In terrestrial ecosystems, lack of nutrients, waterlogged conditions on the tundra, and the lack of water in the Arctic desert areas also limit productivity.
(lack of nutrients; waterlogged; conditions on the tundra; L:In terrestrial ecosystems) (the lack of water in the Arctic desert areas; also limit; productivity)
lack of nutrients the lack of water in the Arctic desert areas
waterlogged also limit
conditions on the tundra L:In terrestrial ecosystems
productivity