蒂奧得螺屬（學名：）是腹足綱蜑螺科軟體動物的一個屬，體型細小，有螺厴，棲息於淡水或半鹹水環境中 Bouchet P 2014 Theodoxus Montfort 1810 Accessed through World Register of Marine Species at httpwwwmarinespeciesorgaphiaphpptaxdetailsid153531 on 20141117。 分佈 本屬物種分佈於歐洲及北非，並往東延伸至。除本屬以外，蜑螺科並沒有其他屬與本屬具。這在蜑螺科相當例外，因為其他蜑螺科物種只分佈於南半球的熱帶。本屬物種均為蜑螺科僅有棲息於溫帶的物種。 Bunje 和 Lindberg 2007 提出了 Theodoxus 分支的第一個系統發育假說。 Bunje P M Lindberg D R 2007 Lineage divergence of a freshwater snail clade associated with postTethys marine basin development Molecular Phylogenetics and Evolution 42 2 373387 doi101016jympev200606026 Theodoxus 屬的進化受到中新世地中海與副特提斯海分離的影響。 形態描述 本屬物種的螺殼呈半卵形，具有平坦的殼口。沒有。及內溶解。 蒂奧得螺屬物種間的差異，無論是大小、外殼膜上的顏色與花紋變化、螺厴及齒舌的細節都有很大差異，這使得本屬物種的辨認比較困難 Zettler M L Frankowski J Bochert R Röhner M 2004 Morphological and ecological features of Theodoxus fluviatilis Linnaeus 1758 from Baltic brackish water and German freshwater populations 38 3 305316 pdf PDF。 生態 These animals live on stones and often also under stones in up to 56 m depth or deeper feeding on algal covers Theodoxus needs rough surfaces in order to be able to digest its food so a stony 基質 生物生活環境 is necessary Green algae are not consumed Theodoxus has no 纖維素酶 s These snails lay egg capsules containing 3070 eggs each usually on the shells of other Theodoxus animals only one juvenile grows the other eggs serve as food 物種 截至 2007 年，還未有對蒂奧得螺屬作系統評價的研究。Bunje 2004 的研究指出本屬至少有 34 個現害種；而在 2015 年度的國際自然保護聯盟瀕危物種紅色名錄列出了本屬包括有至少 23 個物種 The IUCN Red List of Threatened Species Version 20143 wwwiucnredlistorg Downloaded on 23 March 2015 Searching for Theodoxus。 本屬包括下列各物種： Subgenus Brusinaella Andrusov 1912Andrussov 1912 Verh Russ min Ges St Pétersburg 2 49 4 Theodoxus petasatus Seninski 1905 Subgenus Calvertia Bourguignat 1880Bourguignat 1880 Etude Foss Tert Quat Vallée Cettina page 44 Theodoxus acuticarinatus Fuchs 1870 Theodoxus amblygonioides Wenz 1930 Theodoxus amethystinus Brusina 1874 Theodoxus anconae Capellini 1880 Theodoxus angularis Gozhik 2002 Theodoxus anonymus Anistratenko Gozhik 1995 Theodoxus atticus Kühn 1963 Theodoxus barakovici Brusina 1902 Theodoxus bessarabicus Sinzov 1896 Theodoxus brusinai Pavlović 1931 Theodoxus capellinii Pantanelli 1876 Theodoxus capillaceus Brusina 1874 Theodoxus crescens Fuchs 1870 Theodoxus doderleini dAncona 1869 Theodoxus dumortieri Fontannes 1878 Theodoxus fasciatus Gozhik in Gozhik Datsenko 2007 Theodoxus gnezdai Brusina 1884 Theodoxus grateloupianus Férussac 1823 Theodoxus hoernesanus Semper 1867 Theodoxus ingulenzis Gozhik in Gozhik Datsenko 2007 Theodoxus kalodictya Andrusov 1909 Theodoxus koslinskyi Porumbaru 1881 Theodoxus lamellatus Brusina 1892 Theodoxus lineatus Sinzov 1896 Theodoxus lorkovici Brusina 1878 Theodoxus macedonicus Wenz 1943 Theodoxus maculosus Gozhik in Gozhik Datsenko 2007 Theodoxus mayeri Semper 1867 Theodoxus militaris Neumayr 1869 Theodoxus miljkovici Brusina 1902 Theodoxus millepunctatus Brusina 1902 Theodoxus moeschi Locard 1893 Theodoxus morulus Kühn 1963 Theodoxus mutinensis dAncona 1869 Theodoxus neumayri Burgerstein 1877 Theodoxus nitens Gozhik 2002 Theodoxus nivosus Brusina 1874 Theodoxus novorossicus Sinzov 1896 Theodoxus obtusangulaeformis Gozhik 2002 Theodoxus oxytropida Andrusov 1909 Theodoxus pavlovici Milošević 1983 Theodoxus philippianus Locard 1883 Theodoxus pilari Brusina 1884 Theodoxus platystoma Brusina 1874 Theodoxus pseudodanubialis Sinzov 1896 Theodoxus pseudofluviatilis Locard 1893 Theodoxus pseudograteloupanus Sinzov 1884 Theodoxus radmanesti Fuchs 1870 Theodoxus radovanovici Brusina 1893 Theodoxus reiseri Brusina 1902 Theodoxus rugosus Pavlović 1931 Theodoxus sagittiferus Brusina 1874 Theodoxus schachmaticus Andrusov 1909 Theodoxus scoliogramma Brusina 1884 Theodoxus semidentatus Sandberger 1875 Theodoxus serrulatus Brusina 1892 Theodoxus slavonicus Brusina 1878 Theodoxus sophievkaensis Gozhik in Gozhik Datsenko 2007 Theodoxus stanae Brusina 1893 Theodoxus stefanescui Fontannes 1887 Theodoxus stefanescuiformis Gozhik in Gozhik Datsenko 2007 Theodoxus subquadrofasciatus Gozhik 2002 Theodoxus subslavonicus Gozhik 2002 Theodoxus sundicus Andrusov 1909 Theodoxus suskalovici Pavlović 1903 Theodoxus sycophantus Brusina 1878 Theodoxus tropidophorus Brusina 1884 Theodoxus turbinatus Fuchs 1870 Theodoxus veljetinensis Pavlović 1903 Theodoxus venustus Brusina 1897 Theodoxus xanthozona Brusina 1884 Theodoxus zagradovkaensis Gozhik 2002 Theodoxus zivkovici Pavlović 1903 Subgenus Neritonyx Andrusov 1912Andrussov 1912 Verh Russ min Ges St Pétersburg 2 49 page 4 Brusinaella and page 1 Neritonyx Theodoxus unguiculatus Seninski 1905 Subgenus Ninniopsis Tomlin 1930 1930 Some preoccupied generic namesII 19 1 2224 page 24 Theodoxus colchicus Andrusov 1912 Subgenus Neritaea Roth 1855Roth 1855 Malak Bl 2 56 Récluz 1841Neritaea Fauna Europaea last update 27 January 2011 accessed 12 April 2011 Theodoxus groyanus Férussac 1823 Sowerby Theodoxus micans Gaudry Fischer in Gaudry 1867 Reeve 1856 synonym Theodoxus africanus Reeve 1856 Theodoxus subdoricus Schütt 1976 Theodoxus subterrelictus Schütt 1963 Theodoxus peloponensis Récluz 1841 Theodoxus varius Menke 1828 Subgenus Theodoxus Montfort 1810 Schütt 1965Species in genus Theodoxus n20 accessed 11 April 2011 Schütt H 1965 Zur Systematik und Ökologie türkischer Süßwasserprosobranchier Zoologische Mededelingen 41 3 4372 Plate 1 abstract PDF Theodoxus baeticus Lamarck 1822Theodoxus Fauna Europaea last update 27 January 2011 accessed 12 April 2011 Theodoxus becenensis Cobălcescu 1883 Theodoxus brenneri Handmann 1882 Theodoxus carasiensis Jekelius 1944 Theodoxus constantiae Stefanescu 1896 Theodoxus culceri Porumbaru 1881 Theodoxus cyrtocelis Krauss 1852 Pfeiffer 1828 Theodoxus denisluensis Oppenheim 1919 Theodoxus deperditus Almera 1894 Theodoxus elongatulus Philippi 1844 Theodoxus eugenii Jekelius 1944 Clessin 1886 Linnaeus 1758 模式種Anistratenko V V 2005 Lectotypes for Tricolia pullus Gibbula divaricata and Theodoxus fluviatilis Mollusca Gastropoda revisited Vestnik zoologii 39 6 310 PDF synonyms Theodoxus brauneri Lindholm 1908 but as separate species per Theodoxus brauneri f lacrymans Lindholm 1908 Theodoxus brauneri f alboguttata Lindholm 1908 Theodoxus brauneri f pulherrima Lindholm 1908 Theodoxus lutetianus Montfort 1810 Bunje 2005 PDF does not consider Theodoxus velox Anistratenko 1999Anistratenko O Y Anistratenko V V 1999 Mollusks of the genus Theodoxus Gastropoda Pectinibranchia Neritidae from the Black and the Azov seas basin Vestnik Zoologii 33 1119 to be a distinct species from Theodoxus fluviatilis Theodoxus gregarius Thomä 1845 Theodoxus heldreichi Martens 1879 Theodoxus hisingeri Bellardi Michelotti 1841 Theodoxus hispalensis Martens 1879 Theodoxus jekeliusi JurišićPolšak 1979 Theodoxus leobersdorfensis Handmann 1887 Philippi 1836 Theodoxus milessii Papp 1979 Theodoxus moosbrunnensis Papp 1953 Theodoxus morellii Bellardi Michelotti 1841 Lindholm 1924 synonym Theodoxus lituratus Eichwald 1838 Theodoxus pilidei Tournouër 1879 Pfeiffer 1828 Theodoxus punctatolineatus Sinzov 1896 Bourguignat 1852 Theodoxus scamandrius Calvert Neumayr 1880 Theodoxus semiplicatus Neumayr in Herbich Neumayr 1875 Theodoxus simplex Fuchs 1877 Theodoxus subglobosus Eichwald 1853 Bourguignat in Issel 1865 or Theodoxus fluviatilis subthermalis Issel 1865 Pfeiffer 1828 Theodoxus trifasciatus Grateloup 1839 Theodoxus turislavicus Jekelius 1944 Graells 1846 Theodoxus zografi Brusina 1902 Subgenus Theodoxus abnormis Jenkins 1864 Theodoxus almelae Revilla 1958 Theodoxus astrachanicus Starobogatov 1994Kantor Yu I Vinarski M V Schileyko A A Sysoev A V published online on March 2 2010 Catalogue of the continental mollusks of Russia and adjacent territories Version 231 Theodoxus banaticus Jekelius 1944 Theodoxus bohotinensis Simionescu Barbu 1940 Theodoxus bolivari Royo Gómez 1922 Theodoxus bukowskii Oppenheim 1919 Theodoxus cariosus Wood 1828 Leach 1815 WoRMS 2010 Theodoxus coronatus Leach 1815 Accessed through World Register of Marine Species at httpwwwmarinespeciesorgaphiaphpptaxdetailsid216264 on 20110411 Theodoxus crenulatus Klein 1853 Theodoxus cunici Brusina 1892 Theodoxus danasteri Lindholm 1908 Theodoxus doricus Neumayr 1880 Clessin 1886 Theodoxus gloeri Odabaşi Arslan 2015 Theodoxus gurur Sands Glöer 2020 Theodoxus hellenicus Bukowski 1896 Theodoxus imbricatus Brusina 1878 Theodoxus intracarpaticus Jekelius 1944 Theodoxus karasuna Theodoxus licherdopoli Stefanescu 1896 Theodoxus luteofasciatus Miller 1879 Theodoxus macri Sowerby 1849 Bourguignat 1864 Theodoxus marteli Pallary 1920 Theodoxus milachevitchi Golikov Starobogatov 1966 Theodoxus mutinensis Récluz 1841 Theodoxus olivaceus Lindholm 1924 Theodoxus pallidus Dunker 1861 Theodoxus pappi Sauerzopf 1952 Theodoxus paradisii Magrograssi 1928 Theodoxus patrae Esu Girotti 2015 Theodoxus percarinatus Oppenheim 1919 Theodoxus pilidei Tournouêr 1879 Theodoxus politioanei Jekelius 1944 Theodoxus politus Jekelius 1944 Theodoxus poppei Ahuir 2020 Theodoxus postcrenulatus Papp 1953 Theodoxus prozlatarici Jekelius 1944 Theodoxus pseudodacicus Neubauer Harzhauser Georgopoulou Mandic Kroh 2014 Theodoxus quadrifasciatus Bielz 1864 Theodoxus reticularis Sowerby 1838Madhyastha A Dutta J Daniel B A 2010 Theodoxus reticularis In IUCN 2010 IUCN Red List of Threatened Species Version 20104 wwwiucnredlistorg Downloaded on 11 April 2011 IT IS A SYNONYM OF Clithon reticularis httpoldredlistiucnredlistorgdetails1667390 Theodoxus reticulatus Pană 1990 Theodoxus retropictus UNREFERENCED see probably Clithon retropictum Theodoxus rhodiensis Tournouër in Fischer 1877 Theodoxus sarmaticus Lindholm 1901 Theodoxus schultzii Grimm 1877 Zettler M L 2007 A redescription of Theodoxus schultzii Grimm 1877 an endemic neritid gastropod of the Caspian Sea 39 3 245252 abstract Theodoxus scriptus Stefanescu 1896 Theodoxus serratiliniformis Geyer 1914Glöer P 2002 Die Süßwassergastropoden Nord und Mitteleuropas Die Tierwelt Deutschlands ConchBooks Hackenheim 326 pp Theodoxus sinjanus Brusina 1876 Theodoxus soceni Jekelius 1944 Theodoxus sphaeroidalis Revilla 1958 Theodoxus spratti Jenkins 1864 Theodoxus stoicai Neubauer Harzhauser Georgopoulou Mandic Kroh 2014 Bourguignat in Issel 1865 Theodoxus susuzianus Harzhauser Neubauer Hoşgör 2018 Theodoxus timisensis Jekelius 1944 Theodoxus trilophosensis Rust 1997 Theodoxus velascoi Graëlls 1846 V Anistratenko in O Anistratenko Starobogatov V Anistratenko 1999 Theodoxus vespertinus Sowerby II 1849 Theodoxus wesselinghi Sands Glöer 2020 Theodoxus wilkei Sands Glöer 2020 Theodoxus zlatarici Brusina 1902 Synonyms Theodoxus burdigalensis dOrbigny 1852 synonym of Theodoxus Theodoxus trifasciatus Grateloup 1839 Theodoxus cariosus Wood 1828 and Theodoxus cariosa are synonyms of Neritina cariosa Wood 1828 Goodwin D R 2006 The Discovery of Neritina Theodoxus cariosa Wood 1828 on the Island of Maui Hawaii Gastropoda Neritidae Visaya Net 11 pp PDF Theodoxus dacicus Jekelius 1944 synonym of Theodoxus Theodoxus leobersdorfensis dacicus Jekelius 1944 Theodoxus doriae Issel 1865 synonym of Theodoxus fluviatilis Linnaeus 1758 represented as Theodoxus Theodoxus fluviatilis Linnaeus 1758 Theodoxus euphraticus Mousson 1874 Mansoorian A 2001 Freshwater Gastropod of Khuzestan Province SouthWest Iran Iranian Int J Sci 2 2 9 pp PDF is a synonym of Mousson 1874Glöer P 2012 The freshwater snails Gastropoda of Iran with descriptions of two new genera and eight new species ZooKeys 219 1161 Theodoxus intracarpatica Jekelius 1944 synonym of Theodoxus intracarpaticus Jekelius 1944 Theodoxus lutetianus Montfort 1810 synonym of Theodoxus fluviatilis Linnaeus 1758 represented as Theodoxus Theodoxus fluviatilis Linnaeus 1758 Theodoxus mariae Handmann 1887 synonym of Neritina mariae Handmann 1887 Theodoxus martensi Brusina 1884 synonym of Ninnia martensi Brusina 1884 Theodoxus neglectus is a synonym of Neritina neglecta Pease 1861 Theodoxus oslavanensis Rzehak 1893 synonym of Theodoxus Theodoxus cyrtocelis austriacus Rzehak 1893 Theodoxus oualaniense Lesson 1831 synonym of Checked verified by a taxonomic editorClithon oualaniense Lesson 1831 Theodoxus perlongus Jekelius 1944 synonym of Theodoxus Theodoxus zografi perlongus Jekelius 1944 Theodoxus petralbensis Jekelius 1944 synonym of Theodoxus Theodoxus zografi petralbensis Jekelius 1944 Theodoxus pictus Férussac 1823 synonym of Agapilia picta Férussac 1823 Theodoxus reclivatus Say 1822 synonym of Röding 1798 Theodoxus rumanus Stefanescu 1896 synonym of Theodoxus Calvertia stefanescui Fontannes 1887 Theodoxus scamandri Calvert Neumayr 1880 synonym of Theodoxus Theodoxus scamandrius Calvert Neumayr 1880 Theodoxus vespertinus synonym of Neritina vespertinus Sowerby 1849 Theodoxus vetranici Brusina 1902 synonym of Neritina vetranici Brusina 1902 Theodoxus virginea Linnaeus 1758 synonym of Linnaeus 1758 Theodoxus Vittoclithon pictus bizonalis Grateloup 1839 synonym of Agapilia picta bizonalis Grateloup 1839 參考文獻 本條目包含源自公有領域的內容 Genus summary for Theodoxus last modified 24 February 2009 accessed 11 April 2011： Eichhorst TE 2016 Neritidae of the world Volume 2 Harxheim Conchbooks Pp 6961366 外部連結 Category 蜑螺科 Category 有未列明來源語句的條目