Numbers of all stages of M. occidentalis and T. urticae were counted under a dissecting microscope, and the mean number of mites per leaf was determined for each row in each plot.
(Numbers of all stages of M. occidentalis and T. urticae; were counted; ) (the mean number of mites per leaf; was determined; )
Numbers of all stages of M. occidentalis and T. urticae the mean number of mites per leaf
were counted was determined