 Furthermore, prosocial behaviors can evolve that accumulate most of the benefit in a few highly connected nodes while many peripheral nodes receive low or negative payoff.
<coherence>  Furthermore, prosocial behaviors can evolve that accumulate most of the benefit in a few highly connected nodes while many peripheral nodes receive low or negative payoff.
 However, they often accumulate most of the benefit in a few highly connected nodes while many peripheral nodes receive low or negative payoff.