Reptile and amphibian populations at these sites are much smaller in size than one would expect based on the habitat characteristics (Smith, in litt.
(Reptile and amphibian populations at these sites; are; much smaller in size than one would expect based on the habitat characteristics (Smith, in litt) (one; would expect; )
Reptile and amphibian populations at these sites one
are would expect
much smaller in size than one would expect based on the habitat characteristics (Smith, in litt