At its center, the Morosini Fountain dates from Venetian times, though the somber lion statues are three hundred years older.
The Morosini Fountain and the lion statues are both from Venetian times. 
mnli hypothesis: The Morosini Fountain and the lion statues are both from Venetian times.  premise: At its center, the Morosini Fountain dates from Venetian times, though the somber lion statues are three hundred years older.