ヴェロ・コプナー・ウィン＝エドワーズ
Wynne-Edwards, V.C. 1962. Animal Dispersion in Relation to Social Behavior. Oliver & Boyd, London.
Wynne-Edwards, V.C. 1986. Evolution through group selection. Oxford : Blackwell Scientific. 340 pp. ISBN 0632015411 (paperback)