 It can also occur  between species with low contact rates if facilitated by an intermediary species . For example, bats can directly transfer the rabies virus to humans through bites, and also through aerosolization of bat saliva and urine which can then be absorbed by human mucous membranes in the nose, mouth, and eyes. A host shifting event occurs when a strain that was previously zoonotic now circulates exclusively among humans.
<coherence>  It can also occur  between species with low contact rates if facilitated by an intermediary species . For example, bats can directly transfer the rabies virus to humans through bites, and also through aerosolization of bat saliva and urine which can then be absorbed by human mucous membranes in the nose, mouth, and eyes. A host shifting event occurs when a strain that was previously zoonotic now circulates exclusively among humans.
 It can also occur between species with low contact rates if facilitated by an intermediary species 