however, at most sites the extent of the contiguous habitat is small, both increasing the vulnerability of large populations due to their density, and of small populations due to the potential of extirpation by removal of only a few plants (FWS 1986).
(the extent of the contiguous habitat; is; small; L:at most sites) (both; increasing; the vulnerability of large populations; due to their density)
the extent of the contiguous habitat both
is increasing
small L:at most sites
the vulnerability of large populations due to their density