caption death_date death_place birth_place image_size nationality field work_institutions birth_date known_for image article_title name
edouard van beneden 28 april 1910 liège leuven 150px belgian embryologist university of liège 5 march 1846 meiosis edouard musée de zoologie.jpg van beneden in front of the aquarium et edouard van beneden
 edouard van beneden
edouard van beneden

Édouard joseph louis marie van beneden -lrb- 5 march 1846 -- 28 april 1910 -rrb- , son of pierre-joseph van beneden , was a belgian embryologist , cytologist and marine biologist .
he was professor of zoology at the university of liège .
he contributed to cytogenetics by his works on the roundworm `` ascaris '' .
in this work he discovered how chromosomes organized meiosis -lrb- the production of gametes -rrb- .
van beneden elucidated , together with walther flemming and eduard strasburger , the essential facts of mitosis , where , in contrast to meiosis , there is a qualitative and quantitative equality of chromosome distribution to daughter cells .
-lrb- see karyotype -rrb- .
