 While manipulated fire return interval and seasonality moderated both fire spread  and intensity, their overall effects were low in comparison with  hydrological and climatic  variables.
<meaning-changed>  While manipulated fire return interval and seasonality moderated both fire spread  and intensity, their overall effects were low in comparison with  hydrological and climatic  variables.
 While manipulated fire return interval and seasonality moderated both fire spread and intensity, their overall effects were low in comparison with meteorological ( hydrological and climatic variables.