 It has however been suggested that knuckle-walking evolved independently and separately in Pan and gorillas and so was not present in the human ancestors.
<coherence>  It has however been suggested that knuckle-walking evolved independently and separately in Pan and gorillas and so was not present in the human ancestors.
 Knuckle-walking, though has been suggested to have evolved independently and separately in Pan and gorillas and so was not present in the human ancestors.