Spines on the stem are straight but those on the fruits are hooked, enabling seed dispersal in animal fur, wool etc. Leaf underside is covered with dense grey-white tomentum.
(Spines on the stem; are; straight) (Spines on the stem; are; hooked) (those on the fruits; enabling; seed dispersal in animal fur, wool) Context(those on the fruits enabling,List([36, 76))):(etc. Leaf underside; is covered; with dense grey-white tomentum)
those on the fruits enabling,List([36, 76))
Spines on the stem Spines on the stem those on the fruits etc. Leaf underside
are are enabling is covered
straight
hooked
seed dispersal in animal fur, wool
with dense grey-white tomentum