查尔斯·克拉克_(植物学家)

**查爾斯・克拉克**（**Charles M. Clarke**）是專業從事於豬籠草屬食蟲植物的植物學家與分類學家。克拉克出生於澳大利亞墨爾本。他獲得了墨爾本莫納什大學（Monash University）的生物學榮譽學位，同時還在新南威爾斯阿米代爾的新英格蘭大學（University of New England）取得了生態系統管理博士學位。

克拉克第一次到婆羅洲研究豬籠草是在 1987 年。1989 年至 1990 年，他生活在汶萊以研究豬籠草屬植物的生態問題。在這兩次旅行間，克拉克還曾在昆士蘭州的詹姆斯・庫克大學（James Cook University）任教生態學和生物統計學，同時還在香港擔任園藝顧問。

克拉克現就任於莫納什大學馬來西亞分校。克拉克已完成了五本關於豬籠草屬的著作，其中最重要的是《婆羅洲的豬籠草屬植物》（_Nepenthes_ of Borneo）和《蘇門答臘島與西馬來西亞的豬籠草》（_Nepenthes_ of Sumatra and Peninsular Malaysia）。

克拉克描述了 5 個豬籠草屬物種，分別為本斯通豬籠草（_Nepenthes benstonei_）、陳氏豬籠草（_Nepenthes chaniana_）、泉氏豬籠草（_Nepenthes izumiae_）、賈桂琳豬籠草（_Nepenthes jacquelineae_）和堅韌豬籠草（_Nepenthes tenax_）。

## 著作

* Clarke, C.M. 1992. The ecology of metazoan communities in _Nepenthes_ pitcher plants in Borneo, with special reference to the community of _Nepenthes bicalcarata_ Hook.f. Ph.D. thesis, University of New England, Armidale, N.S.W. 269 pp.
* Clarke, C.[M.] 1993. The Possible Functions of the Thorns of _Nepenthes bicalcarata_ (Hook.f.) Pitchers.PDF (194 KiB) _Carnivorous Plant Newsletter_ **22**(1–2): 27–28.
* Clarke, C.M. & R.L. Kitching 1993. The Metazoan Food Webs from Six Bornean _Nepenthes_ Species. _Ecological Entomology_ **18**: 7–16.
* Clarke, C.M. & J.A. Moran 1994. A further record of aerial pitchers in _Nepenthes ampullaria_ Jack. _Malayan Nature Journal_ **47**: 321–323.
* Clarke, C.M. & R.L. Kitching 1995. Swimming Ants and Pitcher Plants: a Unique Ant-Plant Interaction from Borneo. _Journal of Tropical Ecology_ **11**(4): 589–602.
* Clarke, C.M. 1997. _Nepenthes of Borneo_. Natural History Publications (Borneo), Kota Kinabalu. xi + 207 pp.
* Clarke, C.M. 1997. Another nice trip to Sumatra （頁面存檔備份，存於網際網路檔案館）. _Carnivorous Plant Newsletter_ **26**(1): 4–10.
* Clarke, C.M. 1997. The effects of pitcher dimorphism on the metazoan community of the carnivorous pitcher plant _Nepenthes bicalcarata_ Hook.f. _Malayan Nature Journal_ **50**: 149–157.
* Clarke, C.M. 1998. Initial colonisation and prey capture in _Nepenthes bicalcarata_ (Nepenthaceae) pitchers in Brunei. _Sandakania_ **12**: 27–36.
* Clarke, C.M. 1998. The aquatic arthropod community of the pitcher plant, _Nepenthes bicalcarata_ (Nepenthaceae) in Brunei. _Sandakania_ **11**: 55–60.
* Clarke, C.M. 1998. A re-examination of geographical variation in _Nepenthes_ food webs. _Ecography_ **21**: 430–436.
* Clarke, C.M. 1999. _Nepenthes benstonei_ (Nepenthaceae), a new pitcher plant from Peninsular Malaysia. _Sandakania_ **13**: 79–87.
* Clarke, C.M. 2001. Ecology & Conservation of Montane _Nepenthes_ (Nepenthaceae) in Sumatra. Fourth International Flora Malesiana Symposium. 20-24 July 1998, Kuala Lumpur.
* Clarke, C.M. 2001. _Nepenthes of Sumatra and Peninsular Malaysia_. Natural History Publications (Borneo), Kota Kinabalu. ix + 325 pp.
* Clarke, C.M. 2001. _A Guide to the Pitcher Plants of Sabah_. Natural History Publications (Borneo), Kota Kinabalu. iv + 40 pp.
* Moran, J.A., M.A. Merbach, N.J. Livingston, C.M. Clarke & W.E. Booth 2001. Termite Prey Specialization in the Pitcher Plant _Nepenthes albomarginata_ - Evidence from Stable Isotope Analysis.PDF _Annals of Botany_ **88**: 307–311.
* Clarke, C.M. 2002. _A Guide to the Pitcher Plants of Peninsular Malaysia_. Natural History Publications (Borneo), Kota Kinabalu. iv + 32 pp.
* Clarke, C.M., T. Davis & R. Tamin 2003. _Nepenthes izumiae_ (Nepenthaceae): a new species from Sumatra. _Blumea_ **48**: 179–182.
* Moran, J.A., C.M. Clarke & B.J. Hawkins 2003. From Carnivore to Detritivore? Isotopic Evidence for Leaf Litter Utilization by the Tropical Pitcher Plant _Nepenthes ampullaria_. _International Journal of Plant Sciences_ **164**: 635–639.
* Dong, T.T.X., Q.M. Xiao, C.M. Clarke, H.S. Zhong, N.J. Zhao, K.L. Chun & K.W.K. Tsim 2003. Phylogeny of _Astragalus_ in China: Molecular evidence from the DNA sequences of 5S rRNA spacer, ITS, and 18S rRNA. _Journal of Agricultural and Food Chemistry_ **51**(23): 6709–6714.
* Clarke, C.M. & C.C. Lee 2004. _A Pocket Guide: Pitcher Plants of Sarawak_. Natural History Publications (Borneo), Kota Kinabalu. vi + 81 pp.
* Cantley, R., C.M. Clarke, J. Cokendolpher, B. Rice & A. Wistuba 2004. _Nepenthes clipeata_ Survival Project. International Carnivorous Plant Society.
* Clarke, C.M., C.C. Lee & S. McPherson 2006. _Nepenthes chaniana_ (Nepenthaceae), a new species from north-western Borneo. _Sabah Parks Journal_ **7**: 53–66.
* Clarke, C.M & R. Kruger 2006. _Nepenthes tenax_ C.Clarke and R.Kruger (Nepenthaceae), a new species from Cape York Peninsula, Queensland. _Austrobaileya_ **7**(2): 319–324.
* Clarke, C.M. 2006. Introduction. In: Danser, B.H. _The Nepenthaceae of the Netherlands Indies_. Natural History Publications (Borneo), Kota Kinabalu. pp. 1–15
* Clarke, C.M., U. Bauer, C.C. Lee, A.A. Tuen, K. Rembold & J.A. Moran 2009. Tree shrew lavatories: a novel nitrogen sequestration strategy in a tropical pitcher plant.PDF _Biology Letters_ **5**(5): 632–635. doi:10.1098/rsbl.2009.0311
* Chin, L., J.A. Moran & C. Clarke 2010. Trap geometry in three giant montane pitcher plant species from Borneo is a function of tree shrew body size. _New Phytologist_, published online January 22, 2010. doi:10.1111/j.1469-8137.2009.03166.x

## 參考文獻

## 擴展閱讀

* Clarke, C.M. 1997. _Nepenthes of Borneo_. Natural History Publications (Borneo), Kota Kinabalu. xi + 207 pp. ISBN 983-812-057-X