Beneath the surface litter layer, a rich complexity of soil organisms decompose plant residue and dead roots, then release their stored nutrients slowly over time.
(a rich complexity of soil organisms; decompose then; L:Beneath the surface litter layer) Context(a rich complexity of soil organisms decompose then,List([34, 114))):(a rich complexity of soil organisms; decompose then release; their stored nutrients; T:slowly over time)
a rich complexity of soil organisms decompose then,List([34, 114))
a rich complexity of soil organisms a rich complexity of soil organisms
decompose then decompose then release
L:Beneath the surface litter layer
their stored nutrients T:slowly over time