8) This paper makes it clear that at least two applications of glyphosate are required with the use of such GM crops - some of the study data, however, would seem to imply that in practice three applications may be required for 
(This paper; makes; it clear that at least two applications of glyphosate are required with the use of such GM crops - some of the study data) (at least two applications of glyphosate; are required; with the use of such GM crops)
This paper at least two applications of glyphosate
makes are required
it clear that at least two applications of glyphosate are required with the use of such GM crops - some of the study data
with the use of such GM crops