They wanted to determine how different species dispersed as the climate changed.
(how different species; dispersed; T:as the climate changed) (the climate; changed; ) (They; wanted; to determine how different species) Context(They wanted,List([0, 11))):(They; wanted to determine; how different species dispersed)
They wanted,List([0, 11))
how different species the climate They They
dispersed changed wanted wanted to determine
T:as the climate changed
to determine how different species
how different species dispersed