龙虱科

“龙虱”也可以指小型单人龙舟。

龙虱科（学名：Dytiscidae），亦作龙虱科，为水甲虫的一种，隶属于鞘翅目之下的肉食亚目。生活在田野、水沟、小溪等水体中，是一种掠食性水生昆虫。其学名源于希腊文δυτικός（dytikos），意思就是“肉食性的潜水甲虫”。本科物种几乎见于全世界所有淡水栖境，但亦有部分于落叶堆中栖息。成虫体长一般在1and(-)之间，尽管物种间的差异会比较大。当中见于欧洲的及巴西的有最大的体型，可分别长至4.5cm到4.75cm。与之相反，拥有最小体型的物种应当为栖息于澳大利亚地下水中的 atypicali，只有约0.9mm长。

大多数物种的颜色都在深棕色、黑色或深橄榄绿，某些亚科的物种会在其边沿带有金黄色的条状斑纹。

## 食用

本科物种的成虫，特别是Cybister属的物种，均为可食用昆虫。在美国内华达州洪堡干湖的一个山洞内，还在当地史前人类遗下的粪化石内发现有被食用的残骸。在墨西哥，他们在吃C. explanatus时会以墨西哥夹饼夹着来吃。在日本多个地区（例如：长野县）有以日本龙虱（Cybister japonicus）为食的习惯；在中华人民共和国广东省，除了日本龙虱以外，亦会食用C. bengalensis、C. guerini、C. limbatus、C. sugillatus、三点龙虱等物种。很可能也包括在内，但这个物种的肉很少。也有报告指在台湾、泰国及新几内亚亦有食用本科物种的习惯。

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大但在陆地上很慢，成年后也不是特别凶猛，它们也被许多中型鸟类、哺乳动物和其他大型捕食者津津有味地吃掉。幼虫通常比较安全，由于它们的保护色和通过水柱逃跑的能力；它们可能相当难抓，可能成为小池塘的顶级掠食者。

## 系统分类

本科包括下列各属：

Acilius｜Leach, 1817

Aethionectes｜Sharp, 1882

Africodytes｜Biström, 1988

Africophilus｜Guignot, 1948

Agabetes｜Crotch, 1873

Agabinus｜Crotch, 1873

豆龙虱属｜Agabus｜Leach, 1817，端毛龙虱属

Agametrus｜Sharp, 1881

Agaporomorphus｜Zimmermann, 1921

Aglymbus｜Sharp, 1880

Agnoshydrus｜Biström, Nilsson & Wewalka, 1997

Allodessus｜Guignot, 1953

彩斑龙虱属｜Allopachria｜Zimmermann, 1924

Amarodytes｜Régimbart, 1900

Amurodytes｜Fery & Petrov, 2013

Andex｜Sharp, 1882

Andonectes｜Guéorguiev, 1971

Anginopachria｜Wewalka, Balke & Hendrich, 2001

Anodocheilus｜Babington, 1841

Antiporus｜Sharp, 1882

Australphilus｜Watts, 1978

Austrelatus｜Shaverdo, Hájek, Hendrich, Surbakti, Panjaitan & Balke, 2023

Austrodytes｜Watts, 1978

Barrethydrus

Barretthydrus｜Lea, 1927

- Barretthydrus geminatus Lea, 1927
- Barretthydrus stepheni Watts, 1978
- Barretthydrus tibialis Lea, 1927

Batrachomatus｜Clark, 1863

Belladessus｜K.B.Miller & Short, 2015

Bidessini｜Sharp, 1880

Bidessodes｜Régimbart, 1895

Bidessonotus｜Régimbart, 1895

Bidessus｜Sharp, 1882

Boreonectes｜(Fery & Ribera, 2018) Fery & Ribera, 2018

Boreonectes｜Angus, 2010

Borneodessus｜Balke, Hendrich, Mazzoldi & Biström, 2002

Brachyvatus｜Zimmermann, 1919

Brancuporus｜Hendrich, Toussaint & Balke, 2014

Bunites｜Spangler, 1972

Calicovatellus｜K.B.Miller & Lubkin, 2001

Canthyporus｜Zimmermann, 1919

Capelatus｜Turner & Bilton, 2015

Caperhantus｜Balke, Hájek & Hendrich, 2017

Carabdytes｜Balke, Hendrich & Wewalka, 1992

Carabhudrus

Carabhydrus｜Watts, 1978

Carrhydrus

Celina｜Aubé, 1837

Chostonectes｜Sharp, 1880

Clarkhydrus｜Fery & Ribera, 2018

Clemnius｜Villastrigo, Ribera, Manuel, Millán & Fery, 2017

Clypeodytes｜Régimbart, 1894

Coelambus｜Thomson, 1860

- Coelambus cribrarius (Scudder, 1900)
- Coelambus derelictus (Scudder, 1900)
- Coelambus disjectus (Scudder, 1900)
- Coelambus dzieduszyckii (Lomnicki, 1894)
- Coelambus ganglbaueri (Lomnicki, 1894)
- Coelambus infernalis (Scudder, 1900)
- Coelambus latefasciatus (Lomnicki, 1894)
- Coelambus miocenus (Wickham, 1912)
- Coelambus niedzwiedzkii (Lomnicki, 1894)
- Coelambus ozokeriticus (Lomnicki, 1894)
- Coelambus picipoides (Lomnicki, 1894)

Coelhydrus｜Sharp, 1882

Colymbetes｜Clairville, 1806

Colymbetes｜Schellenberg, 1806

Comaldessus｜Spangler & Barr, 1995

Copelatus｜Erichson, 1832

- Copelatus abonnenci Guignot, 1939
- Copelatus acamas Guignot, 1955
- Copelatus adelbert Megna, Atthakor, Manaono, Hendrich & Balke, 2017
- Copelatus advena Sharp, 1882
- Copelatus aemulus Bilardo & Rocchi, 1995
- Copelatus aequatorius Régimbart, 1899
- Copelatus aethiopicus Régimbart, 1906
- Copelatus agaroensis Balke, 2008
- Copelatus agrias Guignot, 1954
- Copelatus alternatus Sharp, 1882
- Copelatus amaroides Guignot, 1952
- Copelatus amatolensis Omer-Cooper, 1965
- Copelatus amazonicus Régimbart, 1889
- Copelatus ambiguus Bertrand & Legros, 1975
- Copelatus amphibius Ranarilalatiana & Bergsten, 2019
- Copelatus anastomosans Guignot, 1952
- Copelatus andobonicus Guignot, 1960
- Copelatus andreinii Régimbart, 1905
- Copelatus andrewesi J.Balfour-Browne, 1939
- Copelatus angolensis Peschet, 1924
- Copelatus angouei Bilardo & Rocchi, 2011
- Copelatus angustus Gschwendtner, 1932
- Copelatus ankaratra Ranarilalatiana & Bergsten, 2019
- Copelatus annobomensis J.Balfour-Browne, 1939
- Copelatus anthracinus Régimbart, 1895
- Copelatus antoniorum Hájek & Brancucci, 2011
- Copelatus aphroditae Balke, 2003
- Copelatus apicinotatus Bilardo & Rocchi, 2018
- Copelatus assalyi Bilardo & Rocchi, 2010
- Copelatus assimilis Régimbart, 1895
- Copelatus ateles Guignot, 1955
- Copelatus atrosulcatus Régimbart, 1906
- Copelatus babyrousa Hájek, Hendrich & Balke, 2018
- Copelatus bacchusi Wewalka, 1981
- Copelatus bacillifer Guignot, 1955
- Copelatus baculiformis Guignot, 1955
- Copelatus badeni Sharp, 1882
- Copelatus bakewelli J.Balfour-Browne, 1939
- Copelatus baoulicus Bilardo & Pederzani, 1978
- Copelatus bapounouensis Bilardo & Rocchi, 2019
- Copelatus baranensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus barbouri Young, 1942
- Copelatus basifasciatus Régimbart, 1895
- Copelatus basilewskyi Bilardo & Pederzani, 1979
- Copelatus bechynei Guignot, 1953
- Copelatus befasicus Guignot, 1956
- Copelatus bengalensis Guignot, 1955
- Copelatus bertrandi Nilsson, Bilardo & Rocchi, 1996
- Copelatus betampona Ranarilalatiana & Bergsten, 2019
- Copelatus bezdeki Sheth, Ghate & Hájek, 2018
- Copelatus bibulus Guignot, 1948
- Copelatus bicolor (Fabricius, 1792)
- Copelatus biformis Sharp, 1882
- Copelatus bimaculatus (Resende & Vanin, 1991)
- Copelatus binaghii Bilardo & Pederzani, 1978
- Copelatus biolleyi Guignot, 1952
- Copelatus biroi Guignot, 1956
- Copelatus birougouensis Bilardo & Rocchi, 2018
- Copelatus biseriatus J.Balfour-Browne, 1939
- Copelatus biswasi Mukherjee & Sengupta, 1986
- Copelatus blancasi Guignot, 1958
- Copelatus blatchleyi Young, 1953
- Copelatus bolivianus Guignot, 1957
- Copelatus bombycinus Guignot, 1952
- Copelatus bonvouloiri Sharp, 1882
- Copelatus bottegoi Régimbart, 1895
- Copelatus bougainvillensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus boukali Hendrich & Balke, 1998
- Copelatus brancuccii Rocchi, 1979
- Copelatus brasiliensis Zimmermann, 1921
- Copelatus brendelli Hájek, Hendrich & Balke, 2018
- Copelatus brevicornis (Sharp, 1882)
- Copelatus brevistrigatus Guignot, 1959
- Copelatus brivioi Rocchi, 1976
- Copelatus bromeliarum (Scott, 1912)
- Copelatus brullei Aubé, 1838
- Copelatus brunneus J.Balfour-Browne, 1939
- Copelatus buqueti Aubé, 1838
- Copelatus burgeoni Gschwendtner, 1930
- Copelatus caelatipennis Aubé, 1838
- Copelatus caelatipennis Chevrolat, 1863
- Copelatus caelatus Guignot, 1952
- Copelatus caffer J.Balfour-Browne, 1939
- Copelatus calaquei Bilardo & Rocchi, 2008
- Copelatus camerunensis Guignot, 1941
- Copelatus capensis Sharp, 1882
- Copelatus carayoni Legros, 1949
- Copelatus carinatus Sharp, 1882
- Copelatus celinoides Guignot, 1952
- Copelatus cessaima Caetano, Bená & Vanin, 2013
- Copelatus chevrolati Aubé, 1838
- Copelatus chibcha Guignot, 1952
- Copelatus chinensis Regimbart, 1899
- Copelatus chipiriricus Guignot, 1957
- Copelatus chloroticus Régimbart, 1899
- Copelatus cinnamomeus Régimbart, 1895
- Copelatus clarki Sharp, 1882
- Copelatus collarti Gschwendtner, 1932
- Copelatus compertus Guignot, 1956
- Copelatus concii Bilardo, 1982
- Copelatus concolor Sharp, 1882
- Copelatus concolorans J.Balfour-Browne, 1939
- Copelatus confinis Bilardo & Rocchi, 1999
- Copelatus congo Gschwendtner, 1938
- Copelatus consimilis Bilardo & Rocchi, 2002
- Copelatus consors Sharp, 1882
- Copelatus cooperae Bilardo & Pederzani, 1972
- Copelatus cordovai Megna & Epler, 2012
- Copelatus cordylinoides J.Balfour-Browne, 1938
- Copelatus coxalis Sharp, 1882
- Copelatus crassus Régimbart, 1895
- Copelatus cryptarchoides Régimbart, 1899
- Copelatus cubaensis Schaeffer, 1908
- Copelatus curtistriatus Bilardo & Rocchi, 1995
- Copelatus curvispinis Bilardo & Rocchi, 2008
- Copelatus cynthiae Bilardo & Rocchi, 2008
- Copelatus daemeli Sharp, 1882
- Copelatus danyi Megna & Epler, 2012
- Copelatus darlingtoni Young, 1942
- Copelatus davidi Wewalka, 2017
- Copelatus debilis Sharp, 1882
- Copelatus deccanensis Sheth, Ghate & Hájek, 2018
- Copelatus decellei Bilardo, 1982
- Copelatus decemsulcatus Régimbart, 1895
- Copelatus deceptor Bilardo & Rocchi, 1995
- Copelatus dentatipenis Jiang, Hajek & Jia, 2022
- Copelatus depressus Sharp, 1882
- Copelatus descarpentriesi Bertrand & Legros, 1975
- Copelatus diffisus Guignot, 1939
- Copelatus dimorphus Sharp, 1882
- Copelatus distinctus Aubé, 1838
- Copelatus distinguendus Régimbart, 1903
- Copelatus diversistriatus Jiang, Hajek & Jia, 2022
- Copelatus djiboutensis Wewalka & Jäch, 2017
- Copelatus dolosus Guignot, 1956
- Copelatus doriae Sharp, 1882
- Copelatus doudouensis Bilardo & Rocchi, 2015
- Copelatus duodecimstriatus Aubé, 1838
- Copelatus duponti Aubé, 1838
- Copelatus efoutensis Bilardo & Rocchi, 1995
- Copelatus ejactus Omer-Cooper, 1965
- Copelatus ellai Bilardo & Rocchi, 1995
- Copelatus elutus Guignot, 1958
- Copelatus enganensis Guignot, 1940
- Copelatus epactus Guignot, 1948
- Copelatus erichsonii Guérin-Méneville, 1847
- Copelatus espinhasso Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024
- Copelatus esteriensis Bilardo & Pederzani, 1978
- Copelatus eucritus Guignot, 1952
- Copelatus evanidus Bilardo & Rocchi, 1995
- Copelatus exaratus Sharp, 1882
- Copelatus externus Kirsch, 1873
- Copelatus falciformis Pederzani & Schizzerotto, 2017
- Copelatus fasciatus Bilardo & Rocchi, 1995
- Copelatus fastidiosus Guignot, 1959
- Copelatus feae Régimbart, 1888
- Copelatus felicis Hajek, Jiang & Jia, 2022
- Copelatus fernandensis Guignot, 1952
- Copelatus ferruginicollis Régimbart, 1895
- Copelatus ferus Guignot, 1953
- Copelatus festae Régimbart, 1899
- Copelatus fidschiensis Zimmermann, 1928
- Copelatus fijiensis Guignot, 1955
- Copelatus filiformis Sharp, 1882
- Copelatus fizpaci Bilardo & Rocchi, 2013
- Copelatus flavicans Guignot, 1952
- Copelatus flavidus Régimbart, 1895
- Copelatus florae Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024
- Copelatus fluviaticus Guignot, 1957
- Copelatus fontanus J.Balfour-Browne, 1950
- Copelatus fossilis Říha, 1974
- Copelatus fractistriatus Bilardo & Rocchi, 1995
- Copelatus freudei Guignot, 1955
- Copelatus fryi J.Balfour-Browne, 1939
- Copelatus fulviceps J.Balfour-Browne, 1938
- Copelatus fuscipennis Sharp, 1882
- Copelatus fuscomaculatus Guignot, 1952
- Copelatus gabonicus Bilardo & Pederzani, 1978
- Copelatus galapagoensis G.R.Waterhouse, 1845
- Copelatus garambanus Guignot, 1955
- Copelatus gardineri Scott, 1912
- Copelatus geayi Régimbart, 1904
- Copelatus geniculatus Sharp, 1882
- Copelatus gentilis Sharp, 1882
- Copelatus gestroi (Sharp, 1882)
- Copelatus gigas Bilardo & Rocchi, 2019
- Copelatus glyphicus (Say, 1823)
- Copelatus griffinii Régimbart, 1899
- Copelatus gschwendtneri Guignot, 1939
- Copelatus guadelupensis Legros, 1948
- Copelatus guerini Aubé, 1838
- Copelatus guineensis Guignot, 1955
- Copelatus haemorrhoidalis Régimbart, 1883
- Copelatus hararensis Guignot, 1952
- Copelatus hardenbergi J.Balfour-Browne, 1950
- Copelatus hebeter Guignot, 1953
- Copelatus heterogynus Régimbart, 1899
- Copelatus hydroporoides (Murray, 1859)
- Copelatus ibrahimi Zalat, Saleh, Angus & Kaschef, 2000
- Copelatus iguelaensis Bilardo & Rocchi, 2002
- Copelatus ilybioides Régimbart, 1895
- Copelatus imitator Bilardo & Rocchi, 2002
- Copelatus inaequalis Sharp, 1882
- Copelatus incognitus Sharp, 1882
- Copelatus indicus Sharp, 1882
- Copelatus inopinatus Bilardo & Rocchi, 1995
- Copelatus inornatus Sharp, 1882
- Copelatus insidiosus Bilardo & Rocchi, 1995
- Copelatus insolitus Chevrolat, 1863
- Copelatus instabilis Régimbart, 1897
- Copelatus instriatus Zimmermann, 1921
- Copelatus insuetus Guignot, 1941
- Copelatus insulanus Guignot, 1939
- Copelatus integer Sharp, 1882
- Copelatus internus Régimbart, 1904
- Copelatus interstriatus J.Balfour-Browne, 1939
- Copelatus inuber Guignot, 1952
- Copelatus ipiformis Régimbart, 1895
- Copelatus irinus Régimbart, 1899
- Copelatus irregularis W.J.MacLeay, 1871
- Copelatus ischius Guignot, 1956
- Copelatus jactator Guignot, 1955
- Copelatus jaechi Bilardo & Rocchi, 2016
- Copelatus jamaicensis Young, 1942
- Copelatus japonicus Sharp, 1884
- Copelatus jarrigei Legros, 1954
- Copelatus javanus Régimbart, 1883
- Copelatus jocosus Guignot, 1955
- Copelatus johannis Guignot, 1954
- Copelatus kalaharii Gschwendtner, 1935
- Copelatus kammuriensis Tamu & Tsukamoto, 1955
- Copelatus kaszabi Guignot, 1956
- Copelatus kely Ranarilalatiana & Bergsten, 2019
- Copelatus kietensis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus kilimandjarensis Bilardo & Pederzani, 1972
- Copelatus kindianus Guignot, 1954
- Copelatus kongouensis Bilardo & Rocchi, 1999
- Copelatus koreanus Mori, 1932
- Copelatus laccophilinus Sharp, 1882
- Copelatus laeticulus Sharp, 1882
- Copelatus laevipennis Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus lamottei Legros, 1954
- Copelatus lanzai Bilardo & Rocchi, 1995
- Copelatus laraensis Bilardo & Rocchi, 1995
- Copelatus lasckonyi Bilardo & Rocchi, 1995
- Copelatus latens Guignot, 1952
- Copelatus laticollis Régimbart, 1899
- Copelatus latifasciatus Bilardo & Rocchi, 1999
- Copelatus latipes Sharp, 1882
- Copelatus latus J.Balfour-Browne, 1950
- Copelatus leonardii Bilardo & Rocchi, 1999
- Copelatus leonensis Legros, 1954
- Copelatus lepersonneae Gschwendtner, 1943
- Copelatus lignosus Guignot, 1955
- Copelatus lineatipennis Guignot, 1955
- Copelatus lineatus (Guérin-Méneville, 1838)
- Copelatus longicornis Sharp, 1882
- Copelatus lootensi Guignot, 1955
- Copelatus louayensis Bilardo & Rocchi, 2004
- Copelatus luctuosus Guignot, 1939
- Copelatus luridescens Régimbart, 1889
- Copelatus luteocinctus Guignot, 1955
- Copelatus luteomaculatus Guignot, 1956
- Copelatus luzonicus Guignot, 1952
- Copelatus macellus Guignot, 1950
- Copelatus madoni Guignot, 1952
- Copelatus mahajanga Pederzani & Hájek, 2005
- Copelatus mahleri Holmen & Vazirani, 1990
- Copelatus makokouensis Bilardo & Rocchi, 1995
- Copelatus malaisei Guignot, 1954
- Copelatus malavergnorum Manuel & Ramahandrison, 2022
- Copelatus mancus Sharp, 1887
- Copelatus marginatus Sharp, 1882
- Copelatus marginipennis (Laporte, 1835)
- Copelatus martinbaehri Hendrich, Shaverdo, Hájek & Balke, 2019
- Copelatus martini Manuel, Deler-Hernández, Megna & Hájek, 2018
- Copelatus masculinus Régimbart, 1899
- Copelatus massaicus Guignot, 1941
- Copelatus mathani Guignot, 1952
- Copelatus maushomi Sheth, Ghate & Hájek, 2018
- Copelatus mbokoensis Bilardo & Rocchi, 2006
- Copelatus melanogrammus Régimbart, 1883
- Copelatus mendax Bilardo & Rocchi, 2011
- Copelatus miaowangi Hajek, Jiang & Jia, 2022
- Copelatus minimus Bilardo & Rocchi, 1995
- Copelatus minor Bilardo & Pederzani, 1978
- Copelatus minutissimus J.Balfour-Browne, 1939
- Copelatus mocquerysi Régimbart, 1895
- Copelatus mohelicus Guignot, 1959
- Copelatus monticola Guignot, 1951
- Copelatus montivagus Young, 1942
- Copelatus mopanshanensis Jiang, Zhao & Hajek, 2022
- Copelatus mulangensis Bameul, 2003
- Copelatus mundus Sharp, 1882
- Copelatus mutabilis Bilardo & Rocchi, 1999
- Copelatus mvoungensis Bilardo & Rocchi, 2004
- Copelatus mysorensis Vazirani, 1970
- Copelatus nakamurai Guéorguiev, 1970
- Copelatus nangaensis Guignot, 1952
- Copelatus nanlingensis Hajek, Jiang & Jia, 2022
- Copelatus nauclerus Guignot, 1939
- Copelatus neavei J.Balfour-Browne, 1950
- Copelatus neelumae Vazirani, 1973
- Copelatus neogestroi Balke, 2008
- Copelatus neoguineensis Zimmermann, 1919
- Copelatus nguemai Bilardo & Rocchi, 2011
- Copelatus nigricans Sharp, 1882
- Copelatus nigricollis Paulino de Oliveira, 1880
- Copelatus nigrolineatus Sharp, 1882
- Copelatus nigropennis Zimmermann, 1927
- Copelatus nigrostriatus Régimbart, 1895
- Copelatus nilssoni Wewalka, 1982
- Copelatus nimbaensis Legros, 1958
- Copelatus nitens Bilardo & Rocchi, 1999
- Copelatus nitidus Sharp, 1882
- Copelatus nodieri Régimbart, 1895
- Copelatus normalis Erichson, 1847
- Copelatus notius Omer-Cooper, 1965
- Copelatus nzei Bilardo & Rocchi, 1999
- Copelatus oblitus Sharp, 1882
- Copelatus obscurus Sharp, 1882
- Copelatus occultus Bilardo & Rocchi, 1995
- Copelatus ogasawarensis Kamiya, 1932
- Copelatus onorei Pederzani & Rocchi, 1982
- Copelatus ornatipennis Zimmermann, 1926
- Copelatus owas Régimbart, 1895
- Copelatus pachys Guignot, 1955
- Copelatus pallidus Régimbart, 1895
- Copelatus paludorum Guignot, 1952
- Copelatus pandanorum Scott, 1912
- Copelatus panguana Hájek, Alarie, Benetti, Hamada, Springer, Hendrich, Villastrigo, Torres, Basantes & Balke, 2024
- Copelatus pantosi Guignot, 1958
- Copelatus papuensis J.Balfour-Browne, 1939
- Copelatus parabaptus Guignot, 1955
- Copelatus parallelipipedus Régimbart, 1895
- Copelatus parallelus Zimmermann, 1920
- Copelatus pardii Rocchi, 1990
- Copelatus parisii Guignot, 1959
- Copelatus parumstriatus Gschwendtner, 1934
- Copelatus paryphes Guignot, 1955
- Copelatus pederzanii Bilardo & Rocchi, 1995
- Copelatus pereirai Guignot, 1955
- Copelatus peridinus Guignot, 1955
- Copelatus pinnifer Guignot, 1952
- Copelatus piriensis Omer-Cooper, 1965
- Copelatus platynotus Régimbart, 1895
- Copelatus polystrigus Sharp, 1882
- Copelatus ponomarenkoi Říha, 1974
- Copelatus portior Guignot, 1956
- Copelatus posticatus (Fabricius, 1801)
- Copelatus poungai Bilardo & Rocchi, 2006
- Copelatus predaveterus Miller, 2003
- Copelatus prolixus Sharp, 1882
- Copelatus prolongatus Sharp, 1882
- Copelatus propino Guignot, 1960
- Copelatus propinquus Régimbart, 1895
- Copelatus proximus Sharp, 1882
- Copelatus pseudostriatus Ranarilalatiana & Bergsten, 2019
- Copelatus pulchellus (Klug, 1834)
- Copelatus pulicarius Régimbart, 1895
- Copelatus pumilus Régimbart, 1895
- Copelatus punctatus Bilardo & Rocchi, 1995
- Copelatus punctulatus Aubé, 1838
- Copelatus puzhelongi Jiang, Hajek & Jia, 2022
- Copelatus quadrisignatus Régimbart, 1877
- Copelatus quadristriatus Pederzani & Schizzerotto, 2017
- Copelatus racenisi Guignot, 1951
- Copelatus ragazzii Régimbart, 1887
- Copelatus regimbarti Branden, 1885
- Copelatus restrictus Sharp, 1882
- Copelatus rimosus Guignot, 1952
- Copelatus rivalis Guignot, 1952
- Copelatus rocchii Bilardo, 1982
- Copelatus romani Zimmermann, 1924
- Copelatus rosulae Hajek, Jiang & Jia, 2022
- Copelatus royi Legros, 1958
- Copelatus rubiginosus Guignot, 1954
- Copelatus ruficapillus Régimbart, 1895
- Copelatus ruteri Legros, 1954
- Copelatus saegeri Guignot, 1955
- Copelatus safiotra Ranarilalatiana & Bergsten, 2019
- Copelatus sahlbergi J.Balfour-Browne, 1939
- Copelatus sallaei Sharp, 1882
- Copelatus sanfilippoi Bilardo & Pederzani, 1972
- Copelatus saverii Pederzani & Schizzerotto, 2017
- Copelatus scalptus Guignot, 1954
- Copelatus scaphites Guignot, 1955
- Copelatus schaefferi Young, 1942
- Copelatus schereri Wewalka, 1981
- Copelatus schuhi Hendrich & Balke, 1998
- Copelatus scutatus Guignot, 1952
- Copelatus scytalotus Guignot, 1956
- Copelatus senegalensis Lagar & Hernando, 1991
- Copelatus sexstriatus Sharp, 1882
- Copelatus sharpi Branden, 1885
- Copelatus sibelaemontis Hájek, Hendrich, Hawlitschek & Balke, 2010
- Copelatus silvestrii Régimbart, 1903
- Copelatus simoni Régimbart, 1889
- Copelatus singularis Bilardo & Rocchi, 1995
- Copelatus sociennus J.Balfour-Browne, 1952
- Copelatus solitarius Sharp, 1882
- Copelatus sordidipennis Régimbart, 1895
- Copelatus speciosus Régimbart, 1892
- Copelatus spoliatus Guignot, 1955
- Copelatus stavropolitanus Říha, 1974
- Copelatus stillicidicola Bilardo & Rocchi, 1995
- Copelatus striatellus Boheman, 1848
- Copelatus striaticollis Lucas, 1857
- Copelatus striatopterus Aubé, 1838
- Copelatus striatulus Aubé, 1838
- Copelatus strigipennis (Laporte, 1835)
- Copelatus strigosulus (Fairmaire, 1878)
- Copelatus strinatii Guignot, 1958
- Copelatus striolatus Peschet, 1917
- Copelatus stygis Guignot, 1958
- Copelatus subdeficiens Régimbart, 1902
- Copelatus subsimilis Guignot, 1958
- Copelatus substriatus Kirsch, 1873
- Copelatus subterraneus Guéorguiev, 1978
- Copelatus sudrei Bameul, 2003
- Copelatus sulcatus Sharp, 1882
- Copelatus sulcipennis (Laporte, 1835)
- Copelatus sumbawensis Régimbart, 1899
- Copelatus suppar Guignot, 1956
- Copelatus supplementaris Régimbart, 1895
- Copelatus sylvaticoides Bilardo & Rocchi, 2013
- Copelatus sylvaticus Guignot, 1952
- Copelatus takakurai Satô, 1985
- Copelatus tanaus Guignot, 1953
- Copelatus taprobanicus Wewalka & Vazirani, 1985
- Copelatus tenebrosis Regimbart, 1880
- Copelatus tenebrosus Régimbart, 1880
- Copelatus tengchongensis Hajek, Jiang & Jia, 2022
- Copelatus teranishii Kamiya, 1938
- Copelatus terminalis Sharp, 1882
- Copelatus ternatensis Régimbart, 1899
- Copelatus terzanii Bilardo & Saverio, 2016
- Copelatus thiriouxi Peschet, 1917
- Copelatus thrasys Guignot, 1952
- Copelatus tibialis Sharp, 1882
- Copelatus tinctor Guignot, 1954
- Copelatus togoensis Régimbart, 1895
- Copelatus tomokunii Satô, 1985
- Copelatus tondangoyei Bilardo & Rocchi, 2013
- Copelatus tostus J.Balfour-Browne, 1950
- Copelatus trifilis Guignot, 1958
- Copelatus triglyphus Guignot, 1955
- Copelatus trilobatus Régimbart, 1895
- Copelatus tschaga Bilardo & Pederzani, 1972
- Copelatus tucuchiensis J.Balfour-Browne, 1939
- Copelatus tulagicus Guignot, 1942
- Copelatus uludanuensis Hendrich & Balke, 1995
- Copelatus undecimstriatus Aubé, 1838
- Copelatus urceolus Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus usagarensis Zimmermann, 1926
- Copelatus vagatus Guignot, 1952
- Copelatus vagestriatus Zimmermann, 1919
- Copelatus vagus Bilardo & Rocchi, 1995
- Copelatus validus Sharp, 1882
- Copelatus vancli Hájek, Hendrich & Balke, 2018
- Copelatus vanninii Bilardo & Rocchi, 1999
- Copelatus variegatus Régimbart, 1895
- Copelatus variistriatus Hájek, Shaverdo, Hendrich & Balke, 2021
- Copelatus venustus Bilardo & Rocchi, 1995
- Copelatus vigintistriatus Fairmaire, 1869
- Copelatus vigintisulcatus Régimbart, 1895
- Copelatus villiersi Guignot, 1950
- Copelatus virungaensis Bilardo, 1982
- Copelatus vitraci Legros, 1948
- Copelatus vivax Guignot, 1953
- Copelatus vokoka Ranarilalatiana & Bergsten, 2019
- Copelatus wallacei J.Balfour-Browne, 1939
- Copelatus waltoni J.Balfour-Browne, 1950
- Copelatus wayanadensis Manivannan & Madani, 2011
- Copelatus wewalkai Holmen & Vazirani, 1990
- Copelatus weymarni J.Balfour-Browne, 1947
- Copelatus xanthocephalus Régimbart, 1899
- Copelatus xiniudong Jiang, Hajek & Jia, 2022
- Copelatus yacumensis Guignot, 1957
- Copelatus yaguarete Scheers, 2018
- Copelatus yombensis Bilardo & Rocchi, 2015
- Copelatus zadiensis Bilardo & Rocchi, 1995
- Copelatus zanabato Manuel & Ramahandrison, 2022
- Copelatus zanatanensis Ranarilalatiana & Bergsten, 2019
- Copelatus zimmermanni Gschwendtner, 1934

Copelatus｜Sheth, Ghate & Hájek, 2018

Coptotomus｜Say, 1830

Cretodytes｜Ponomarenko, 1977

Crinodessus｜K.B.Miller, 1997

大龙虱属｜Cybister｜Curtis, 1827，真龙虱属

Darwinhydrus｜Sharp, 1882

Deronectes｜Sharp, 1882

Derovatellus｜Sharp, 1882

Desmopachria｜Babington, 1841

Deuteronectes｜Guignot, 1945

Deuteronectus

Dimitshydrus｜Uéno, 1996

Dyschirioides

Dyticus｜Geoffroy, 1762

Dytiscus｜Linnaeus, 1758

Electruphilus｜Balker, Toledo & Hendrich, 2019

Ereboporus｜K.B.Miller, Gibson & Alarie, 2009

Eretes｜Laporte, 1833

Etruscodytes｜Mazza, Cianferoni & Rocchi, 2013

Exocelina｜Broun, 1886

Fontidessus｜K.B.Miller & Spangler, 2008

Geodessus｜Brancucci, 1979

Gibbidessus｜Watts, 1978

Glareadessus｜Wewalka & Biström, 1998

Graphoderus｜Dejean, 1833

Graptodytes｜Seidlitz, 1887

Haideoporus｜Young & Longley, 1976

Hamadiana｜Benetti, Short & Michat, 2019

Hemibidessus｜Zimmermann, 1921

Heroceras｜Guignot, 1950

Heterhydrus｜Fairmaire, 1869

Heterosternuta｜Strand, 1935

Hintonia｜Young, 1980

Hoperius｜Fall, 1927

Hornectes｜Fery & Ribera, 2018

Hovahydrus｜Biström, 1982

Huxelhydrus｜Sharp, 1882

Hydaticus｜Leach, 1817

Hyderodes｜Hope, 1838

Hydrocolus｜Roughley & Larson, 2000

Hydrodessus｜J.Balfour-Browne, 1953

Hydrodytes｜K.B.Miller, 2001

短褶龙虱属｜Hydroglyphus｜Motschulsky, 1853

Hydronebrius｜Jakovlev, 1897

Hydropeplus｜Sharp, 1882

Hydroporus｜Clairville, 1806

Hydroporus｜Schellenberg, 1806

Hydrotrupes｜Sharp, 1882

Hydrovatus｜Motschulsky, 1853

水龙虱属｜Hygrotus｜Stephens, 1828

Hyphoporus｜Sharp, 1882

Hyphovatus｜Wewalka & Biström, 1994

异爪龙虱属｜Hyphydrus｜Illiger, 1802

Hypodessus｜Guignot, 1939

Iberonectes｜Fery & Ribera, 2018

Iberoporus｜Castro & Delgado, 2001

Ilybiosoma｜Crotch, 1873

Ilybius｜Erichson, 1832

Incomptodessus｜K.B.Miller & García, 2011

Isonotus｜Houlbert, 1934

Japanolaccophilus｜Satô, 1972

Kakadudessus｜Hendrich & Balke, 2009

- Kakadudessus tomweiri Hendrich & Balke, 2009

Kuschelydrus｜Ordish, 1976

Laccodytes｜Régimbart, 1895

Laccomimus｜Toledo & Michat, 2015

Lacconectus｜Motschulsky, 1855

粒龙虱属｜Laccophilus｜Leach, 1815

Laccoporus｜J.Balfour-Browne, 1939

- Laccoporus nigritulus (Gschwendtner, 1936)

Laccornellus｜Roughley & Wolfe, 1987

Laccornis｜Gozis, 1914

Laccosternus｜Brancucci, 1983

Lancetes｜Sharp, 1882

Laodytes｜Queney, Lemaire & Ferrand, 2020

Larsonectes｜Fery & Ribera, 2018

Leconectes｜Fery & Ribera, 2018

Leiodytes｜Guignot, 1936

Leuronectes｜Sharp, 1882

Liadyxianus｜Prokin, Petrov, Wang & Ponomarenko, 2013

Limbodessus｜Guignot, 1939

Liodessus｜Guignot, 1939

Lioporeus｜Guignot, 1950

Liopterus｜Dejean, 1833

Madaglymbus｜Shaverdo & Balke, 2008

Matus｜Aubé, 1836

Megadytes｜Sharp, 1882

Megaporus｜Brinck, 1943

Meladema｜Laporte, 1835

Melanodytes｜Seidlitz, 1887

Meridiorhantus｜Balke, Hájek & Hendrich, 2017

Mesodytes｜Prokin, Petrov, Wang & Ponomarenko, 2013

Metaporus｜Guignot, 1945

Methles｜Sharp, 1882

Microdessus｜Young, 1967

Microdytes｜J.Balfour-Browne, 1946

Microhydrodytes｜K.B.Miller, 2002

Miodytiscus｜Wickham, 1911

Morimotoa｜Uéno, 1957

Mystonectes｜Fery & Ribera, 2018

Napodytes｜Steiner, 1981

Nartus｜Zaitzev, 1907

孔龙虱属｜Nebrioporus｜Régimbart, 1906

Necterosoma｜Sharp, 1882

Necterosoma｜W.J.MacLeay, 1871

Nectoboreus｜Fery & Ribera, 2018

Nectomimus｜Fery & Ribera, 2018

Nectoporus｜Guignot, 1950

Neobidessodes｜Hendrich & Balke, 2009

Neobidessus｜Young, 1967

Neoclypeodytes｜Young, 1967

Neonectes｜J.Balfour-Browne, 1944

Neonectes｜Zimmermann, 1932

Neoporus｜Guignot, 1931

Neoscutopterus｜J.Balfour-Browne, 1943

彩裳粒龙虱属｜Neptosternus｜Sharp, 1882

Notaticus｜Zimmermann, 1928

Novadessus｜K.B.Miller, 2016

Onychohydrus｜Schaum & White, 1847

Oreodytes｜Seidlitz, 1887

Pachydrus｜Sharp, 1882

Pachynectes｜Régimbart, 1903

Palaeodytes｜Ponomarenko, 1987

Papuadessus｜Balke, 2001

Paroster｜Sharp, 1882

Peschetius｜Guignot, 1942

Petrodessus｜K.B.Miller, 2012

Philaccolilus｜Guignot, 1937

Philaccolus｜Guignot, 1937

Philodytes｜J.Balfour-Browne, 1939

Phreatodessus｜Ordish, 1976

Platambus｜Thomson, 1859

Platydytes｜Biström, 1988

Platynectes｜Gueorguiev, 1972

Platynectes｜Régimbart, 1879

Porhydrus｜Guignot, 1945

Primospes｜Sharp, 1882

Procoelambus｜Théobald, 1937

Prodaticus｜Sharp, 1882

缝龙虱属｜Pseuduvarus｜Biström, 1988

Psychopomporus｜Jean, Telles & K.B.Miller, 2012

Pteroporus｜Guignot, 1933

Queda｜Sharp, 1882

Regimbartina｜Chatanay, 1911

Rhantaticus｜Sharp, 1880

Rhantus｜Dejean, 1833

Rhithrodytes｜Bameul, 1989

Rompindessus｜Balke, Bergsten & Hendrich, 2017

Sandracottus｜Sharp, 1882

Sanfilippodytes｜Franciscolo, 1979

Scarodytes｜Gozis, 1914

Schistomerus｜Palmer, 1957

Sekaliporus｜Watts, 1997

Sharphydrus｜Omer-Cooper, 1958

Siamoporus｜Spangler, 1996

Siettitia｜Abeille de Perrin, 1904

Sinodytes｜Spangler, 1996

Spanglerodessus｜K.B.Miller & García, 2011

Spencerhydrus｜Sharp, 1882

Sternhydrus｜Brinck, 1945

Sternopriscus｜Sharp, 1880

Stictonectes｜Brinck, 1943

Stictotarsus｜Zimmermann, 1919

Stygoporus｜Larson & LaBonte, 1994

Suphisellus｜Zimmermann, 1921

Tassilodytes｜Fery & Bouzid, 2016

Tepuidessus｜Spangler, 1981

Thermonectus｜Dejean, 1833

- Thermonectus alfredi Griffini, 1898
- Thermonectus basillaris (Harris, 1829)
- Thermonectus batesi Sharp, 1882
- Thermonectus circumscriptus (Latreille, 1809)
- Thermonectus cuneatus Sharp, 1882
- Thermonectus depictus Sharp, 1882
- Thermonectus duponti (Aubé, 1838)
- Thermonectus intermedius Crotch, 1873
- Thermonectus laporti (Aubé, 1838)
- Thermonectus leprieuri J.Balfour-Browne, 1944
- Thermonectus margineguttatus (Aubé, 1838)
- Thermonectus marmoratus (Gray, 1831)
- Thermonectus nigrofasciatus (Aubé, 1838)
- Thermonectus nobilis Zimmermann, 1924
- Thermonectus sibleyi Goodhue-McWilliams, 1981
- Thermonectus simulator Sharp, 1882
- Thermonectus succinctus (Aubé, 1838)
- Thermonectus tremouillesi Michat & Torres, 2016
- Thermonectus variegatus (Dejean, 1836)
- Thermonectus variegatus (Laporte de Castelnau, 1835)
- Thermonectus zimmermani Goodhue-McWilliams, 1981

Tikoloshanes｜Omer-Cooper, 1956

Tiporus｜Watts, 1985

Trichonectes｜Guignot, 1941

Trogloguignotus｜Sanfilippo, 1958

Tyndallhydrus｜Sharp, 1882

Typhlodessus｜Brancucci, 1985

Uvarus｜Guignot, 1939

Vatellus｜Aubé, 1837

Yola｜Gozis, 1886

Yolina｜Guignot, 1936

Zaitzevhydrus｜Fery & Ribera, 2018

Zimpherus｜K.B.Miller & Wheeler, 2015

## 参考文献

- 联邦科学与工业研究组织 (CSIRO) (2004): Water for a Healthy Country - Family Dytiscidae . Version of 2004-JUL-02. Retrieved 2008-AUG-04
- De Foliart, Gene R. (2002): Chapter 26 - Eastern Asia: China, Japan, and other countries. In: The Human Use of Insects as a Food Resource: A Bibliographic Account in Progress.
- Jäch, Manfred A. (2003): Fried water beetles Cantonese style. American Entomologist 49(1): 34-37. PDF fulltext
- Larson, D.J., Alarie, Y., and Roughley, R.E. (2000): Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa. .

## 基本资料

- 说明："Cybister lateralimarginalis"
- fossil_range：侏罗纪晚期至今
- 学名：Dytiscidae
- 命名者：Leach, 1815
- 模式属：龙虱属 Dytiscus
- type_genus_authority：Linnaeus，1758
- subdivision_ranks：亚科、族与属
- 栽培群成员：见内文
- diversity：至少160个属，约4000物种
- diversity_link：龙虱科物种表