 Furthermore, prosocial behaviors can evolve that accumulate most of the benefit in a few highly connected nodes while many peripheral nodes receive low or negative payoff.
<clarity>  Furthermore, prosocial behaviors can evolve that accumulate most of the benefit in a few highly connected nodes while many peripheral nodes receive low or negative payoff.
 Furthermore, prosocial behaviors can evolve that accumulate most of the benefits in the hands of a few highly-connected individuals, while many others receive low or negative payoff.