Although climatic factors may be a proximate factor in regional salmon decline, the ultimate causes may involve a longer history of change affecting the resilience of species and populations.
(climatic factors; may be; a proximate factor in regional salmon decline) (the ultimate causes; may involve; a longer history of change) (change; affecting; the resilience of species and populations)
climatic factors the ultimate causes change
may be may involve affecting
a proximate factor in regional salmon decline
a longer history of change
the resilience of species and populations