One-third to one-half of acorn losses have been attributed to removal by birds and squirrels while the acorns were still on the tree.
(One-third to one-half of acorn losses; have been attributed; to removal by birds and squirrels; T:while the acorns were still on the tree) (the acorns; were; T:still; on the tree)
One-third to one-half of acorn losses the acorns
have been attributed were
to removal by birds and squirrels T:while the acorns were still on the tree
T:still on the tree