Indeed, since depletion of fat reserves, predation, and fungal disease are the principal causes of winter death in this species, [ 14 ] it would seem that mortality should be more likely to increase rather than decrease in abnormally warm, wet conditions.
(depletion of fat reserves, predation, and fungal disease; are; the principal causes of winter death in this species) (that mortality; to increase; )
depletion of fat reserves, predation, and fungal disease that mortality
are to increase
the principal causes of winter death in this species