勇蝸科

勇蝸科（學名：Helicarionidae），又名鱉甲蝸牛科，鱉甲螺科，是一個由陸生肺螺類半蛞蝓組成的科，皆為真有肺類柄眼類支序的成員，屬於勇蝸總科（Helicarionoidea）。

## 分佈

本科物種分佈於古北界東部，馬爾加什（馬達加斯加、留尼旺及葛摩）、印度、東南亞及澳大利亞。

## 解剖

本科的蝸牛都會在交配時伸出幾丁質的交配器戀矢（love dart）插進對方的生殖孔，以交換精子。

此外，本科物種的單倍體染色體介乎21到30之間。

## 下級分類

本科包括以下屬：

Advena｜Gude, 1913

Aenigmatoconcha｜C.Tumpeesuwan & S.Tumpeesuwan, 2017

- Aenigmatoconcha clivicola C.Tumpeesuwan & S.Tumpeesuwan, 2017
- Aenigmatoconcha eunetis Pholyotha & Panha, 2021
- Aenigmatoconcha mitis (L.Pfeiffer, 1863)
- Aenigmatoconcha sumonthai C.Tumpeesuwan & S.Tumpeesuwan, 2018

Amenixesta｜Iredale, 1941

- Amenixesta fifensis Iredale, 1941
- Amenixesta hunsteini (E.A.Smith, 1887)

Antiquarion｜Stanisic, 2010

- Antiquarion boondah Hyman & Köhler, 2020
- Antiquarion chooreechillum Hyman & Köhler, 2020
- Antiquarion collinsae J.Stanisic, 2010
- Antiquarion evelynensis Hyman & Köhler, 2020
- Antiquarion limbus Hyman & Köhler, 2020
- Antiquarion occultus Hyman & Köhler, 2020
- Antiquarion ravenshoe Hyman & Köhler, 2020
- Antiquarion swausi Hyman & Köhler, 2020

Apothapsia｜D.T.Holyoak & G.A.Holyoak, 2020

Attenborougharion｜Hyman & Köhler, 2017

奧氏蛞蝓屬｜Austenia｜G.Nevill, 1878

Bathia｜Robson, 1914

Belopygmaeus｜Solem, 1989

Belopygmeus｜Baker, 1938

Brevisentis｜Hyman, 2007

Buffetia｜Iredale, 1945

Burmochlamys｜Pholyotha & Panha, 2022

Burnettia｜Stanisic, 2010

Caldwellia｜H.Adams, 1873

Caracolla｜Ehrenberg, 1831

疑難螺屬｜Chalepotaxis｜Ancey, 1887

- 黑線小鱉甲蝸牛 Chalepotaxis spadix cinctus

Colmanarion｜Stanisic, 2010

隱奧氏蛞蝓屬｜Cryptaustenia｜Cockerell, 1891

Ctenoglypta｜Ancey, 1904

Ctenophila｜Ancey, 1882

Cucullarion｜Stanisic, 1998

Dancea｜Zilch, 1960

Dedrolamellaria

Delinitesta｜Iredale, 1933

Dendrolamellaria｜Preston, 1913

Dendronitor｜Iredale, 1933

Dupontia｜Godwin-Austen, 1908

小杜螺屬｜Durgella｜W.T.Blanford, 1863

- Durgella aborense Godwin-Austen, 1916
- Durgella assamica Gondwin-Austen, 1881
- Durgella birmanica (Pfeiffer, 1847)
- Durgella christianae (Theobald, 1864)
- Durgella concinna Blanford & Godwin-Austen, 1908
- Durgella densestriata Vermeulen, T.-S.Liew & Schilthuizen, 2015
- Durgella dwipana (Gude, 1903)
- Durgella edeana Godwin-Austen, 1910
- Durgella erratica (Godwin-Austen, 1888)
- Durgella hosei Godwin-Austen, 1891
- Durgella khasiaca Godwin-Austen, 1883
- Durgella kumahensis (Theobald & Stoliczka, 1872)
- Durgella levicula (W.H.Benson, 1859)
- Durgella libas Solem, 1966
- Durgella mairangensis Godwin-Austen, 1898
- Durgella naharaniensis Godwin-Austen, 1914
- Durgella nulla Pholyotha & Panha, 2024
- Durgella pentata Pholyotha & Panha, 2024
- Durgella rhaphiellus (E.von Martens, 1867)
- Durgella rogersi Godwin-Austen, 1907
- Durgella russeola (Morelet, 1865)
- Durgella salius (W.H.Benson, 1859)
- Durgella seposita (W.H.Benson, 1859)
- Durgella siamensis Möllendorff, 1902
- Durgella sundana B.Rensch, 1930

Durgellina｜Thiele, 1928

Echonitor｜Iredale, 1937

Eddiella｜Stanisic, 2010

Einasleighana｜Stanisic, 2010

Elatonitor｜Stanisic, 2010

Ellarion｜Iredale, 1941

Epiglypta｜Pilsbry, 1893

Erepta｜Albers, 1850

Eufretum｜H.B.Baker, 1941

Eurychlamys｜Godwin-Austen, 1899

Expocystis｜Iredale, 1937

Fastosarion｜Iredale, 1933

Fenestrarion｜Stanisic, 2010

Fijia｜Gude, 1913

吉拉蛞蝓屬｜Girasia｜Gray, 1855

Gudeconcha

Gudeoconcha｜Iredale, 1944

Haleakala｜Baker, 1940

Harmogenanina｜Germain, 1918

勇蝸屬｜Helicarion｜Férussac, 1821

Hiloaa｜Baker, 1940

Hiona｜Cooke, 1940

Hionarion｜Baker, 1940

Hionella｜Baker, 1940

Holkeion｜Godwin-Austen, 1908

Howearion｜Iredale, 1944

伊比蛞蝓屬｜Ibycus｜Heynemann, 1863

- 小玉蝸牛 Ibycus hiraseanus

Insulorbis｜Baker, 1940

Irenella｜Gude, 1913

Kalidos｜Gude, 1911

Kermarion｜Iredale, 1944

Kipua｜Baker, 1940

Laconia｜Gray, 1855

Laocaia｜Kuzminykh, 1999

Levidens｜Hyman, 2007

Limpidarion｜Hyman & Köhler, 2020

Luinarion｜Iredale, 1933

Malandena｜Iredale, 1933

Mauka｜Baker, 1940

Medyla｜Martens, 1860

Minororbis｜Baker, 1940

Mistarion｜Iredale, 1941

Montanocystis｜Stanisic, 2010

Muangnua｜Solem, 1966

Mysticarion｜Iredale, 1941

Nesaecia｜Gude, 1911

Nesarion｜Baker, 1940

Nesocyclus｜Baker, 1940

Nesonanina｜C.R.Boettger, 1918

Neutra｜Baker, 1940

Nitor｜Gude, 1911

Nukupiena｜Baker, 1940

Oafatua｜Baker, 1940

Opara｜Baker, 1940

圓鱉甲蝸牛屬｜Ovachlamys｜Habe, 1946

Pachystyla｜Mörch, 1852

Palmervillea｜Stanisic, 2010

Panulena

Parmacochlea｜E.A.Smith, 1884

Parmavitrina｜Iredale, 1937

Parmella｜H.Adams, 1867

Parmellops｜Iredale, 1944

Peloparion｜Iredale, 1937

Periclocystis｜Iredale, 1937

扁楯蛞蝓屬｜Petalochlamys｜Godwin-Austen, 1907，葉楯蛞蝓屬

Pitcairnia｜Baker, 1940

Pittoconcha｜Preston, 1913

Plegma｜Gude, 1911

Pravonitor｜Iredale, 1937

Pseudaustenia｜Cockerell, 1891

Pseudoguppya｜Baker, 1925

Pseudophasis｜Germain, 1918

Pseudosaphtia de｜Winter, 2008

Pseudosesara｜Solem, 1962

Quirosella｜Clench, 1958

Rapafila｜Baker, 1940

曙螺屬｜Rasama｜Laidlaw, 1932

Rhadella｜Godwin-Austen, 1914

Rhyssotopsis｜Ancey, 1887

若通螺屬｜Rotungia｜Godwin-Austen, 1918

Saphtia de｜Winter, 2008

Satiella｜W.T.Blanford & Godwin-Austen, 1908

Sesara｜Martens, 1860

Sheaia｜Hyman, 2007

Sigaloeista｜Shea & Griffiths, 2010

昔塔螺屬｜Sitala｜H.Adams, 1865

Sitalarion｜H.B.Baker, 1941

Sivella｜W.T.Blanford, 1863

Sodaleta｜Iredale, 1937

Sophina｜Benson, 1859

Stanisicarion｜Hyman & Ponder, 2010

Tarocystis｜Iredale, 1937

Teraia｜Solem, 1966

Thularion｜Stanisic, 1993

Torrecystis｜Stanisic, 2010

Tropicystis｜Stanisic, 2010

Uafatua｜Baker, 1940

Uapuneutra｜Baker, 1940

Ubiquitarion｜Hyman, Lamborena & Köhler, 2017

Urazirochlamys｜Habe, 1946

Vanmolia de｜Winter, 2008

Vercularion｜Iredale, 1933

Waihoua｜Baker, 1940

Westracystis｜Iredale, 1939

Zagmena｜Iredale, 1941

The following two subfamilies were recognized in the taxonomy of Bouchet & Rocroi (2005):

- 亞科 Helicarioninae Bourguignat, 1877 - synonyms: Pseudotrochatellinae A. J. Wagner, 1905; Ereptinae Godwin-Austen, 1908; Xestinae Gude & B. B. Woodward, 1921; Sesarinae Thiele, 1931; Nitoridae Iredale, 1937; Epiglyptidae Iredale, 1944; Gudeoconchidae Iredale, 1944
- 亞科 Durgellinae Godwin-Austen, 1888
- 族 Durgellini Godwin-Austen, 1888 - synonyms: Sitalinae Godwin-Austen, 1900; Sophininae W. Blanford & Godwin-Austen, 1908; Satiellini Schileyko, 2003
- 族 Girasiini Collinge, 1902

### 屬

下列本科所有的屬：

#### Durgellinae亞科

族 Durgellini

- 族 Girasiini

## 外部連結

- Helicarionidae
- Video of a Helicarionid in Sulawesi

## 基本資料

- 學名：Helicarionidae
- 命名者：Bourguignat, 1877