Assertion: giardia's nucleus | /r/HasProperty | et al.
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text: Interestingly, none of the Enteromonads were basal to the Diplomonads (as suggested by Brugerolle, 1975; Siddall et al., 1992) but were instead polyphyletic within the lineage including Spironucleus, suggesting either multiple origins of diplokarya or multiple instances of secondary loss of the duplicated nucleus (Kolisko et al., 2008).