Assertion: quetzal | /r/CapableOf | colonize the central american isthmus
Facets:
value: in tandem
type: MANNER
value: during the pliocene epoch
type: TEMPORAL
Source sentences:
text: It has been theorized that the quetzals and Lauraceae (whose fruits they depend on and whose seeds they disperse) colonized the Central American Isthmus in tandem during the Pliocene Epoch (Raven and Axelrod, 1974).