doctoral_advisor thesis_year death_date death_place birth_place thesis_title awards nationality fields workplaces birth_date thesis_url known_for spouse image article_title name alma_mater
richard gardner virginia papaioannou 1981 18 may 2001 great tew , oxfordshire hampshire studies postimplantation mammalian embryo on cell fate and cell potency in the waddington 1999 -rrb- medal -lrb- 1999 -rrb- lister fellowship frs -lrb- united kingdom developmental biology university harbor laboratory national institute for medical research centre for genome of research , edinburgh oxford imperial cancer research fund cold spring 23 march 1956 http://ethos.bl.uk/orderdetails.do?uin=uk.bl.ethos.237750 anterior-posterior patterning in mammalian embryos robin denniston rosa beddington -lrb- 1956-2001 -rrb- . tif rosa beddington
 rosa beddington brasenose college , oxford -lrb- ba , phd -rrb-
rosa beddington

rosa susan penelope beddington frs -lrb- 23 march 1956 -- 18 may 2001 -rrb- was a british biologist whose career had a major impact on developmental biology .
