This effect, combined with more efficient killing of insect pests, may result in less food for their predators and other animals on the food chain.
(This effect; combined; with more efficient killing of insect pests) (This effect, combined with more efficient killing of insect pests; may result; in less food)
This effect This effect, combined with more efficient killing of insect pests
combined may result
with more efficient killing of insect pests
in less food