  The spatial segregation of species is fundamental to ecosystem formation and
stability. Behavioural strategies may determine where species are located and
how their interactions change the local environment arrangement. In response to
stimuli in the environment, individuals may move in a specific direction
instead of walking randomly. This behaviour can be innate or learned from
experience, and allow the individuals to conquer or the maintain territory,
foraging or taking refuge. We study a generalisation of the spatial
rock-paper-scissors model where individuals of one out of the species may
perform directional movement tactics. Running a series of stochastic
simulations, we investigate the effects of the behavioural tactics on the
spatial pattern formation and the maintenance of the species diversity. We also
explore a more realistic scenario, where not all individuals are conditioned to
perform the behavioural strategy or have different levels of neighbourhood
perception. Our outcomes show that self-preservation behaviour is more
profitable in terms of territorial dominance, with the best result being
achieved when all individuals are conditioned and have a long-range vicinity
perception. On the other hand, invading is more advantageous if part of
individuals is conditioned and if they have short-range neighbourhood
perception. Finally, our findings reveal that the self-defence strategy is the
least jeopardising to biodiversity which can help biologists to understand
population dynamics in a setting where individuals may move strategically.
