Neornithes

{{evolution}}
'''Neornithes''' Gadow 1893<ref>Gadow, H. 1893. Vogel II: Systematicher Theil. In: Bronn, H. G. (ed.), ''Klassen und Ordnungen dees Thier-Reichs'', vol. 6(4). C. F.Winter, Leipzig.</ref> (=[[Aves]] ''sensu'' Gauthier 1986),<ref>Gauthier, J. 1986.  Saurischian monophyly and the origin of birds. ''Memoires of the California Academy of Sciences'' 8: 1-55.</ref> meaning "new birds". This term has fallen out of use for many people as some argue that all modern birds are in aves. Neornithine dates back to an outdated notion that creatures like archeopteryx and the enantiornithines are part of the aves group. 

Neornithes are the most recent common ancestral species of all extant birds and all of said species' descendants. Neornithine anatomy has been reviewed most recently by Hope (2002),<ref>Hope, S. 2002. The Mesozoic radiation of Neornithes. In: Chiappe, L. & Witmer, L. (eds.), ''Mesozoic Birds: Above the Heads of Dinosaurs'', 339-388.</ref> who presented a succinct but precise account of the morphological changes associated with the derivation of crown clade birds. Synapomorphies (common features) of Neornithes include: 
#Premaxilla/maxilla fused
#Maxilla greatly reduced and primarily restricted to the palatal region
#Mandibular symphysis fused
#Dentary and surangular fused
#Teeth absent
#Humeral facet of coracoid does not extend laterally beyond the scapular facet of the coracoid
#Scapular humeral facet is oriented laterally or craniolaterally
#Humeral facet of coracoid separated slightly or completely from the scapular facet 
#Humeral head large
#Sulcus for transverse humeral ligament distinct
#Deltopectoral crest of humerus inflected cranially
#Pneumotricipital fossa pierced by a pneumatic foramen

The origin of Neornithes is one of the most contested issues in paleornithology, with two widely disparate models advanced to account for the evolution of this group (reviewed in [[Neornithine 'Big Bang']]). Classically, Neornithes has been considered to cleave into two divergent lineages, the Paleognathae and the Neognathae, first delineated by Huxley on the basis of palatal configuration.

Neornithes are divided into the following extant orders (based on Jarvis, E.D. et al. (2014)<ref name=Jarvis2014>{{cite journal | last1 = Jarvis | first1 = E.D. |display-authors=etal  | year = 2014 | title = Whole-genome analyses resolve early branches in the tree of life of modern birds | url = http://www.sciencemag.org/content/346/6215/1320.abstract | journal = Science | volume = 346 | issue = 6215| pages = 1320–1331 | doi=10.1126/science.1253451 | pmid=25504713 | pmc=4405904}}</ref>):
*Struthioniformes
*Rheiformes
*Casuariiformes
*Apterygiformes
*Tinamiformes
*Anseriformes
*Galliformes
*Phoenicopteriformes
*Podicipediformes
*Mesitornithiformes
*Pterocliformes
*Columbiformes
*Musophagiformes
*Otidiformes
*Cuculiformes
*Caprimulgiformes
*Steatornithiformes
*Nyctibiiformes
*Podargiformes
*Aegotheliformes
*Apodiformes
*Opisthocomiformes
*Gruiformes
*Charadriiformes
*Eurypgiformes
*Phaethoniformes
*Gaviiformes
*Sphenisciformes
*Procellariiformes
*Ciconiiformes
*Suliformes
*Pelecaniformes
*Cathartiformes
*Accipitriformes
*Strigiformes
*Coliiformes
*Leptosomiformes
*Trogoniformes
*Bucerotiformes
*Coraciiformes
*Piciformes
*Cariamiformes
*Falconiformes
*Psittaciformes
*Passeriformes

The constitution and phylogenetic placement of these taxa remains hotly contested in systematic ornithology, but some general conclusions can be made:
*The ratite assemblage (Tinamiformes + Struthioniformes + Casuariiformes + Rheiformes) may be holophyletic.<ref name="sibley1990">Sibley, C. G. & Ahlquist, J. E. 1990. ''Phylogeny and Classification of Birds''. Yale University Press, New Haven.</ref>
*Charadriiform-like taxa (e.g., ''Graculavus'') are the wellspring of the principal neornithine adaptive radiation.<ref>Martin, L. D. 1983. The origin and early radiation of birds. In: Brush, A. H. & Clark, Jr., G. A. (eds.), ''Perspectives in Ornithology'', 291-338.</ref><ref>Olson, S. L. 1985. The fossil record of birds. In: Farner, D. S., King, J. R., and Parkes, K. C. (eds.), ''Avian Biology'' Volume VIII, 79-252.</ref><ref>Feduccia, A. 1995. Explosive evolution in Tertiary birds and mammals. ''Science'' 267: 637-638.</ref><ref name="feduccia1996">Feduccia, A. 1996. ''The Origin and Evolution of Birds, First Edition''. Yale University Press, New Haven.</ref><ref>Feduccia, A. 1999. ''The Origin and Evolution of Birds, Second Edition.'' Yale University Press, New Haven.</ref><ref>Feduccia, A. 2003. 'Big bang' for Tertiary birds? ''Trends in Ecology and Evolution'' 18(4): 172-176.</ref>
*Ciconiiformes as classically defined is polyphyletic.<ref name="feduccia1996"/> 
*Strigiformes and Caprimulgiformes are closely allied.<ref name="sibley1990"/>
*Falconiformes as classically defined is polyphyletic.<ref name="sibley1990"/>
*Gaviiformes and Podicipediformes are not closely related.<ref name="sibley1990"/><ref>Storer, R. W. 1960. Evolution in the diving birds. ''Proceedings of the Twelfth International Ornithological Congress'', 694-707.</ref>
*Passeriformes is not a basal radiation within Neornithes.<ref>van Tuinen, M., Sibley, C. G., & Hedges, S. B. 2000. The early history of modern birds inferred from DNA sequences of nuclear and mitochondrial ribosomal genes. ''Molecular and Biological Evolution'' 17(3): 451-457.</ref>

Further consensus is not readily possible and indeed even these conclusions presented here are contentious.  Higher level systematics of the neornithine assemblage remain difficult to interpret despite a century of work on these birds, as noted by Streseman in the 1950s. Thus, neornithine evolution remains a fertile field for students of avian phylogeny.

==References==
{{reflist|2|90%}}

[[Category:EvoWiki ports]]
[[Category:Evolution]]
[[Category:Biology]]

Categories: EvoWiki ports, Evolution, Biology