While the populations of most large predators are declining, coyote populations are thriving and expanding their ranges.
(the populations of most large predators; are declining; ) (coyote populations; are thriving; T:While the populations of most large predators are declining) (coyote populations; expanding; their ranges)
the populations of most large predators coyote populations coyote populations
are declining are thriving expanding
T:While the populations of most large predators are declining
their ranges