Red-backed voles have dense, long, soft fur in winter but this changes to shorter, coarser fur in summer.
(Red-backed voles; have; dense, long, soft fur in winter but this changes to shorter, coarser fur in summer) (this; changes; to shorter, coarser fur in summer)
Red-backed voles this
have changes
dense, long, soft fur in winter but this changes to shorter, coarser fur in summer
to shorter, coarser fur in summer