Costal short-cells neither distinctly grouped into long rows nor predominantly paired (&#145;short-cells&#146; much longer than usual, and the files interrupted by prickles).
(Costal short-cells; distinctly grouped; into long rows nor predominantly paired (&#145;short-cells&#146; much longer than usual, and the files) (Costal short-cells; predominantly paired; ) (the files; interrupted; by prickles)
Costal short-cells Costal short-cells the files
distinctly grouped predominantly paired interrupted
into long rows nor predominantly paired (&#145;short-cells&#146 much longer than usual, and the files
by prickles