厚壁蕨属

**膜蕨屬**（學名：_Hymenophyllum_），是膜蕨科下的一個科。

## 下屬分類

本屬包括以下物種：

* _Hymenophyllum abruptum_ Hook.
* _Hymenophyllum acanthoides_ (Bosch) Rosenst.
* _Hymenophyllum acutum_ (C. Presl) Ebihara & K. Iwats.
* _Hymenophyllum adiantoides_ Bosch
* _Hymenophyllum aeruginosum_ (Poir.) Carmich.
* _Hymenophyllum album_ (Blume) Parris
* _Hymenophyllum alveolatum_ C. Chr.
* _Hymenophyllum amabile_ Morton
* _Hymenophyllum andinum_ Bosch
* _Hymenophyllum angulosum_ Christ
* _Hymenophyllum angustum_ Bosch
* _Hymenophyllum antillense_ (Jenman) Jenman
* _Hymenophyllum apiculatum_ Mett. ex Kuhn
* _Hymenophyllum applanatum_ (A. M. Gray & R. G. Williams) Ebihara & K. Iwats.
* _Hymenophyllum apteryx_ M. Kessler & Sundue
* _Hymenophyllum archboldii_ (Copel.) C. V. Morton
* _Hymenophyllum armstrongii_ (Baker) Kuhn
* _Hymenophyllum asplenioides_ (Sw.) Sw.
* _Hymenophyllum assurgens_ M. Kessler & A. R. Sm.
* _Hymenophyllum australe_ Willd.
* _Hymenophyllum axillare_ Sw.
* 蕗蕨 _Hymenophyllum badium_ Hook. & Grev.
* _Hymenophyllum baileyanum_ Domin
* _Hymenophyllum balfourii_ Baker
* 華東膜蕨 _Hymenophyllum barbatum_ (Bosch) Baker
* _Hymenophyllum bartlettii_ (Copel.) C. V. Morton
* _Hymenophyllum batuense_ Rosenst.
* _Hymenophyllum bicolanum_ Copel.
* _Hymenophyllum bivalve_ (G. Forst.) Sw.
* 爪哇厚壁蕨 _Hymenophyllum blandum_ Racib.
* _Hymenophyllum bontocense_ Copel.
* _Hymenophyllum brachyglossum_ A. Braun ex Kunze
* 短柄膜蕨 _Hymenophyllum brachypus_ Sodiro
* _Hymenophyllum braithwaitei_ Ebihara & K. Iwats.
* _Hymenophyllum brasilianum_ (Fée) Rosenst.
* _Hymenophyllum brassii_ C. Chr.
* _Hymenophyllum breve_ Rosenst.
* _Hymenophyllum brevidens_ Alderw.
* _Hymenophyllum brevifrons_ Kunze
* _Hymenophyllum brevistipes_ Liebm.
* _Hymenophyllum bryophilum_ C. Chr.
* _Hymenophyllum caespitosum_ Gaudich.
* _Hymenophyllum caparaoense_ Brade
* _Hymenophyllum capense_ Schrad.
* _Hymenophyllum capillaceum_ Roxb.
* _Hymenophyllum capillare_ Desv.
* _Hymenophyllum capurroi de la_ Sota
* _Hymenophyllum cardunculus_ C. Chr.
* _Hymenophyllum caudiculatum_ Mart.
* _Hymenophyllum cernuum_ A. Gepp
* _Hymenophyllum ciliatum_ (Sw.) Sw.
* _Hymenophyllum cincinnatum_ A. Gepp
* _Hymenophyllum cocosense_ A. Rojas
* _Hymenophyllum compactum_ Bonap.
* _Hymenophyllum consanguineum_ C. V. Morton
* _Hymenophyllum copelandii_ C. V. Morton
* _Hymenophyllum cordobense_ (Hieron.) C. Larsen & Arana
* 皺葉蕗蕨 _Hymenophyllum corrugatum_ Christ
* _Hymenophyllum crassipetiolatum_ Stolze
* _Hymenophyllum crispatoalatum_ Hayata
* _Hymenophyllum crispatum_ Wall. ex Hook. & Grev.
* _Hymenophyllum crispum_ Kunth
* _Hymenophyllum cristatum_ Hook. & Grev.
* _Hymenophyllum cruentum_ Cav.
* _Hymenophyllum cuneatum_ Kunze
* _Hymenophyllum cupressiforme_ Labill.
* _Hymenophyllum darwinii_ Hook. fil.
* _Hymenophyllum delicatulum_ Sehnem
* _Hymenophyllum deltoideum_ C. Chr.
* _Hymenophyllum demissum_ (G. Forst.) Sw.
* _Hymenophyllum densipilosum_ A. Rojas
* _Hymenophyllum dentatum_ Cav.
* 厚壁蕨 _Hymenophyllum denticulatum_ Sw.
* _Hymenophyllum dependens_ C. V. Morton
* _Hymenophyllum deplanchei_ Mett. ex Kuhn
* 台灣膜蕨 _Hymenophyllum devolii_ M. J. Lai
* _Hymenophyllum dicranotrichum_ (C. Presl) Sadeb.
* 指狀細口團扇蕨 _Hymenophyllum digitatum_ (Sw.) Fosberg
* _Hymenophyllum dilatatum_ (G. Forst.) Sw.
* _Hymenophyllum dimidiatum_ Mett. ex Kuhn
* _Hymenophyllum diversilobum_ (C. Presl) Fée
* _Hymenophyllum eboracense_ Croxall
* _Hymenophyllum ectocarpon_ Fée
* _Hymenophyllum edanoi_ (Copel.) C. V. Morton
* _Hymenophyllum edentulum_ (Bosch) C. Chr.
* _Hymenophyllum elbertii_ Rosenst.
* _Hymenophyllum elegans_ Spreng.
* _Hymenophyllum ellipticosorum_ Alderw.
* _Hymenophyllum elongatum_ J. W. Grimes
* _Hymenophyllum emarginatum_ Sw.
* 優美膜蕨 _Hymenophyllum exquisitum_ T. C. Hsu & Y. S. Chao
* 毛蕗蕨 _Hymenophyllum exsertum_ Wall. ex Hook.
* _Hymenophyllum falklandicum_ Baker
* _Hymenophyllum farallonense_ Hieron.
* _Hymenophyllum feejeense_ Brack.
* _Hymenophyllum fendlerianum_ Sturm
* _Hymenophyllum ferax_ Bosch
* _Hymenophyllum ferrugineum_ Colla
* _Hymenophyllum filmenofilicum_ Christenh. & Schwartsb.
* 流蘇苞蕗蕨 _Hymenophyllum fimbriatum_ J. Sm.
* _Hymenophyllum firmum_ Alderw.
* _Hymenophyllum flabellatum_ Labill.
* _Hymenophyllum flexuosum_ A. Cunn.
* _Hymenophyllum foersteri_ Rosenst.
* _Hymenophyllum foxworthyi_ Copel.
* _Hymenophyllum fragile_ (Hedw.) Morton
* _Hymenophyllum francii_ (Christ) Ebihara & K. Iwats.
* _Hymenophyllum frankliniae_ Colenso
* _Hymenophyllum fuciforme_ Sw.
* _Hymenophyllum fucoides_ (Sw.) Sw.
* _Hymenophyllum fujisanense_ Nakai
* _Hymenophyllum fumarioides_ Willd.
* _Hymenophyllum fuscum_ (Blume) Bosch
* _Hymenophyllum gardneri_ Bosch
* _Hymenophyllum geluense_ Rosenst.
* _Hymenophyllum glaziovii_ Baker
* _Hymenophyllum gorgoneum_ Copel.
* _Hymenophyllum gracilescens_ Domin
* _Hymenophyllum hallierii_ Rosenst.
* _Hymenophyllum hastatum_ A. Rojas
* _Hymenophyllum heimii_ Tardieu
* _Hymenophyllum helicoideum_ Sodiro
* _Hymenophyllum hemidimorphum_ R. C. Moran & B. Øllg.
* _Hymenophyllum hemipteron_ Rosenst.
* _Hymenophyllum herterianum_ Brause
* _Hymenophyllum herzogii_ Rosenst.
* _Hymenophyllum hieronymi_ (Brause) C. Chr.
* _Hymenophyllum hirsutum_ (L.) Sw.
* _Hymenophyllum hirtellum_ Sw.
* 南洋厚壁蕨 _Hymenophyllum holochilum_ (Bosch) C. Chr.
* _Hymenophyllum horizontale_ C. V. Morton
* _Hymenophyllum hosei_ Copel.
* _Hymenophyllum humatoides_ (Copel.) comb. ined.
* _Hymenophyllum humbertii_ C. Chr.
* _Hymenophyllum humboldtianum_ E. Fourn.
* _Hymenophyllum hygrometricum_ (Poir.) Desv.
* _Hymenophyllum imbricatum_ Blume
* _Hymenophyllum inaequale_ (Poir.) Desv.
* _Hymenophyllum integrivalvatum_ C. Sánchez
* _Hymenophyllum interruptum_ Kunze
* _Hymenophyllum involucratum_ Copel.
* _Hymenophyllum ivohibense_ Tardieu
* _Hymenophyllum jamesonii_ Hook.
* 爪哇蕗蕨 _Hymenophyllum javanicum_ Spreng.
* _Hymenophyllum jimenezii_ A. R. Sm. & M. Kessler
* _Hymenophyllum johorense_ Holttum
* _Hymenophyllum junghuhnii_ Bosch
* _Hymenophyllum kaieteurum_ Jenman
* _Hymenophyllum karstenianum_ J. W. Sturm
* _Hymenophyllum kerianum_ Watts
* 頂果膜蕨 _Hymenophyllum khasyanum_
* _Hymenophyllum klabatense_ Christ
* _Hymenophyllum krauseanum_ Phil.
* _Hymenophyllum kuhnii_ C. Chr.
* _Hymenophyllum lamellatum_ Stolze
* _Hymenophyllum laminatum_ Copel.
* _Hymenophyllum lanatum_ Fée
* _Hymenophyllum lanceolatum_ Hook. & Arn.
* _Hymenophyllum latifrons_ Bosch
* _Hymenophyllum latisorum_ M. Kessler & A. R. Sm.
* _Hymenophyllum laxum_ (Copel.) C. V. Morton
* _Hymenophyllum ledermannii_ Brause
* _Hymenophyllum lehmannii_ Hieron.
* _Hymenophyllum leptocarpum_ Copel.
* _Hymenophyllum leratii_ Rosenst.
* 鱗蕗蕨 _Hymenophyllum levingei_ C. B. Clarke
* _Hymenophyllum lherminieri_ Mett. ex Kuhn
* _Hymenophyllum lindenii_ Hook.
* _Hymenophyllum lineare_ (Sw.) Sw.
* _Hymenophyllum lobatoalatum_ Klotzsch
* _Hymenophyllum lobbii_ Moore
* _Hymenophyllum longifolium_ Alderw.
* 線葉蕗蕨 _Hymenophyllum longissimum_ (Ching & P. S. Chiu) K. Iwats.
* _Hymenophyllum lyallii_ Hook. fil.
* _Hymenophyllum macgillivrayi_ (Baker) Copel.
* _Hymenophyllum macroglossum_ Bosch
* _Hymenophyllum macrosorum_ Alderw.
* _Hymenophyllum macrothecum_ Fée
* _Hymenophyllum maderense_ Gibby & Lovis
* _Hymenophyllum malingii_ (Hook.) Mett.
* _Hymenophyllum marginatum_ Hook. & Grev.
* _Hymenophyllum matthewsii_ Bosch
* _Hymenophyllum maxonii_ Christ ex Morton
* _Hymenophyllum megistocarpum_ (Copel.) C. V. Morton
* _Hymenophyllum melanosorum_ (Copel.) C. V. Morton
* _Hymenophyllum microcarpum_ Desv.
* _Hymenophyllum microchilum_ (Baker) C. Chr.
* _Hymenophyllum mikawanum_ (Seriz.) comb. ined.
* _Hymenophyllum minimum_ A. Rich.
* _Hymenophyllum mirificum_ Morton
* _Hymenophyllum mnioides_ Baker
* _Hymenophyllum molle_ C. V. Morton
* _Hymenophyllum mortonianum_ Lellinger
* _Hymenophyllum mossambicense_ (Schelpe) Schippers
* _Hymenophyllum multialatum_ C. V. Morton
* _Hymenophyllum multicristatum_ A. Rojas
* _Hymenophyllum multifidum_ (G. Forst.) Sw.
* _Hymenophyllum myriocarpum_ Hook.
* _Hymenophyllum nahuelhuapiense_ Diem & J. S. Licht.
* _Hymenophyllum nanostellatum_ Lellinger
* _Hymenophyllum nanum_ Sodiro
* _Hymenophyllum neocaledonicum_ (C. Chr.) Rouhan & C. Del Rio
* _Hymenophyllum nephrophyllum_ Ebihara & K. Iwats.
* _Hymenophyllum nitiduloides_ Copel.
* 細口團扇蕨 _Hymenophyllum nitidulum_ (Bosch) Ebihara & K. Iwats.
* _Hymenophyllum novoguineense_ (Rosenst.) K. Iwats.
* _Hymenophyllum nutantifolium_ Alderw.
* _Hymenophyllum obtusum_ Hook. & Arn.
* _Hymenophyllum okadai_ Masam.
* 長毛蕗蕨 _Hymenophyllum oligosorum_ Makino
* _Hymenophyllum ooides_ F. Muell. & Baker ex Baker
* _Hymenophyllum opacum_ Copel.
* _Hymenophyllum ovatum_ Copel.
* _Hymenophyllum pachydermicum_ Ces.
* 毛葉蕨 _Hymenophyllum pallidum_ (Blume) Ebihara & K. Iwats.
* _Hymenophyllum palmatifidum_ (Müll. Berol.) Ebihara & K. Iwats.
* _Hymenophyllum paniculiflorum_ C. Presl
* _Hymenophyllum paniense_ Ebihara & K. Iwats.
* _Hymenophyllum pantotactum_ Alderw.
* _Hymenophyllum parallelocarpum_ Hayata
* _Hymenophyllum paucicarpum_ Jenman
* _Hymenophyllum pectinatum_ Cav.
* _Hymenophyllum pedicularifolium_ Ces.
* _Hymenophyllum peltatum_ (Poir.) Desv.
* _Hymenophyllum penangianum_ Matthew & Christ ex Christ
* _Hymenophyllum perparvulum_ Alderw.
* _Hymenophyllum perrieri_ Tardieu
* 星毛膜蕨 _Hymenophyllum pilosissimum_ C. Chr.
* _Hymenophyllum pilosum_ Alderw.
* _Hymenophyllum platylobum_ Bosch
* _Hymenophyllum pleiocarpum_ Alderw.
* _Hymenophyllum plicatum_ Kaulf.
* _Hymenophyllum plumieri_ Hook. & Grev.
* _Hymenophyllum plumosum_ Kaulf.
* _Hymenophyllum pluviatile_ Perrie & Brownsey
* _Hymenophyllum pollemonianum_ Rosenst.
* 長柄蕗蕨 _Hymenophyllum polyanthos_ (Sw.) Sw.
* _Hymenophyllum poolii_ Baker
* _Hymenophyllum praetervisum_ Christ
* _Hymenophyllum prionema_ Kunze
* _Hymenophyllum proctoris_ C. Sánchez
* 吊羅蕗蕨 _Hymenophyllum productum_ Kunze
* _Hymenophyllum protrusum_ Hook.
* _Hymenophyllum pulchellum_ Schltdl. & Cham.
* _Hymenophyllum pulcherrimum_ Colenso
* _Hymenophyllum pumilio_ Rosenst.
* _Hymenophyllum pumilum_ C. Moore
* _Hymenophyllum punctisorum_ Rosenst.
* _Hymenophyllum pyramidatum_ Desv.
* _Hymenophyllum quetrihuense_ Diem & J. S. Licht.
* _Hymenophyllum ramosii_ Copel.
* _Hymenophyllum rarum_ R. Br.
* _Hymenophyllum recurvum_ Gaudich.
* _Hymenophyllum reinwardtii_ Bosch
* _Hymenophyllum retusum_ (Copel.) comb. ined.
* _Hymenophyllum revolutum_ Colenso
* _Hymenophyllum ridleyi_ (Copel.) comb. ined.
* _Hymenophyllum ringens_ Christ
* 琉球蕗蕨 _Hymenophyllum riukiuense_ Christ
* _Hymenophyllum rolandi-principis_ Rosenst.
* _Hymenophyllum roraimense_ C. V. Morton
* _Hymenophyllum rosenstockii_ Brause
* _Hymenophyllum rubellum_ Rosenst.
* _Hymenophyllum rufescens_ Kirk
* _Hymenophyllum rufifolium_ Alderw.
* _Hymenophyllum rufifrons_ Alderw.
* _Hymenophyllum rufum_ Fée
* _Hymenophyllum rugosum_ C. Chr. & Skottsb.
* _Hymenophyllum ruizianum_ (Klotzsch) Kunze
* _Hymenophyllum saenzianum_ L. D. Gómez
* _Hymenophyllum salakense_ Racib.
* _Hymenophyllum sampaioanum_ Brade & Rosenst.
* _Hymenophyllum sanguinolentum_ (G. Forst.) Sw.
* _Hymenophyllum scabrum_ A. Rich.
* _Hymenophyllum secundum_ Hook. & Grev.
* 半翼柄蕗蕨 _Hymenophyllum semialatum_ T. C. Hsu
* _Hymenophyllum semiglabrum_ Rosenst.
* _Hymenophyllum senterreanum_ Dubuisson & Deblauwe
* _Hymenophyllum sericeum_ (Sw.) Sw.
* _Hymenophyllum serrulatum_ (C. Presl) C. Chr.
* _Hymenophyllum seselifolium_ C. Presl
* _Hymenophyllum sibthorpioides_ (Bory ex Willd.) Mett. ex Kuhn
* _Hymenophyllum sieberi_ (C. Presl) Bosch
* _Hymenophyllum silvaticum_ C. V. Morton
* _Hymenophyllum silveirae_ Christ
* 寬片膜蕨 _Hymenophyllum simonsianum_ Hook.
* _Hymenophyllum simplex_ C. V. Morton
* _Hymenophyllum sodiroi_ C. Chr.
* _Hymenophyllum soriemersum_ Rouhan & C. Del Rio
* _Hymenophyllum splendidum_ Bosch
* 撕苞蕗蕨 _Hymenophyllum stenocladum_ (Ching & P. S. Chiu) K. Iwats.
* _Hymenophyllum subdemissum_ Christ
* _Hymenophyllum subobtusum_ Rosenst.
* _Hymenophyllum subrigidum_ Christ
* _Hymenophyllum superbum_ Morton
* 華東膜蕨 _Hymenophyllum taiwanense_ (Tagawa) C. V. Morton
* _Hymenophyllum talamancanum_ A. Rojas
* _Hymenophyllum taliabense_ Alderw.
* _Hymenophyllum tarapotense_ Stolze
* _Hymenophyllum tayloriae_ Farrar & Raine
* _Hymenophyllum tegularis_ (Desv.) Proctor & Lourteig
* _Hymenophyllum tenellum_ (Jacq.) Kuhn
* _Hymenophyllum tenerum_ Bosch
* _Hymenophyllum thuidium_ Harr.
* _Hymenophyllum todjambuense_ Kjellb.
* _Hymenophyllum tomaniiviense_ (Brownlie) Ebihara & K. Iwats.
* _Hymenophyllum tomentosum_ Kunze
* _Hymenophyllum torricellianum_ Alderw.
* _Hymenophyllum tortuosum_ Hook. & Grev.
* _Hymenophyllum trapezoidale_ Liebm.
* _Hymenophyllum treubii_ Racib.
* _Hymenophyllum triangulare_ Baker
* _Hymenophyllum trichomanoides_ Bosch
* _Hymenophyllum trichophorum_ (Alderw.) Ebihara & K. Iwats.
* _Hymenophyllum trichophyllum_ Kunth
* _Hymenophyllum trifoliatum_ (L.) Proctor & Lourteig
* _Hymenophyllum tunbrigense_ (L.) Sm.
* _Hymenophyllum turquinense_ C. Sánchez
* _Hymenophyllum ulei_ Christ & Giesenh.
* _Hymenophyllum umbratile_ Diem & J. S. Licht.
* _Hymenophyllum undulatum_ (Sw.) Sw.
* _Hymenophyllum urbanii_ Brause
* _Hymenophyllum vacillans_ Christ
* _Hymenophyllum valvatum_ Hook. & Grev.
* _Hymenophyllum verecundum_ Morton
* _Hymenophyllum veronicoides_ C. Chr.
* _Hymenophyllum viguieri_ Tardieu
* _Hymenophyllum villosum_ Colenso
* _Hymenophyllum vittatum_ Copel.
* _Hymenophyllum walleri_ Maiden & Betche
* _Hymenophyllum whitei_ Goy
* _Hymenophyllum wilsonii_ Hook.
* 萊氏蕗蕨 _Hymenophyllum wrightii_ Bosch
* _Hymenophyllum zamboanganum_ (Copel.) comb. ined.

**雜交種：**

* _Hymenophyllum × scopulorum_ Rumsey & F. J. Roberts
* _Hymenophyllum × tucuchense_ (Jermy & T. G. Walker) C. D. Adams & Baksh.-Com.

## 參考資料