瘤拟步行虫科

瘤拟步行虫科（学名：Zopheridae）是属于鞘翅目拟步总科的昆虫。近年来，有其他两个科被划分到本科之中，使得本科的成员数量有了相当的成长；这两个科分别是拟吉丁虫科（Monommatidae）以及细坚虫科（Colydiidae）。某些学者接受本科划分了6个亚科的系统，然而有些则将细坚虫科从本科中剔除为一独立科。本科有超过100个属群需要被重新定义；据估计在世界各地有着数百个种类。这个科没有固定使用的英文俗名，不过多数亚科都有各自传统的英文俗名，例如 ironclad beetles、cylindrical bark beetles等。至2018年为止，约有116个物种已被描述。

## 系统分类学

目前所使用的瘤拟步行虫科级分类元定义主要来自史利宾斯基和劳伦斯于 1999 年 基于成虫和幼生期形态所做的拟步行虫总科系统发育分析，近年的分子系统发育分析研究则显示本科可能并非单系群。

本科包括以下属：

Ablabus｜Broun, 1880

Acolobicus｜Sharp, 1894

Acolophoides｜Slipinski & Lawrence, 1997

Acolophus｜Sharp, 1885

Acostonotus｜Slipinski & Lawrence, 1997

Acropis｜Burmeister, 1840

Afrorthocerus｜Pope, 1959

Allobitoma｜Broun, 1921

Alluauditoma｜Dajoz, 1980

Anarmostes｜Pascoe, 1860

Anisopaulax｜Reitter, 1877

Anosyana｜Dajoz, 1980

Antilissus｜Sharp, 1879

Antillemonomma｜Freude, 1955

Aplanetes｜Sharp, 1899

Aprostoma｜Guérin-Méneville, 1839

Ascomma｜Waterhouse, 1876

Aspathines｜Champion, 1888

Aspinatimonomma｜Freude, 1955

Asprotera｜Pascoe, 1860

Asynchita｜Sharp, 1894

Atyscus｜Pascoe, 1863

Aulonium｜Erichson, 1845

Bhutania｜Dajoz, 1975

Bitoma｜Herbst, 1793

Bolcocius｜Dajoz, 1977

Bulasconotus｜Slipinski & Lawrence, 1997

- Bulasconotus scaccarius Bernard & Gillett, 2020

Bupala｜Pascoe, 1863

Caanthus｜Champion, 1894

Cacotarphius｜Sharp, 1894

Canaricerus｜Oromi & Garcia, 1987

Caprodes｜Pascoe, 1863

Carbothrus｜Aoki, 2012

Catolaemus｜Sharp, 1894

Cebia｜Pascoe, 1863

Cerchanotus｜Erichson, 1845

Chorasus｜Sharp, 1882

Cicablabus｜Slipinski & Lawrence, 1997

Ciconissus｜Broun, 1893

Cleteomonomma｜Freude, 1957

Cleteus｜Fairmaire, 1906

Coeloderes｜Mulsant & Rey, 1859

Colobicones｜Grouvelle, 1918

Colobicus｜Latreille, 1807

Colydium｜Fabricius, 1792

Colydodes｜Motschulsky, 1855

Corticus｜Latreille, 1829

Coslonatus｜Li & Cai, 2021

Cotulades｜Pascoe, 1860

Coxelus｜Dejean, 1821

Cretomysteria｜Deng, Ślipiński, Ren & Pang, 2017

Curtemonomma

Dechomus｜Jacquelin Du Val, 1859

Denophloeus｜Stephan, 1989

Denophoelus｜Stephan, 1989

Diodesma｜Latreille, 1829

Diplagia｜Reitter, 1882

Diplotoma｜Erichson, 1845

Ditomoidea｜Grouvelle, 1906

Docalis｜Pascoe, 1860

Doderonymus｜Binaghi, 1937

Dryptops｜Broun, 1882

Emilka｜Slipinski, 1982

Enarsus｜Pascoe, 1866

Endeitoma｜Sharp, 1894

Endestes｜Pascoe, 1863

Endocoxelus｜Waterhouse, 1876

Endophloeus｜Dejean, 1834

Enhypnon｜Carter, 1919

Epistranodes｜Slipinski & Lawrence, 1997

Epistranus｜Sharp, 1878

Erylus｜Dajoz, 1969

Eucicones｜Sharp, 1894

Eudesma｜LeConte, 1863

Faecula｜Slipinski & Lawrence, 1997

Fenerivia｜Dajoz, 1980

Franzorphius｜Schuh, 1998

Gathocles｜Broun, 1893

Gempylodes｜Pascoe, 1863

Glenentela｜Broun, 1893

Glyphocryptus｜Sharp, 1885

Grandemonomma｜Freude, 1957

Heterargus｜Sharp, 1886

Holopleuridia｜Reitter, 1876

Hyberis｜Pascoe, 1860

Hybonotus｜Slipinski & Lawrence, 1997

Hyboonotus｜Slipinski & Lawrence, 2016

Hyporhagus｜Dejean, 1833

Hyporhagus｜Thomson, 1860

Hyporrhagus｜Thomson, 1860

Hystricones｜Sharp, 1894

Inscutomonomma｜Pic, 1951

Insulcicleteus｜Freude, 1957

Isotarphius｜Heller, 1916

Kanantsia｜Dajoz, 1980

Kraatziana｜Pic, 1934

Labrotrichus｜Sharp, 1894

Langelandia｜Aubé, 1842

Lascobitoma｜Slipinski & Lawrence, 1997

Lasconotus｜Erichson, 1845

Lascotonus｜Grouvelle, 1895

Lascotrichus｜Pope, 1961

Lastrema｜Reitter, 1882

Latometus｜Erichson, 1842

Lemmis｜Pascoe, 1860

Lepidosynchis｜Kolbe, 1897

Linophloeus｜Dajoz, 1980

Lobogestoria｜Reitter, 1878

Lobomesa｜Slipinski & Lawrence, 1997

Lyreus｜Aubé, 1861

Madacones｜Dajoz, 1980

Madadesia｜Dajoz, 1986

Madenphloeus｜Dajoz, 1980

Mamakius｜Pope, 1961

Mauritianopidium｜Dajoz, 1977

Mecedanum｜Erichson, 1845

Megataphrus｜Casey, 1890

Metopiestes

Micromonomma｜Pic, 1933

Microprius｜Fairmaire, 1868

Microsicus｜Sharp, 1894

Microvonus

Monoedus｜Horn, 1882

Monomma｜Klug, 1833

Munaria｜Reitter, 1882

Myrmecoxenus

Namunaria｜Reitter, 1882

Nematidium｜Erichson, 1845

Neotrichus｜Sharp, 1886

Niphopelta｜Reitter, 1882

Norix｜Broun, 1893

Noserinus｜Casey, 1907

Nosoderma｜Guérin-Méneville, 1838

Nosodomodes｜Reitter, 1922

Notocoxelus｜Slipinski & Lawrence, 1997

Notorthocerus｜Slipinski & Lawrence, 1999

Opostirus｜Kirsch, 1865

Orthocerodes｜Slipinski & Lawrence, 1999

Orthocerus｜Latreille, 1796

Orvoenia｜Dajoz, 1980

Paha｜Dajoz, 1984

Papuatrichus｜Aoki, 2013

Paratarphius｜Dajoz, 1971

Paryphus｜Erichson, 1845

Paxillobitoma｜Lord & Ivie, 2016

Phaennis｜Champion, 1894

Pharax｜Pascoe, 1860

Phellopsis｜LeConte, 1862

Phloeodalis｜Erichson, 1845

Phloeodes｜LeConte, 1862

Phloeonemus｜Erichson, 1845

Phloeopsidius｜Gebien, 1925

Phormesa｜Pascoe, 1863

Phorminx｜Carter & Zeck, 1937

Phreatus｜Pascoe, 1863

Plagiope｜Erichson, 1845

Priolomopsis｜Dajoz, 1982

Priolomus｜Erichson, 1845

Pristoderes

Pristoderus｜Hope, 1840

Prostheca｜Wollaston, 1860

Protarphius｜Broun, 1893

Pseudaulonium｜Reitter, 1877

Pseudendestes｜Lawrence, 1980

Pseudocorticus｜Hinton, 1935

Pseudotarphius｜Wollaston, 1873

Pycnomerodes｜Broun, 1886

Pycnomerus｜Erichson, 1842

Rechodes｜Erichson, 1845

Recyntus｜Broun, 1882

Rhagodera｜Mannerheim, 1843

Rhagoderidea｜Wickham, 1914

Rhizonium｜Sharp, 1876

Rhopalocerus｜Redtenbacher, 1842

Rytinotus｜Broun, 1880

Sallachius｜Pope, 1959

Sasajia｜Masumoto & Akita, 2007

Sassaka｜Pope, 1961

Scoriaderma｜Fairmaire, 1894

Sechellotoma｜Dajoz, 1980

Semicoxelus｜Alekseev & Pankowski, 2020

Sesaspis

Sparactus｜Erichson, 1845

Spinatimonomma｜Freude, 1955

Spinhyporhagus｜Freude, 1984

Sprecodes｜Pope, 1961

Stenomonoedus｜Heinze, 1954

Stephaniolus｜Ivie, Slipinsky & Wegrzynowicz, 2001

Sympanotus｜Sharp, 1885

Synagathis｜Carter & Zeck, 1937

Synchita｜Hellwig, 1792

Tarphiablabus｜Slipinski & Lawrence, 1997

Tarphiomimus｜Wollaston, 1873

Tarphiosoma｜Wollaston, 1862

Tarphius｜Erichson, 1845

Tentablabus｜Slipinski & Lawrence, 1997

Todima｜Grouvelle, 1893

Todimopsis｜Slipinski & Lawrence, 1997

Trachypholis｜Erichson, 1845

Trigonophloeus｜Dajoz, 1980

Uloncrus

Usechimorpha｜Blaisdell, 1929

Usechus｜Motschulsky, 1845

Verodes｜Casey

Vitiacus｜Broun, 1893

Xylolaemus｜Dejean, 1835

Xylolaemus｜Redtenbacher, 1857

Zanclea｜Pascoe, 1863

Zopher｜Slipinski & Lawrence, 1999

Zopherosis｜White, 1859

瘤拟步行虫属｜Zopherus｜Laporte, 1840

## 古昆虫学

目前所知的瘤拟步行虫科化石纪录相当稀少，多数化石纪录来自波罗的海琥珀（始新世），最早的化石纪录为来自上白垩纪缅甸琥珀的 Paleoendeitoma antennata、Paleoendeitoma minuta（属于细坚虫亚科 Colydiinae）和 Cretomysteria burmanica （亚科未定论）。

## 基本资料

- 说明：Ironclad beetle,（Zopherus nodulosus haldemani），摄于美国德州
- 母公司：3
- 学名：Zopheridae
- 命名者：安托万·约瑟夫·简·索里尔, 1834
- 栽培群成员：Colydiinae - cylindrical bark beetles、Latometinae (sometimes in Zopheridae)、Monommatinae (sometimes in Zopheridae)、Pycnomerinae (sometimes in Zopheridae)、Usechinae (sometimes in Zopheridae)、Zopherinae - ironclad beetles
- subdivision_ranks：亚科