 To suppress recombination, balancers usually harbor multiple, nested chromosomal inversions so that synapsis between  homologous chromosomes is disrupted.
<fluency>  To suppress recombination, balancers usually harbor multiple, nested chromosomal inversions so that synapsis between  homologous chromosomes is disrupted.
 To suppress recombination, balancers usually harbor multiple, nested chromosomal inversions so that synapsis between the homologous chromosomes is disrupted.