What he meant was that for any pair of characteristics there is only one gene in a gamete even though there are two genes in ordinary cells.
(he; meant; ) (What he meant; was; that for any pair of characteristics there is only one gene in a gamete even though there are two genes in ordinary cells)
he What he meant
meant was
that for any pair of characteristics there is only one gene in a gamete even though there are two genes in ordinary cells