Pterostichinae
Pterostichinae is a subfamily of ground beetles (family Carabidae). It belongs to the advanced harpaline assemblage, and if these are circumscribed "sensu lato" as a single subfamily, Pterostichinae are downranked to a tribe Pterostichini. However, as the former Pterostichitae supertribe of the Harpalinae as loosely circumscribed does seem to constitute a lineage rather distinct from "Harpalus", its core group is here considered to be the present subfamily and the Harpalinae are defined more narrowly. They are usually mid-sized and rather stout ground beetles. Coloration is typically dark and without conspicuous patterns, but often with a strong sheen like  polished metal. They are widely distributed and inhabit a wide range of terrestrial habitats. Unlike the more basal ground beetles which only eat small animals, the Pterostichinae include a large proportion of omnivorous or even herbivorous taxa. Systematics. This group includes the following tribes and genera: Tribe Abacetini Tribe Catapieseini Tribe Chaetodactylini Tribe Cnemalobini Tribe Cratocerini Tribe Drimostomatini Tribe Microcheilini Tribe Morionini Tribe Pterostichini (including Metiini) Tribe Zabrini