This led to significant involvement in major ecosystem level studies in grassland systems ranging from tundra regions to temperate zone prairies to the tropics.
(This; led; to significant involvement in major ecosystem level studies in grassland systems) (grassland systems; ranging; from tundra regions; to temperate zone prairies)
This grassland systems
led ranging
to significant involvement in major ecosystem level studies in grassland systems
from tundra regions to temperate zone prairies