Tomiostrobus
Tomiostrobus is an extinct quillwort genus from the Early Triassic of Australia and Russia, which was especially widespread in the aftermath of Permian Triassic mass extinctions. Description. "Tomiostrobus australis" is preserved as whole plants closely spaced within bedding planes, and lived as an early successional weed in lake and pond sedimentary environments, like living "Isoetes". Unlike living "Isoetes", "Tomiostrobus" formed closed cones with sporophylls that were distinctly shouldered and woody. This may have been an adaptation to heavy grazing by herbivorous therapsids.