1959, last paragraph , On the Origin of Species Graham Bell (1982) first used the term "tangled bank" to describe the conditions of heavy competition and many different microsites Spatial variation in the environment provides selection for variable offspring Intra-specific competition causes selection for sexually-produced offspring Red Queen
(tangled bank; provides; selection for variable offspring Intra-specific competition)
tangled bank
provides
selection for variable offspring Intra-specific competition