 However, open questions about URLanization mechanisms in animal groups remain: Evolutionary adaptation towards  group-level optima (group-level selection), often implicitly assumed in the criticality hypothesis, is in general not a reasonable mechanism in fission-fusion groups composed of non-related individuals.
<fluency>  However, open questions about URLanization mechanisms in animal groups remain: Evolutionary adaptation towards  group-level optima (group-level selection), often implicitly assumed in the criticality hypothesis, is in general not a reasonable mechanism in fission-fusion groups composed of non-related individuals.
 However, open questions about URLanization mechanisms in animal groups remain: Evolutionary adaptation towards a group-level optima (group-level selection), often implicitly assumed in the criticality hypothesis, is in general not a reasonable mechanism in fission-fusion groups composed of non-related individuals.