Assertion: lignin's effect | /r/HasProperty | observed for hct
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text: On the other hand, the strongest effects on H/G/S lignin composition have been observed for trees downregulated in C3’H and HCT, which deposit lignin with large increases in H unit content (Coleman et al., 2008a; Ralph et al., 2012; Vanholme et al., 2013a), whereas trees that overexpress F5H produce lignin strongly enriched in S units (Franke et al., 2000; Li L. et al., 2003; Stewart et al., 2009), and trees that are downregulated in COMT have dramatically reduced S unit biosynthesis (Van Doorsselaere et al., 1995; Lapierre et al., 1999).