The crop traits that would be most useful to subsistence farmers tend to be very complex.
(The crop traits; would be; most useful to subsistence farmers) (The crop traits that would be most useful to subsistence farmers; tend; to be very complex) (The crop traits that would be most useful to subsistence farmers; to be; very complex)
The crop traits The crop traits that would be most useful to subsistence farmers The crop traits that would be most useful to subsistence farmers
would be tend to be
most useful to subsistence farmers
to be very complex
very complex