This impact on the coral reefs (measure of impact on a resource) can be measured in a number of the components of the reef ecosystem including coral growth, survival, spawning/settlement at selected sites as well as other reef inhabitants (e.g., fish).
(This impact on the coral reefs; can be measured; in a number of the components of the reef ecosystem including coral growth, survival, spawning/settlement at selected sites as well as other reef inhabitants (e.g., fish)
This impact on the coral reefs
can be measured
in a number of the components of the reef ecosystem including coral growth, survival, spawning/settlement at selected sites as well as other reef inhabitants (e.g., fish