 Our hypothesis leads to an experimentally testable prediction that a system with both types of invariance detects the relative steepness even in spatio-temporal gradient stimuli such as waves .
<meaning-changed>  Our hypothesis leads to an experimentally testable prediction that a system with both types of invariance detects the relative steepness even in spatio-temporal gradient stimuli such as waves .
 Our hypothesis leads to an experimentally testable prediction that a system with both types of invariance detects the relative steepness even in dynamic gradient stimuli as well as in static gradients .