Human-caused changes, such as livestock grazing, predator control, mule deer population growth, and fire suppression have contributed to the change.
(Human-caused changes, such as livestock grazing, predator control, mule deer population growth, and fire suppression; have contributed; to the change)
Human-caused changes, such as livestock grazing, predator control, mule deer population growth, and fire suppression
have contributed
to the change