The decrease in soil nutrient supply may appear to reduce the suitability of the soil for the tree, but the overall effect could be an increase in tree fitness.
(The decrease in soil nutrient supply; may appear; ) (The decrease in soil nutrient supply; to reduce; the suitability of the soil for the tree) (the overall effect; could be; an increase in tree fitness)
The decrease in soil nutrient supply The decrease in soil nutrient supply the overall effect
may appear to reduce could be
the suitability of the soil for the tree
an increase in tree fitness