Thus, protected areas exclude most of the warm temperate broad-leaved forest, which is the major habitat of the macaques and deer.
(protected areas; exclude; most of the warm temperate broad-leaved forest) (the warm temperate broad-leaved forest; is; the major habitat of the macaques and deer)
protected areas the warm temperate broad-leaved forest
exclude is
most of the warm temperate broad-leaved forest
the major habitat of the macaques and deer