Linda Holland
Linda Zimmerman Holland is a research biologist at Scripps Institution of Oceanography known for her work examining the evolution of vertebrates. Education and career. Holland has a B.A. (1962) and an M.A. (1964) from Stanford University. She worked as a research associate at the University of California, San Diego, Scripps Clinic, and Scripps Institution of Oceanography in the period from 1970 until 1998. She earned her Ph.D. from the University of California, San Diego in 2001. She started as a research biologist at Scripps in 1998, and is emeritus as of 2022. Research. Holland's early research examined  anatomical structures in purple sea urchins, and a protein involved in clotting, Von Willebrand factor. She went on to examine reproduction in sea urchins, salps, and amphioxus, known as lancelet. Holland began collecting amphioxus in Tampa, Florida in 1988, which enabled her to use them as a model system to study evolutionary biology. Holland was the lead scientist on the project analyzing the genome of amphioxus, and her work revealed reuse and copying of genes by amphioxus. Her research also addressed the evolution of bilaterian animals as in her 2013 Holland et al. publication (see image). In 2017 she wrote  a history of the use of amphioxus in biological research. Awards and honors. In 2014 Holland, and her husband Nick Holland, received the A.O. Kovalevsky Medal for their work on amphioxus. This award also includes being named an honorary member of the Saint Petersburg Society of Naturalists.