Variability in the latter can be almost completely explained by year-to-year changes in climate and river flow, and a discussion is included in the aforementioned manuscript.
(Variability in the latter; can be explained; by year-to-year changes in climate and river flow) (a discussion; is included; in the aforementioned manuscript)
Variability in the latter a discussion
can be explained is included
by year-to-year changes in climate and river flow
in the aforementioned manuscript