Question:  What species has an extinct subspecies that are classified as Trogloxene and are known from fossils and prehistoric art?

Passages:
Bernard's wolf ("Canis lupus bernardi"), also known as the Banks Island tundra wolf, Banks Island wolf, and the Victoria's Island wolf, was a subspecies of the gray wolf, "Canis lupus", that was limited to the Banks and Victoria Island of the Canadian arctic.  An extinct subspecies, it was described as "white with black-tipped hair along the ridge of the back".  It was formally discovered, classified, and named after Peter Bernard and Joseph F. Bernard, his nephew, after an adult male skin and skull was collected by them and brought to the National Museum of Canada.  There were very few specimens of this subspecies that were recovered, around three or four in total.  A survey was conducted in March 1993 by the Department of Renewable Resources that was to catalog the wolf and caribou populations of the area.  While a number of caribou were found and recorded, along with many other indigenous animal species, not a single wolf was found.  The Victoria's Island population is believed to have died out in between 1918 and 1952.  They were previously widespread in their native habitat but were annihilated by excessive hunting. <P2-S0> The Merriam's elk ("Cervus canadensis merriami") is an extinct subspecies of elk once found in the arid lands of the southwestern United States, predominantly Arizona. <P2-S1>  Since the arrival of the Europeans uncontrolled hunting and cattle grazing had driven the subspecies into extinction around the beginning of the 20th century, with the exact presumed date being 1906. <P2-S2>  Another subspecies of elk, the eastern elk ("Cervus canadensis canadensis") also became extinct at roughly the same time. <P2-S3>  Not much else is known about this subspecies as it became extinct before studies were done. <P2-S4>  Elk from Yellowstone National Park were introduced to this area in 1913, and are reasonably common in the area today. Rhinoceros sondaicus annamiticus is an extinct subspecies of "Rhinoceros sondaicus", that lived in Vietnam, Laos, Cambodia, Thailand and Malaysia.  The term Annamiticus derives from the Annamite name of the Indochinese Mountains in Indochina, part of the distribution of this species, also known as the Javanese Javan rhinoceros or simply Vietnamese rhinoceros for Vietnam.  Only this species, the Stieng called them Pai-mhai once lived across South China, Vietnam, Cambodia, Laos, Thailand, and Malaysia.  The subspecific annamiticus is derived from the Annamite Mountain Range in Southeast Asia, part of this subspecies' range.  In 2006, a single population, estimated at fewer than 12 remaining rhinos, lived in an area of lowland forest in the Cat Tien National Park in Vietnam.  Genetic analysis suggested this subspecies and the Indonesian Javan rhinoceros last shared a common ancestor between 300,000 and 2 million years ago.  The last individual of this population was shot by a poacher in 2010. Portable art (sometimes called mobiliary art) refers to the small examples of Prehistoric art that could be carried from place to place, which is especially characteristic of the Art of the Upper Palaeolithic.  It is one of the two main categories of Prehistoric art, the other being the immobile Parietal art, effectively synonymous with rock art. As of September 2016, the International Union for Conservation of Nature (IUCN) lists 116 extinct species, 132 possibly extinct species, 35 extinct in the wild species, 13 possibly extinct in the wild species, five extinct subspecies, one extinct in the wild subspecies, and four extinct varieties of plant. <P3-S0> Prehistoric art in Scotland is visual art created or found within the modern borders of Scotland, before the departure of the Romans from southern and central Britain in the early fifth century CE, which is usually seen as the beginning of the early historic or Medieval era. <P3-S1>  There is no clear definition of prehistoric art among scholars and objects that may involve creativity often lack a context that would allow them to be understood. As of September 2016, the International Union for Conservation of Nature (IUCN) lists 310 extinct species, 117 possibly extinct species, 14 extinct in the wild species, eight extinct subspecies, and five extinct in the wild subspecies of mollusc. Trogloxenes, also called cave guests, are animal species which live close to caves or at the very entrance of the cave, but cannot live exclusively in a cave.  This group consists of both extant animals, such as raccoons, rats, foxes and bears, opiliones (a.k.a. Daddy-Long-Legs) and extinct animals like cave bears, cave lions, cave leopards, and cave hyenas.  Indications trusted by geologists and archaeologists combine to show that these animals lived there in the latter part, at least, of the third interglacial epoch, and on through the fourth and last glacial advance, when, although central Europe was free from an ice cap, an almost Arctic climate prevailed, with much rain.  This is what is known as the Upper Paleolithic, when humanity in Europe was represented by the Neanderthals. As of July 2016, the International Union for Conservation of Nature (IUCN) lists 394 extinct species, 206 possibly extinct species, 16 extinct in the wild species, eight extinct subspecies, and five extinct in the wild subspecies of invertebrate. Panthera leo spelaea or P. spelaea, commonly known as the European or Eurasian cave lion, is an extinct subspecies of lion.  It is known from fossils and many examples of prehistoric art.

Supporting sentences:
Panthera leo spelaea or P. spelaea, commonly known as the European or Eurasian cave lion, is an extinct subspecies of lion.  It is known from fossils and many examples of prehistoric art.