As time went by, fatty, stale-smelling substances accumulated in both breads, but the traditional bread was more protected from staleness by its greater store of malty compounds.
(time; went; by) (stale; smelling; substances accumulated in both breads, but the traditional bread was more protected from staleness by its greater store of malty compounds) (fatty, stale-smelling substances; accumulated; L:in both breads) (the traditional bread; was more protected; from staleness; by its greater store of malty compounds)
time stale fatty, stale-smelling substances the traditional bread
went smelling accumulated was more protected
by
substances accumulated in both breads, but the traditional bread was more protected from staleness by its greater store of malty compounds
L:in both breads
from staleness by its greater store of malty compounds