However, what traits were inherited, the specific particles of their inheritance, and how these particles acted to control or regulate the development of phenotypic traits were all unknown.
(the specific particles of their inheritance, and how these particles acted to control or regulate the development of phenotypic traits; were; unknown) (these particles; acted; to control or regulate the development of phenotypic traits) Context(these particles acted,List([90, 111))):(these particles; acted to control; ) Context(these particles acted,List([90, 111))):(these particles; acted to regulate; the development of phenotypic traits)
these particles acted,List([90, 111)) these particles acted,List([90, 111))
the specific particles of their inheritance, and how these particles acted to control or regulate the development of phenotypic traits these particles these particles these particles
were acted acted to control acted to regulate
unknown
to control or regulate the development of phenotypic traits
the development of phenotypic traits