褶颊虫目

褶颊虫目（学名：Ptychopariida）是三叶虫纲下的其中一目，出现于寒武纪，包括的三叶虫种类相当的繁多。

褶颊虫类拥有较大的胸部，通常是由8个或更多的环节组成的，而且褶颊虫类的胸部通常是比尾部还要长的，它们的尾部是比较小型的。

## 下属分类

本科包括以下属：

Acrocephalitidae Agraulidae Alokistocaridae

无序虫科｜Antagmidae

Asaphiscidae Atopidae Aulacodigmatidae Avoninidae Bigotinidae Bolaspididae Catillicephalidae Cedariidae Changshaniidae Conocoryphidae Conokephalinidae Coosellidae Crepicephalidae Dokimocephalidae Ellipsocephalidae Ellipsocephaloididae Elviniidae Emmrichellidae Estaingiidae Eulomatidae Eulomidae Holanshaniidae Housiidae Hypermecaspididae Ignotogregatidae Inouyiidae Isocolidae Ityophoridae Kingstoniidae Komaspididae Lisaniidae Llanoaspididae Loganellidae Lonchocephalidae Lorenzellidae Luaspididae Mapaniidae Menomoniidae Nepeidae Norwoodiidae Olenidae Onchonotopsidae Palaeolenidae Paleo Papyriaspididae Phylacteridae Plethopeltidae Proasaphiscidae Ptychopariidae Ptychoparioidae Raymondinidae Shumardiidae Tengfengiidae Thysanopeltidae Tricrepicephalidae Utiidae Wuaniidae Yunnanocephalidae

科的地位未定的属：

Acanthometopus｜Lu & Qian, 1983

Aedotes｜Öpik, 1967

Afghanocare｜Wolfart, 1975

Agelagma｜Öpik, 1967

Agraulopsis｜Ružicka, 1940

Aiaiaspis｜Nazarov, 1973

Alceste｜Hawle & Corda, 1847

Amicus｜Koroleva, 1967

Amphorella｜Rozova, 1963

Anemocephalops｜Ivshin, 1962

Angustaeva

Annamitella｜Mansuy, 1916

Anopocodia｜Öpik, 1967

Apatokephaloides｜Raymond, 1924

Apatokephalops｜Lu, 1975

Aplotaspis｜Henderson, 1976

Apomodocia｜Hu, 1971

Aristoharpes｜Whittington, 1950

Arraphus｜Angelin, 1854

Asaphoon｜Hutchison & Ingham, 1967

Asiocephalus｜Palmer, 1968

Aulacodigma｜Öpik, 1967

Austosinia Badainjaranspis Batchatiella

Berkutaspis｜Lisogor, 1977

Bestjubella｜Ivshin, 1962

Bicella｜Rasetti, 1955

Bifodina｜Robison & Pantoja-Alor, 1968

Blosyropsis｜Whittington, 1965

Botrioides｜Stetson, 1927

Bowmania｜Walcott, 1925

Bucksella｜Howell, 1957

Bulnibarbi Bvalbardites

Bynumiella｜Resser, 1942

Cambroproteus｜Geyer, 2015

Carrickia｜Tripp, 1965

Catasolenopleura｜Qiu, 1983

Cedarapis

Changqingia｜Lu & Zhu, 1983

Chencunia Chondrastaulina

Chosenia｜Kobayashi, 1934

Chuangiformis

Ciliocephalus｜Liu, 1977

Ciliscutellum｜Linnaeus, 1987

Circulocrania｜Fortey & Owens, 1987

Conococheaquea｜Rasetti, 1959

Corbina

Craigheadia｜Hupé, 1955

Croixana｜Nelson, 1951

Crusoia Damaiaoaspis

Dentaloscutellum｜Chatterton, 1971

Dipyrgotes｜Öpik, 1967

Dorypygella｜Walcott, 1905

Dubhglasina

Ejinaspis｜Lu, Zhou & Zhou, 1986

Elatius｜Orlowski, 1985

Elegantaspis｜Ivshin, 1962

Elkanaspis｜Ludvigsen, 1982

Emmrichops｜Marek, 1961

Eokosovopeltis｜Pribyl & Vanek, 1972

Epichalnipsus｜Geyer, Popp, Weidner & Förster, 2004

Erbiella｜Fedyanina, 1962

Erixanium｜Öpik, 1963

Eshelmania

Faciura｜Rozova, 1963

Fatocephalus｜Duan & An, 1982

Forchhammeria｜Lermontova & Chernysheva, 1950

Furcalithus｜Fortey & Owens, 1987

Gastropolus｜Whittard, 1966

Girvanopyge｜Kobayashi, 1960

Gloriagloria｜Jell, 2003

Groenwallia Groenwallina

Gymnostomix｜Fortey & Owens, 1987

Hadrocephalites｜Sundberg & McCollum, 2002

Hanivella｜Sivov, 1955

Harpesoides｜Koroleva, 1978

Harpidoides｜Petrunina, Apollonov & Tschugaeva, 1983

Hartshillina｜Lake, 1940

Helioharpes｜Pribyl & Vanek, 1981

Heptabronteus｜Webby, 1974

Hibbertia｜Jones & Woodward, 1899

Hidascutellum｜Kobayashi & Hamada, 1977

Holmdalia｜R.A.Robison, 1988

Huaiaspis｜Li & Peng, 2006

Hunanopyge｜Peng, 1984

Hyperbolochilus｜Ross, 1951

Hyperoparia｜Zhang, 1983

Idiorapha

Iohomia｜Hu, 1971

Irvingelloides｜Kobayashi, 1935

Ithyektyphus｜Shaw, 1956

Janxilithus

Jiia｜Zhou Zhiyi & Zhang Jinlin, 1978

- Jiia talowaois

Kabukiia｜Zhang, 1987

Karagandoides｜Ivshin, 1983

Kathleenella｜Ludvigsen, 1983

Kathrynia｜Westrop, 1986

Kaufmannella｜Sdzuy, 1979

Kazelina

Kepisis｜Hu, 1971

Kielania｜Vanek, 1963

Kirengelia

Kirkdomina｜Tripp, 1962

Kodymaspis｜Prantl & Pribyl, 1950

Komaspis｜Kobayashi, 1935

Kulyambopeltis

Lacorsalina｜Bursky, 1970

Lampropeltis｜Öpik, 1967

Langyashania｜Lu & Zhu, 1980

Lauzonella｜Rasetti, 1944

Lecanoaspis｜Ivshin, 1983

Leioscutellum｜Wu, 1977

Leocephalus｜Ivshin, 1983

Liocare｜Suvorova, 1981

Liopelta｜Suvorova, 1981

Liosolenopleura｜Thoral, 1947

Litaspis｜Suvorova, 1960

Lobocephalina｜Ružicka, 1940

Lochmanaspis｜Ivshin, 1983

Loganopeltis｜Rasetti, 1943

Lordshillia｜Whittard, 1966

Lorensella

Lunacephalus｜Ivshin, 1962

Luottuolingia

Lydiaspis｜Suvorova, 1981

Macronoda｜Lochman, 1964

Magnacephalus｜Stitt, 1971

Maladia

Malinaspis｜Andreeva, 1985

Malongullia｜Webby, Moors & McLean, 1970

Manchurocephalus｜Endo, 1944

Megalopsis｜Qian, 1983

Mekynophrys｜Harrington, 1938

Meniscocorphye

Meniscopsia｜Robison & Babcock, 2011

Menocephalites｜Kobayashi, 1935

Meroperix｜Lane, 1972

Metabowmania｜Kobayashi, 1955

Metaharpes｜Lamont, 1948

Miaopopsis｜Lu, 1965

Minicephalus｜Lochman & Hu, 1960

Minkella｜Lochman & Hu, 1959

Moxomia｜Walcott, 1924

Munijia

Nahannia｜Chatterton & Ludvigsen, 1976

Nambeetella｜Legg, 1976

Naustia｜Ludvigsen, 1982

Neoacrocephalites｜Sivov, 1955

Neoanomocerella｜Xiang, 1963

Neohedinaspis｜Xiang & Zhang, 1984

Neoshirakiella｜Sun, 1979

Neosolenopleurella｜Ju, 1983

Neotaenicephalus｜Kobayashi, 1955

Nilcgna

Noelaspis｜Ludvigsen, Westrop & Kindle, 1989

Novakella｜Whittard, 1961

Octobronteus｜Weber, 1945

Oculeus｜Poletaeva & Romanenko, 1970

Onchonotina｜Lu, 1964

Onchonotopis

Oopsites｜Fortey, 1975

Opsupharus

Oreisator｜Geyer & Malinky, 1997

Parabathycheilus｜Mergl, 1984

Paracrocephalites｜Poulsen, 1954

Paraharpes｜Whittington, 1950

Parairvingella｜Kobayashi, 1938

Parakingasidoides

Paralorenzangella｜Jell, 2003

Paranumia

Parapeishania｜Qiu, 1983

Paraplethopeltis｜Bridge & Cloud, 1947

Paratermierella｜Hupé, 1953

Paratretaspis｜Dean, 1973

Pardailhania｜Thoral, 1947

Parkesolithus｜Campbell & Durham, 1970

Patronaspis｜Ivshin, 1983

Pedinaspis｜Ivshin, 1983

Pelicephalus｜Kurtz, 1975

Perioura｜Resser, 1938

Perischoclonus｜Raymond, 1925

Pesaiella｜Rozova, 1964

Petruninaspis｜Ivshin, 1983

Phorocephala｜Lu, 1965

Pinglusapis

Placosema｜Öpik, 1967

Platyharpes｜Whittington, 1950

Plectrella｜Ludvigsen, Westrop & Kindle, 1989

Polycertaspis Populsenella

Portaginus｜Lesperance, 1968

Princetonella｜Lochman, 1953

Proapatokephaloides｜Ergaliev, 1980

Proaulacopleura｜Kobayashi, 1936

Promesus｜Lochman, 1964

Propatokephalops Propeltura

Proricephalus｜Westrop, 1986

Prospectatrix｜Fortey, 1981

Protostygina｜Prantl & Pribyl, 1949

Pseudoalokistocare｜Sundbe, 1994

Pseudokingstonia｜Palmer, 1965

Pseudokoldinia｜Rozova, 1968

Pseudomegalaspis｜Jaanusson, 1953

Ptychoparoides｜Rka, 1940

Ptychopleurites｜Kobayashi, 1936

Pulchricapitus｜Kurtz, 1975

Pytine｜Fortey, 1975

Pyttstrigis

Rampartaspis｜Loch, 1993

Randicephalus｜Ivshin, 1983

Rasettaspis｜Ivshin, 1983

Reticuloharpes｜Vanek, 1963

Reuscholithus｜Bancroft, 1929

Rimouskia｜Resser, 1938

Sagavia｜Koroleva, 1967

Sagitaspis｜Ivshin, 1983

Sagitoides｜Ivshin, 1983

Salteria｜Thomson, 1864

Sanaschtykgolia｜Poletaeva, 1960

Sangzhiscutellum｜Linnaeus, 1987

Schyilaspis｜Ivshin, 1983

Seletella｜Ivshin, 1983

Seletoides｜Ivshin, 1983

Shenjiawania｜Peng, 1984

Shickshockia｜Rasetti, 1946

Sinoharpes｜Sheng, 1974

Songtaoia｜Yin, 1978

Stegnopsis｜Whittington, 1965

Stenorhachis｜Hintze & Jaanusson, 1956

Striticephala

Sunwaptia｜Westrop, 1986

Syspacheilus｜Resser, 1938

Szechuanella｜Cheng Wen-t'ang, 1960

Taishania｜Sun, 1935

Taoyuania｜Liu, 1977

Tatonaspis｜Kobayashi, 1935

Tonopahella｜Sundberg & McCollum, 2003

Torifera｜Wolfart, 1974

Torridella｜Lochman & Hu, 1962

Tosotychia｜Öpik, 1961

Triarthrella｜Hall, 1863

Troedssonia｜Poletaeva, 1960

Tropidopyge｜Harrington & Kay, 1951

Trymataspis｜Robison, 1964

Tuberaspis

Turgicephalus｜Fortey, 1980

Tuvanella｜Pokrovskaya, 1959

Uxunella｜Romanenko, 1985

Verkholenella｜Vasilenko, 1965

Vermilionites｜Kobayashi, 1955

Wilcoxaspis｜Westrop, 1986

Xianquianaspis

Xiushuilithus｜Zhou, 1976

Yinpanolithus｜Lu & Chang, 1974

Yosimuraspis｜Kobayashi, 1960

Yuehsieszella｜Zhang, 1963

Yuknessaspis｜Rasetti, 1951

Yurakia｜Rozova, 1963

Zdicella｜Šnajdr, 1957

## 基本资料

- fossil_range：520寒武纪早期-奥陶纪晚期
- 说明：Elrathia kingii
- 学名：Ptychopariida
- 命名者：1915
- subdivision_ranks：亚目
- 栽培群成员：褶颊虫亚目 Ptychopariina、椭圆头虫总科 Ellipsocephaloidea、褶颊虫总科 Ptychoparioidea、油栉虫亚目 Olenina、亚目未定、Raymondinacea总科