He then carefully transferred pollen from the anthers of another plant and using a small paintbrush, dusted the pollen onto the stigma of the flowers that lacked anthers.
(He; carefully transferred; pollen; from the anthers of another plant; T:then) (He; using; a small paintbrush) (He; dusted; the pollen; onto the stigma of the flowers) (the flowers; lacked; anthers)
He He He the flowers
carefully transferred using dusted lacked
pollen from the anthers of another plant T:then
a small paintbrush
the pollen onto the stigma of the flowers
anthers