
* _Copris novaki_ Zelenka, 1992
* _Copris nubilosus_ Kohlmann, Cano & Delgado, 2003
* _Copris numa_ Lansberge, 1886
* _Copris obenbergeri_ Balthasar, 1933
* _Copris (Copris) obesus_ Boheman, 1857
* _Copris ochus_ (Motschulsky, 1860)
* _Copris (Paracopris) olivieri_ Ochi, Kon & Kawahara, 2019
* _Copris orion_ Klug, 1835
* _Copris orphanus_ Guérin-Méneville, 1847
* _Copris parapecuarius_ Ochi, Kon & Kawahara, 2008
* _Copris (Copris) parasonensis_ Ochi, Kon & Pham, 2019
* _Copris (Copris) paraszechouanicus_ Ochi, Kon & Pham, 2018
* _Copris pauliani_ Nguyen-phung & Cambefort, 1986
* _Copris pecuarius_ Lewis, 1884
* _Copris pedarioides_ Lansberge, 1886
* _Copris phungae_ Cambefort, 1992
* _Copris phylax_ Gillet, 1908
* _Copris poggii_ Ochi & Kon, 2005
* _Copris (Paracopris) popaensis_ Ochi, Kon & Kawahara, 2019
* _Copris potanini_ Semenov, 1891
* _Copris (Copris) prevosti_ Josso, 2011
* _Copris (Copris) pristinus_ Pierce, 1946
* _Copris pseudobootes_ Ferreira, 1962
* _Copris pseudochus_ Ochi & Kon, 2004
* _Copris (Copris) pseudomoffartsi_ Moretto, 2011
* _Copris pseudosinicus_ Balthasar, 1958
* _Copris (Sinocopris) pueli_ Mollandin De Boissy, 1905
* _Copris punctatus_ Gillet, 1910
* _Copris puncticollis_ Boheman, 1857
* _Copris punctipennis_ Boucomont, 1914
* _Copris punctulatus_ Wiedemann, 1823
* _Copris punjabensis_ Gillet, 1921
* _Copris quasilaevigatus_ Ochi, Kon & Bai, 2007
* _Copris ramosiceps_ Gillet, 1921
* _Copris rebouchei_ Harold, 1869
* _Copris remotus_ LeConte, 1866
* _Copris renwarti_ Nguyen-phung & Cambefort, 1986
* _Copris repertus_ Walker, 1858
* _Copris ribbei_ Lansberge, 1886
* _Copris riekoae_ Ochi & Kon, 2003
* _Copris ritsemae_ Harold, 1875
* _Copris rugosus_ Gillet, 1908
* _Copris rupicapra_ Gravenhorst, 1807
* _Copris ruricola_ Balthasar, 1933
* _Copris (Sinocopris) sabinus_ Gillet, 1910
* _Copris (Sinocopris) sacontala_ Redtenbacher, 1848
* _Copris (Copris) sallei_ Harold, 1869
* _Copris sarpedon_ Harold, 1868
* _Copris saundersi_ Harold, 1869
* _Copris (Copris) schoolmeestersi_ Ochi & Kon, 2010
* _Copris scorpio_ Zelenka, 1993
* _Copris serius_ Nguyen-phung, 1987
* _Copris (Copris) sexdentatus_ Thunberg, 1818
* _Copris (Copris) siamensis_ Gillet, 1910
* _Copris (Copris) siangensis_ Biswas, 1980
* _Copris sierrensis_ Matthews, 1961
* _Copris signatus_ Walker, 1858
* _Copris simonettai_ Cambefort, 1992
* _Copris simulator_ Nguyen-phung & Cambefort, 1986
* _Copris (Copris) singularis_ Nguyen-phung & Cambefort, 1986
* _Copris sinicus_ Hope, 1842
* _Copris (Copris) sinon_ (Olivier, 1789)
* _Copris sodalis_ Walker, 1858
* _Copris sonensis_ Bui, Dumack & Bonkowski, 2018
* _Copris sorex_ Balthasar, 1942
* _Copris sphaeropterus_ Harold, 1877
* _Copris spinator_ Harold, 1881
* _Copris subdolus_ Balthasar, 1958
* _Copris subpunctatus_ Gillet, 1910
* _Copris subsidens_ Péringuey, 1901
* _Copris (Copris) subterraneus_ Heer, 1862
* _Copris sumatrensis_ Gillet, 1921
* _Copris surdus_ Arrow, 1931
* _Copris susanae_ Darling & Génier, 2018
* _Copris szechouanicus_ Balthasar, 1958
* _Copris (Sinocopris) takanoi_ Ochi, Kon & Bai, 2020
* _Copris (Sinocopris) teramachii_ Ochi, Kon & Bai, 2020
* _Copris tetraodon_ Gillet, 1910
* _Copris (Copris) tricorniprothoracicus_ Ochi, Kon & Kawahara, 2020
* _Copris tridentatus_ Solis & Kohlmann, 2003
* 三開蜣螂 _Copris tripartitus_ Waterhouse, 1875
* _Copris truncatus_ Felsche, 1901
* _Copris (Sinocopris) tsukamotoi_ Ochi, Kon & Kawahara, 2008
* _Copris typhoeus_ Gerstaecker, 1884
* _Copris uenoi_ Hanboonsong, Masumoto & Ochi, 2003
* _Copris (Sinocopris) umbilicatus_ Abeille De Perrin, 1901
* _Copris urus_ Boheman, 1857
* _Copris usambaricus_ Gillet, 1908
* _Copris (Copris) vankhaii_ Nguyen-phung, 1988
* _Copris victorini_ Boheman, 1857
* _Copris (Copris) vietnamicus_ Kabakov, 1994
* _Copris (Copris) vietnamimontanus_ Ochi, Kon & Pham, 2018
* _Copris (Copris) vilhenai_ Ferreira, 1962
* _Copris (Copris) vrydaghi_ Ferreira, 1962
* _Copris warneri_ Mccleve & Kohlmann, 2005
* _Copris wiesei_ Kolbe, 1914
* _Copris (Sinocopris) xingkeyangi_ Ochi, Kon & Bai, 2020
* _Copris (Copris) youngai_ Balthasar, 1967
* 張氏蜣螂 _Copris (Copris) zhangi_ Ochi, Kon & Bai, 2009

## 參考文獻