Assertion: squid's ink sac | /r/AtLocation | saundersites
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text: The earliest ink sacs appear in the fossil record in the Carboniferous period around 330 million years ago in cephalopods such as Donovaniconus, Gordoniconus and Saundersites which show a mix of features from older and more modern groups and are placed in their own order, Donovaniconida (Doguzhaeva 2012).