翼狀豬籠草（學名：），又名紅豬籠草，是菲律賓特有的熱帶食蟲植物。其種加詞 alata 來源於拉丁文 alatus，意為 翼狀的。翼狀豬籠草形態多樣性極高，其下類群分類地位不斷更新。 在園藝上，翼狀豬籠草是最容易栽培且栽培得最廣泛的物種之一 Catalano M 2009 Nepenthes In Growing Carnivores an Italian perspective Prague pp 5057，在臺灣曾廣泛栽培。但因其與葫蘆豬籠草的雜交種 紅瓶豬籠草更易種植且外觀更優越。園藝上，翼狀豬籠草逐漸被紅瓶豬籠草代替，已少有人栽培。然而，近期有觀點表明，園藝上曾被廣泛栽培的 翼狀豬籠草 已經不再屬於翼狀豬籠草這一物種，很有可能隨著翼狀豬籠草分類地位的更新被分配到了小花豬籠草名下。 形態特徵 翼狀豬籠草個體的顏色和形態差異巨大。雄花花程式為 K4 A441 G0（雄花頂端 可能不存在），雌花為 K4 A0 4。 分類關係 翼狀豬籠草長期以來被視為是一種形態多樣性極高且分佈範圍涵蓋整個菲律賓群島的物種（巴拉望島除外）。McPherson SR VB Amoroso 2011 Field Guide to the Pitcher Plants of the Philippines Redfern Natural History Productions Poole 在這個寬廣的界定下其分佈海拔 0 至 1900 米，範圍包括保和島、卡米金島、宿霧島、庫利昂島、萊特島、呂宋島、棉蘭老島、民都洛島、內格羅斯島、班乃島、薩馬島 Suarez W 2011 Samars Nepenthes alata Carnivorous Plants in the tropics January 12 2011Robinson A 2012 Nepenthes merrilliana on Samar Carnivorous Plants in the tropics June 29 2012 及錫布延島。 2013 年，馬修傑布與馬丁奇克重新分類了僅分佈於呂宋島北部及中部的物種，這些物種的捕蟲籠均具有顯著的毛被（這些類群在園藝上被稱之為 多毛翼狀豬籠草）。根據這種分類，分佈於更南部的類群則被指向一個被恢復的物種 小花豬籠草及新描述的內格羅斯島豬籠草與拉莫斯豬籠草。Cheek M M Jebb 2013 Nepenthes ramos Nepenthaceae a new species from Mindanao Philippines Willdenowia 43 1 107111 分佈於迪納加特島與薩馬島的綠豬籠草也為其近親 Micheler M T Gronemeyer A Wistuba D Marwinski W Suarez V Amoroso 2013 Nepenthes viridis eine neue NepenthesArt von der Insel Dinagat Philippinen Das Taublatt 76 421，此外還有塞西爾豬籠草、科普蘭豬籠草、絕滅豬籠草、漢密吉伊坦山豬籠草、奇坦蘭山豬籠草、倉田豬籠草、萊特島豬籠草、薩蘭加尼豬籠草及超基豬籠草。Cheek M M Jebb 2013 Recircumscription of the Nepenthes alata group Caryophyllales Nepenthaceae in the Philippines with four new species European Journal of Taxonomy 69 123 Cheek M M Jebb 2013 Nepenthes ultra Nepenthaceae a new species from Luzon Philippines Blumea published online on October 24 2013 Mey FS 2013 The Nepenthes alata group resurrection of N graciliflora N ramos and N negros described as new species Strange Fruits A Gardens Chronicle August 27 2013 這些物種屬於 翼狀豬籠草組 分類群，之間存在著許多共同特徵，比如葉柄具翼，捕蟲籠籠蓋下表面基部存在附屬物，及上位籠基部通常較寬。 翼狀豬籠草與許多其他的豬籠草物種間存在著密切的近緣關係，包括科普蘭豬籠草、棉蘭老島豬籠草 和薩蘭加尼豬籠草。產自蘇門答臘的真穗豬籠草曾一度因模式標本的誤釋而被認為是翼狀豬籠草的一個變種；實際上這兩個物種間並沒有密切的近緣關係。 翼狀豬籠草與真穗豬籠草之間形態特徵的區別 翼狀豬籠草與真穗豬籠草之間形態特徵的區別 翼狀豬籠草與真穗豬籠草之間形態特徵的區別 形態特徵 翼狀豬籠草 真穗豬籠草 葉片 披針形－卵形 披針形 葉尖 急尖或漸尖 圓形至近盾形 葉柄 呈槽狀 略呈或根本不呈槽狀 籠蔓尾 不分叉，尖銳 不分叉或分叉 毛被 紅色或白色 通體無毛 捕蟲籠基部形狀 結構與捕蟲籠的其餘部分類似，漸尖至籠蔓 角狀，木質，漸尖至籠蔓 種下類群 N alata f variegata Hort ex PMann 1996 nomnud N alata var biflora Macfarl 1908 Macfarlane JM 1908 Nepenthaceae In A Engler Das Pflanzenreich IV III Heft 36 191N negros N alata var ecristata Macfarl 1908N mindanaoensis 自然雜交種 N alata N burkeiMcPherson SR 2009 Pitcher Plants of the Old World 2 volumes Redfern Natural History Productions Poole N alata N merrilliana N merrilliata Mann P 1998 A trip to the Philippines Carnivorous Plant Newsletter 27 1 611 N alata N merrilliana N mirabilis N tsangoya N alata N mindanaoensis N alata N mirabilis N mirabilata Cheek MR MHP Jebb 2001 Nepenthaceae Flora Malesiana 15 1157Kurata S M Toyoshima 1972 Philippine species of Nepenthes The Gardens Bulletin Singapore 26 1 155158 Abstract N alata N petiolata N alata N pulchra N alata N truncata N truncalata N alata N ventricosa N ventrata Fleming R 1979 Carnivorous Plant Newsletter 8 1 1012 參考文獻 Alejandro GJD JPC Baysa BOC Lemana GM Madulara RS Madulid DA Madulid 2007 Conspecificity of Nepenthes alata Blco population found in Mt Guisguis Zambales inferred from Internal Transcribed Spacer nrDNA sequence data Acta Manilana 55 1521 Alejandro GJD RS Madulid DA Madulid 2008 The utility of Internal Transcribed Spacer nrDNA sequence data for phylogenetic reconstruction in endemic Philippine Nepenthes L Nepenthaceae The Philippine Scientist 45 99110 Amoroso VB RA Aspiras 2011 Hamiguitan Range a sanctuary for native flora Saudi Journal of Biological Sciences 18 1 715 Amoroso VB LD Obsioma JB Arlalejo RA Aspiras DP Capili JJA Polizon EB Sumile 2009 Inventory and conservation of endangered endemic and economically important flora of Hamiguitan Range southern Philippines Blumea 54 13 7176 An CI EI Fukusaki A Kobayashi 2001 Plasmamembrane HATPases are expressed in pitchers of the carnivorous plant Nepenthes alata Blanco Planta 212 4 547555 An CI EI Fukusaki A Kobayashi 2002 Aspartic proteinases are expressed in pitchers of the carnivorous plant Nepenthes alata Blanco Planta 214 5 661667 An CI S Takekawa A Okazawa EI Fukusaki A Kobayashi 2002 Degradation of a peptide in pitcher fluid of the carnivorous plant Nepenthes alata Blanco Planta 215 3 472477 Bauer U CJ Clemente T Renner W Federle 2012 Form follows function morphological diversification and alternative trapping strategies in carnivorous Nepenthes pitcher plants Journal of Evolutionary Biology 25 1 90102 Bauer U W Federle 2009 The insecttrapping rim of Nepenthes pitchers surface structure and function Plant Signaling Behavior 4 11 10191023 Benz MJ EV Gorb SN Gorb 2012 Diversity of the slippery zone microstructure in pitchers of nine carnivorous Nepenthes taxa ArthropodPlant Interactions 6 1 147158 Blanco FM 1845 Nepenthes In Flora de Filipinas segun el sistema sexual de Linneo Segunda impresion corregida y aumentada por el mismo autor D Miguel Sanchez Manila pp 555557 Buch F M Rott S Rottloff C Paetz I Hilke M Raessler A Mithöfer 2012 Secreted pitfalltrap fluid of carnivorous Nepenthes plants is unsuitable for microbial growth Annals of Botany published online on December 21 2012 Cantley R 2000 Nepenthes of the Philippines video The 3rd Conference of the International Carnivorous Plant Society San Francisco USA Cheers G 1992 Nepenthes alata In Letts Guide to Carnivorous Plants of the World Letts of London House London p 92 Clarke C CC Lee 2012 A revision of Nepenthes Nepenthaceae from Gunung Tahan Peninsular Malaysia Gardens Bulletin Singapore 64 1 3349 Clarke CM 2006 Introduction In Danser BH The Nepenthaceae of the Netherlands Indies Natural History Publications Borneo Kota Kinabalu pp 115 Co L W Suarez 2012 Nepenthaceae Cos Digital Flora of the Philippines Eilingsfeld A 2008 EndoskopAufnahmen aus der Verdauungsflüssigkeit einer Nepenthes alata Das Taublatt 60 1 47 Fontanilla JR R Madulid 2002 Morphometric analysis of Nepenthes alata Blco in Mt Guisguis Sta Cruz Zambales Acta Manilana 50 2531 Gaume L S Gorb N Rowe 2002 Function of epidermal surfaces in the trapping efficiency of Nepenthes alata pitchers New Phytologist 156 3 479489 Cheek MR MHP Jebb 1999 Nepenthes Nepenthaceae in Palawan Philippines Kew Bulletin 54 4 887895 Gorb EV K Haas A Henrich S Enders N Barbakadze S Gorb 2005 Composite structure of the crystalline epicuticular wax layer of the slippery zone in the pitchers of the carnivorous plant Nepenthes alata and its effect on insect attachment Journal of Experimental Biology 208 46514662 Gorb EV SN Gorb 2006 Physicochemical properties of functional surfaces in pitchers of the carnivorous plant Nepenthes alata Blanco Nepenthaceae Plant Biology Stuttgart Germany 8 6 841848 Gorb EV SN Gorb 2009 Chapter 8 Functional Surfaces in the Pitcher of the Carnivorous Plant Nepenthes alata A CryoSem Approach In SN Gorb ed Functional Surfaces in Biology Adhesion Related Phenomena Volume 2 Springer pp 205238 Gronemeyer T 2008 Nepenthes auf den Philippinen Ein Reisebericht Das Taublatt 60 1 1527 Gronemeyer T V Heinrich 2008 Wiederentdeckung von Nepenthes surigaoensis am Naturstandort auf den Philippinen Das Taublatt 60 1 2833 Hatano N T Hamada 2008 Proteome analysis of pitcher fluid of the carnivorous plant Nepenthes alata Journal of Proteome Research 7 2 809816 Hatano N T Hamada 2012 Proteomic analysis of secreted protein induced by a component of prey in pitcher fluid of the carnivorous plant Nepenthes alata Journal of Proteomics 75 15 48444852 Ishisaki K Y Honda H Taniguchi N Hatano T Hamada 2012 Heterogonous expression and characterization of a plant class IV chitinase from the pitcher of the carnivorous plant Nepenthes alata Glycobiology 22 3 345351 Ishisaki K S Arai T Hamada Y Honda 2012 Biochemical characterization of a recombinant plant class III chitinase from the pitcher of the carnivorous plant Nepenthes alata Carbohydrate Research 361 170174 Kajii E T Kamesaki S Ikemoto Y Miura 1988 Decomposing enzymes against human bloodgroup antigens in the extract of Nepenthes alata Die Naturwissenschaften 75 5 258259 Kajii E T Kamesaki S Ikemoto 1991 The effect of the Nepenthes alata extract on the cold agglutininassociated antigens Nihon Hōigaku Zasshi 45 1 3032 Kamesaki T E Kajii S Ikemoto 1989 Purification of the decomposing enzyme from Nepenthes alata against glycophorin B of human red blood cells by highperformance liquid chromatography Journal of Chromatography 489 2 384389 Kato M M Hotta R Tamin T Itino 1993 Inter and intraspecific variation in prey assemblages and inhabitant communities in Nepenthes pitchers in Sumatra Tropical Zoology 6 1 1125 Abstract Kiew RG 1990 Pitcher plants of Gunung TahaN Journal of Wildlife and National Parks Malaysia 10 3437 Lecoufle M 1990 Nepenthes alata In Carnivorous Plants Care and Cultivation Blandford London pp 118121 Macfarlane JM 1927 The Philippine species of Nepenthes The Philippine Journal of Science 33 2 127140 Mansur M 2001 In Prosiding Seminar Hari Cinta Puspa dan Satwa Nasional Lembaga Ilmu Pengetahuan Indonesia Bogor pp 244253 McPherson S T Gronemeyer 2008 Die Nepenthesarten der Philippinen Eine Fotodokumentation Das Taublatt 60 1 3478 Meimberg H P Dittrich G Bringmann J Schlauer G Heubl 2000 Molecular phylogeny of Caryophyllidae sl based on matK sequences with special emphasis on carnivorous taxa Plant Biology 2 2 218228 Meimberg H A Wistuba P Dittrich G Heubl 2001 Molecular phylogeny of Nepenthaceae based on cladistic analysis of plastid trnK intron sequence data Plant Biology 3 2 164175 Meimberg H 2002 PhD thesis Ludwig Maximilian University of Munich Munich Meimberg H G Heubl 2006 Introduction of a nuclear marker for phylogenetic analysis of Nepenthaceae Plant Biology 8 6 831840 Merrill ED 1905 Nepenthaceæ In A review of the identifications of the species described in Blancos Flora de Filipinas Bureau of Public Printing Manila p 72 Merrill ED 1918 Nepenthaceae In Species Blancoanae a critical revision of the Philippine species of plants described by Blanco and by Llanos Bureau of Printing Manila p 160 Mey FS 2013 Neotypification of Nepenthes blancoi and description of N abalata a new species from the Philippines Strange Fruits A Gardens Chronicle February 11 2013 Mey FS 2013 Nepenthes alzapan and an army of new species from the Philippines Strange Fruits A Gardens Chronicle July 15 2013 Mithöfer A 2011 Carnivorous pitcher plants insights in an old topic Phytochemistry 72 13 16781682 Owen TP KA Lennon 1999 American Journal of Botany 86 10 13821390 Owen TP KA Lennon MJ Santo AN Anderson 1999 Annals of Botany 84 459466 Pavlovič A E Masarovičová J Hudák 2007 Carnivorous syndrome in Asian pitcher plants of the genus Nepenthes Annals of Botany 100 3 527536 Renner T CD Specht 2011 A sticky situation assessing adaptations for plant carnivory in the Caryophyllales by means of stochastic character mapping International Journal of Plant Sciences 172 7 889901 Riedel M A Eichner R Jetter 2003 Slippery surfaces of carnivorous plants composition of epicuticular wax crystals in Nepenthes alata Blanco pitchers Planta 218 1 8797 Rottloff S R Stieber H Maischak FG Turini G Heubl A Mithöfer 2011 Functional characterization of a class III acid endochitinase from the traps of the carnivorous pitcher plant genus Nepenthes Journal of Experimental Botany 62 13 46394647 Santo MJ JS Massa TP Owen 1998 Glandular secretion and absorption in the carnivorous pitcher plant Nepenthes alata American Journal of Botany 85 supplement 92 Schlauer J 2000 Carnivorous Plant Newsletter 29 2 53 Scholz I M Bückins L Dolge T Erlinghagen A Weth F Hischen J Mayer S Hoffmann M Riederer M Riedel W Baumgartner 2010 Slippery surfaces of pitcher plants Nepenthes wax crystals minimize insect attachment via microscopic surface roughness Journal of Experimental Biology 213 11151125 Slack A 1979 Nepenthes alata In Carnivorous Plants Ebury Press London pp 8586 Sota T M Mogi K Kato 1998 Local and regionalscale food web structure in Nepenthes alata pitchers Biotropica 30 1 8291 Syamsuardi R Tamin 1994 Kajian kekerabatan jenisjenis Nepenthes di Sumatera Barat Project report Andalas University Padang Abstract Syamsuardi 1995 Klasifikasi numerik kantong semar Nepenthes di Sumatera Barat Numerical classification of pitcher plants Nepenthes in West Sumatra Journal Matematika dan Pengetahuan Alam 4 1 4857 Abstract Thornhill AH IS Harper ND Hallam 2008 The development of the digestive glands and enzymes in the pitchers of three Nepenthes species N alata N tobaica and N ventricosa Nepenthaceae International Journal of Plant Sciences 169 5 615624 Uy BD 1990 Carnivorous Plant Newsletter 19 12 1819 Wang LX Q Zhou 2010 Numerical characterization of surface structures of slippery zone in Nepenthes alata pitchers and its mechanism of reducing locusts attachment Advances in Natural Science 3 2 152160 Wu X J Li Q Zhong X Wu 2005 豬籠草組培快繁技術 Tissue culture and fast propagation in Nepenthes alata Nonwood Forest Research 23 4 4850 擴充套件閱讀 食蟲植物照片搜尋引擎中翼狀豬籠草的照片 Nepenthes alata alata Category 菲律賓植物