Coelodonta
Coelodonta (, from the Greek κοιλία, "koilía" and οδούς, "odoús", "hollow tooth", in reference to the deep grooves of their molars) is an extinct genus of rhinoceros that lived in Eurasia between 3.7 million years to 10,000 years before the present, in the Pliocene and the Pleistocene epochs. It is best known from the type species, the woolly rhinoceros ("Coelodonta antiquitatis"), which ranged throughout northern Eurasia during the Pleistocene. The earliest known species, "Coelodonta thibetana," lived in Tibet during the Pliocene, with the genus spreading to the rest of Eurasia during the Pleistocene. Species. Species recognised as members of "Coelodonta , according to Deng et al". (2011), include: Phylogeny. The authors of the description of the species "C. thibetana" (Deng "et al"., 2011) proposed a cladogram to place phylogenetically their position in relation to other members of Rhinocerotidae, using the five extant species of rhinoceros and 13 extinct species. They found that "Coelodonta" was the sister taxon of the species "Stephanorhinus hemitoechus". A 2019 study based on a 1.78 million year old "Stephanorhinus. sp" dental proteome from Georgia suggested the paraphyly of "Stephanorhinus." Later DNA studies have found the woolly rhinoceros is closely related to Merck's rhinoceros ("Stephanorhinus kirchbergensis"), with  an estimated divergence 5.5 million years ago. These two species in turn are most closely related to the Sumatran rhinoceros ("Dicerorhinus sumatrensis") amongst living rhinoceros. Cladogram after Liu et al, 2021: