Sphaerotilus natans
Sphaerotilus natans is an aquatic periphyton organism associated with polluted water. It forms colonies commonly known as "sewage fungus", but later identified as tightly sheathed filamentous bacteria. Morphology. Straight or smoothly curved filaments 1.5 µm in diameter and 100 to more than 500 µm in length are formed by rod-shaped cells with clear septa growing within a long, tubular sheath. An adhesive basal element at one end of the filament can aid attachment to solid surfaces. The sheath offers some protection from predators, and the ability to anchor in flowing water allows access to a passing stream  of food and nutrients. Individual mature cells swarm out of the protective tube to colonize new sites. Each motile mature cell has an intertwined bundle of flagella appearing as a single flagellum consisting of a long filament with a short hook and a basal body complex, but it is distinguishable by electron microscope as 10 to 30 strands with diameters of 12.5 to 16 nm each. "S. natans" stores reserves of poly- "beta" -hydroxybutyrate as internal globules making up 30 to 40% of the dry weight of a colony. Gram and Neisser staining reactions are negative. Habitat. "S. natans" requires  dissolved simple sugars or organic acids as a food supply, but needs less phosphorus than many competing organisms and can tolerate low oxygen concentrations. Capability to deposit elemental sulfur intracellularly in the presence of hydrogen sulfide is believed to be a detoxifying mechanism. "S. natans" requires either cobalamin or methionine as a trace nutrient. "S. natans" filaments can aid development of a periphyton biofilm trapping suspended particles and stabilizing colonies of other organisms including "Klebsiella" and "Pseudomonas". Significance. "Sphaerotilus natans" is often associated with a buoyant floc (or "bulking sludge") causing poor solids separation in activated sludge clarifiers of secondary  sewage treatment. Metal surfaces covered with "S. natans" may experience accelerated corrosion if the slime creates a barrier causing differential oxygen concentrations. "S. natans" slimes may reduce quality of paper produced by paper mills using recycled water streams.