In brief, Morgan had discovered that eye color in Drosophila expressed a sex-linked trait.
(Morgan; had discovered; that eye color in Drosophila expressed a sex-linked trait; T:In brief) Context(Morgan had discovered,List([10, 31))):(eye color in Drosophila; expressed; a sex-linked trait)
Morgan had discovered,List([10, 31))
Morgan eye color in Drosophila
had discovered expressed
that eye color in Drosophila expressed a sex-linked trait T:In brief
a sex-linked trait