One of their goals was to determine which ecosystems, among deserts, grasslands and forests, had more variable production and which responded most to precipitation variability.
(One of their goals; was; to determine which ecosystems, among deserts, grasslands and forests, had more variable production and which responded most to precipitation variability)
One of their goals
was
to determine which ecosystems, among deserts, grasslands and forests, had more variable production and which responded most to precipitation variability