These are produced by small caterpillars that first feed on the sap, producing a series of small dots visible on the upper leaf surface.
(These; are produced; by small caterpillars) (small caterpillars; feed; on the sap; T:first) (small caterpillars; producing; a series of small dots visible on the upper leaf surface)
These small caterpillars small caterpillars
are produced feed producing
by small caterpillars
on the sap T:first
a series of small dots visible on the upper leaf surface