To adapt, they had to either remain in low salinity environments, such as bays and estuaries, or they had to evolve mechanisms to replace water lost through osmosis to the seawater and to remove salts absorbed from the increasingly saline oceans.
(mechanisms; to replace; water lost through osmosis to the seawater) (water; lost; to the seawater) (they; to remain; in low salinity environments, such as bays and estuaries) (salts; absorbed; from the increasingly saline oceans) (they; to evolve; mechanisms to replace water lost through osmosis to the seawater and to remove salts)
mechanisms water they salts they
to replace lost to remain absorbed to evolve
water lost through osmosis to the seawater
to the seawater
in low salinity environments, such as bays and estuaries
from the increasingly saline oceans
mechanisms to replace water lost through osmosis to the seawater and to remove salts