Eutetramorium
Eutetramorium is a small genus of ants within the subfamily Myrmicinae. To date it contains three species found in Madagascar and Comoros. Description. The queen is extremely ergatoid in "E. mocquerysi", alate in "E. monticellii". In "E. parvum" a normal dealate queen is present in the type-series, but there is also a specimen that may be an ergatoid or a worker-queen intercaste, or possibly merely an oversized worker. Males are known only for "E. mocquerysi", where it is smaller than conspecific workers. Taxonomy. This small genus was established by Emery (1899) for two conspicuous, large, darkly coloured  Madagascan species that superficially resemble "Tetramorium". Initially, Emery (1912, 1914) was of the opinion that the genus was referable to the tribe Myrmecinini, but Ashmead (1905) and Wheeler (1910) had already referred it to Tetramoriini, an opinion that Emery (1915, 1924) also came to accept. Bolton (1976) excluded "Eutetramorium" from Tetramoriini on morphological grounds, and tentatively transferred it to Myrmicini. This placement persisted until recently, when a molecular analysis of Myrmicini by Jansen & Savolainen (2010) showed that the monophyly of the tribe was dubious, and that "Eutetramorium" formed a clade with "Huberia". The yet unpublished, more detailed, DNA analysis of  Myrmicinae by Philip S. Ward has established that "Eutetramorium" belongs in a strongly supported, endemic Madagascan clade that also includes the other four genera: "Malagidris", "Myrmisaraka", "Royidris" and "Vitsika".