查爾斯·克拉克 (植物學家)

查爾斯·克拉克（Charles M. Clarke）是專業從事於豬籠草屬食蟲植物的植物學家與分類學家。克拉克出生於澳大利亞墨爾本。他獲得了墨爾本莫納什大學（Monash University）的生物學榮譽學位，同時還在新南威爾斯阿米代爾的新英格蘭大學（University of New England）取得了生態系統管理博士學位。

克拉克第一次到婆羅洲研究豬籠草是在1987年。1989年至1990年，他生活在汶萊以研究豬籠草屬植物的生態問題。在這兩次旅行間，克拉克還曾在昆士蘭州的詹姆斯·庫克大學（James Cook University）任教生態學和生物統計學，同時還在香港擔任園藝顧問。

克拉克現就任於莫納什大學馬來西亞分校。克拉克已完成了五本關於豬籠草屬的著作，其中最重要的是《婆羅洲的豬籠草屬植物》（Nepenthes of Borneo）和《蘇門答臘島與西馬來西亞的豬籠草》（Nepenthes of Sumatra and Peninsular Malaysia）。

克拉克描述了5個豬籠草屬物種，分別為本斯通豬籠草（Nepenthes benstonei）、陳氏豬籠草（Nepenthes chaniana）、泉氏豬籠草（Nepenthes izumiae）、賈桂琳豬籠草（Nepenthes jacquelineae）和堅韌豬籠草（Nepenthes tenax）。

## 著作

- Clarke, C.M. 1992. The ecology of metazoan communities in Nepenthes pitcher plants in Borneo, with special reference to the community of Nepenthes bicalcarata Hook.f. Ph.D. thesis, University of New England, Armidale, N.S.W. 269 pp.
- Clarke, C.M. 1993. Carnivorous Plant Newsletter 22(1–2): 27–28.
- Clarke, C.M. & R.L. Kitching 1993. The Metazoan Food Webs from Six Bornean Nepenthes Species. Ecological Entomology 18: 7–16.
- Clarke, C.M. & J.A. Moran 1994. A further record of aerial pitchers in Nepenthes ampullaria Jack. Malayan Nature Journal 47: 321–323.
- Clarke, C.M. & R.L. Kitching 1995. Swimming Ants and Pitcher Plants: a Unique Ant-Plant Interaction from Borneo. Journal of Tropical Ecology 11(4): 589–602.
- Clarke, C.M. 1997. Nepenthes of Borneo. Natural History Publications (Borneo), Kota Kinabalu. xi + 207 pp.
- Clarke, C.M. 1997. Another nice trip to Sumatra . Carnivorous Plant Newsletter 26(1): 4–10.
- Clarke, C.M. 1997. The effects of pitcher dimorphism on the metazoan community of the carnivorous pitcher plant Nepenthes bicalcarata Hook.f. Malayan Nature Journal 50: 149–157.
- Clarke, C.M. 1998. Initial colonisation and prey capture in Nepenthes bicalcarata (Nepenthaceae) pitchers in Brunei. Sandakania 12: 27–36.
- Clarke, C.M. 1998. The aquatic arthropod community of the pitcher plant, Nepenthes bicalcarata (Nepenthaceae) in Brunei. Sandakania 11: 55–60.
- Clarke, C.M. 1998. A re-examination of geographical variation in Nepenthes food webs. Ecography 21: 430–436.
- Clarke, C.M. 1999. Nepenthes benstonei (Nepenthaceae), a new pitcher plant from Peninsular Malaysia. Sandakania 13: 79–87.
- Clarke, C.M. 2001. Ecology & Conservation of Montane Nepenthes (Nepenthaceae) in Sumatra. Fourth International Flora Malesiana Symposium. 20-24 July 1998, Kuala Lumpur.
- Clarke, C.M. 2001. Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu. ix + 325 pp.
- Clarke, C.M. 2001. A Guide to the Pitcher Plants of Sabah. Natural History Publications (Borneo), Kota Kinabalu. iv + 40 pp.
- Moran, J.A., M.A. Merbach, N.J. Livingston, C.M. Clarke & W.E. Booth 2001. Annals of Botany 88: 307–311.
- Clarke, C.M. 2002. A Guide to the Pitcher Plants of Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu. iv + 32 pp.
- Clarke, C.M., T. Davis & R. Tamin 2003. Nepenthes izumiae (Nepenthaceae): a new species from Sumatra. Blumea 48: 179–182.
- Moran, J.A., C.M. Clarke & B.J. Hawkins 2003. From Carnivore to Detritivore? Isotopic Evidence for Leaf Litter Utilization by the Tropical Pitcher Plant Nepenthes ampullaria. International Journal of Plant Sciences 164: 635–639.
- Dong, T.T.X., Q.M. Xiao, C.M. Clarke, H.S. Zhong, N.J. Zhao, K.L. Chun & K.W.K. Tsim 2003. Phylogeny of Astragalus in China: Molecular evidence from the DNA sequences of 5S rRNA spacer, ITS, and 18S rRNA. Journal of Agricultural and Food Chemistry 51(23): 6709–6714.
- Clarke, C.M. & C.C. Lee 2004. A Pocket Guide: Pitcher Plants of Sarawak. Natural History Publications (Borneo), Kota Kinabalu. vi + 81 pp.
- Cantley, R., C.M. Clarke, J. Cokendolpher, B. Rice & A. Wistuba 2004. Nepenthes clipeata Survival Project. International Carnivorous Plant Society.
- Clarke, C.M., C.C. Lee & S. McPherson 2006. Nepenthes chaniana (Nepenthaceae), a new species from north-western Borneo. Sabah Parks Journal 7: 53–66.
- Clarke, C.M & R. Kruger 2006. Nepenthes tenax C.Clarke and R.Kruger (Nepenthaceae), a new species from Cape York Peninsula, Queensland. Austrobaileya 7(2): 319–324.
- Clarke, C.M. 2006. Introduction. In: Danser, B.H. The Nepenthaceae of the Netherlands Indies. Natural History Publications (Borneo), Kota Kinabalu. pp. 1–15
- Clarke, C.M., U. Bauer, C.C. Lee, A.A. Tuen, K. Rembold & J.A. Moran 2009. Biology Letters 5(5): 632–635. 10.1098/rsbl.2009.0311
- Chin, L., J.A. Moran & C. Clarke 2010. Trap geometry in three giant montane pitcher plant species from Borneo is a function of tree shrew body size. New Phytologist, published online January 22, 2010. 10.1111/j.1469-8137.2009.03166.x

## 擴展閱讀

- Clarke, C.M. 1997. Nepenthes of Borneo. Natural History Publications (Borneo), Kota Kinabalu. xi + 207 pp. ISBN 983-812-057-X

## 基本資料

- 民族：ALL