 First, we consider the movements of susceptible (S), exposed (E), and recovered (R) individuals who may get infected after movement , as well as infected (I)  . Second, the infectious contacts between people from different regions are dealt with separately from the first factor.
<meaning-changed>  First, we consider the movements of susceptible (S), exposed (E), and recovered (R) individuals who may get infected after movement , as well as infected (I)  . Second, the infectious contacts between people from different regions are dealt with separately from the first factor.
 First, we consider the movements of susceptible (S), exposed (E), and recovered (R) individuals who may get infected and infect others in the destination region , as well as infected (I) . Second, the infectious contacts between people from different regions are dealt with separately from the first factor.