 While manipulated fire return interval and seasonality moderated both fire spread  and intensity, their overall effects were low in comparison with  hydrological and climatic  variables.
<meaning-changed>  While manipulated fire return interval and seasonality moderated both fire spread  and intensity, their overall effects were low in comparison with  hydrological and climatic  variables.
 While manipulated fire return interval and seasonality moderated both fire spread rate and intensity, their overall effects were low in comparison with hydrological and climatic variables.