Please read the following table in Markdown format and then answer the question according to the table. Table cells in one row are seperated by '|', and different rows are seperated by '
'.

Table:
| Species | Country of origin | Total released | First release | Last release | Areas of release | Areas established | Pasture type |
| --- | --- | --- | --- | --- | --- | --- | --- |
| Bubas bison | France, Spain | 1,613 | April 1983 | 1996 (not by CSIRO) | WA | NSW, SA, WA | Winter rainfall |
| Copris elphenor Klug | South Africa | 2,287 | January 1977 | May 1983 | QLD | QLD | Summer rainfall |
| Copris hispanus Linnaeus | Spain | 294 | October 1983 | June 1994 | WA | WA | Winter rainfall |
| Euoniticellus africanus Harold | South Africa | 49,009 | October 1971 | February 1984 | NSW, QLD, SA, Tas, Vic, WA | NSW, QLD | Summer rainfall |
| Euoniticellus fulvus Goeze | France, Turkey | 76,944 | March 1978 | February 1983 | NSW, SA, Tas, Vic, WA | NSW, SA, Tas, Vic, WA | Winter rainfall |
| Euoniticellus intermedius Reiche | South Africa | 248,637 | November 1971 | February 1984 | ACT, NSW, NT, QLD, SA, Vic, WA | NSW, NT, QLD, SA, Vic, WA | Summer rainfall |
| Euoniticellus pallipes Fabricius | Iran, Turkey | 46,642 | March 1977 | September 1982 | NSW, SA, WA | NSW, SA, Vic, WA | Winter rainfall |
| Geotrupes spiniger Marsham | France | 12,082 | April 1979 | December 1983 | ACT, NSW, Tas, Vic | ACT, NSW, SA, Tas, Vic | Winter rainfall |
| Liatongus militaris Castelanu | South Africa (via Hawaii) | 70,450 | January 1968 | November 1979 | NSW, NT, QLD, WA | NSW, NT, QLD | Summer rainfall |
| Onitis alexis Fabricius | South Africa | 186,441 | August 1972 | February 1984 | NSW, NT, QLD, WA | NSW, NT, QLD, SA, Vic, WA | Rainfall, summer rainfall, winter rainfall |
| Onitis aygalus Fabricius | South Africa | 18,682 | January 1977 | January 1982 | NSW, SA, WA | NSW, SA, Vic, WA | Winter rainfall |
| Onitis caffer Boheman | South Africa | 8,738 | October 1979 | April 1984 | WA | QLD, NSW, SA | Summer rainfall, winter rainfall |
| Onitis pecuarius Lansberge | South Africa | 11,395 | November 1976 | May 1979 | NSW, QLD | NSW, QLD | Summer rainfall |
| Onitis vanderkelleni Lansberge | Kenya, Rwanda, Zaire | 10,852 | October 1974 | February 1982 | NSW, QLD | QLD | Summer rainfall |
| Onitis viridulus Bohemann | South Africa | 8,008 | September 1976 | July 1980 | NSW, NT, QLD | NSW, NT, QLD, WA | Summer rainfall |
| Onthophagus binodis Thunberg | South Africa | 173,018 | October 1971 | February 1982 | NSW, QLD, SA, Tas, Vic, WA, Norfolk Islands | NSW, QLD, SA, Tas, Vic, WA, Norfolk Islands | Winter rainfall |
| Onthophagus gazella Fabricius | South Africa | 420,415 | February 1968 | February 1984 | ACT, NSW, NT, QLD, SA, Tas, Vic, WA, Norfolk Islands | ACT, NSW, NT, QLD, SA, WA, Norfolk Islands | Summer rainfall |
| Onthophagus nigiventris d'Orbigny | East Africa | 29,960 | May 1975 | March 1983 | NSW | NSW | Summer rainfall |
| Onthophagus obliquus | Nigeria, Senegal, Zaire | 9,300 | January 1976 | November 1977 | QLD, NT | QLD | Summer rainfall |
| Onthophagus sagittarius | Sri Lanka (via Hawaii) | 9,075 | January 1968 | March 1977 | NSW, NT, QLD, WA | NSW, NT | Summer rainfall |
| Onthophagus taurus Schreber | Spain, Greece, Italy, Turkey | 164,499 | February 1975 | January 1984 | NSW, SA, Tas, Vic, WA | NSW, SA, Tas, Vic, WA | Winter rainfall |
| Sisyphus rubrus Paschalidis | South Africa | 85,933 | March 1973 | February 1980 | NSW, QLD, WA | NSW, QLD | Summer rainfall |
| Sisyphus spinipes | South Africa | 36,125 | March 1972 | December 1978 | NSW, NT, QLD, WA | NSW, QLD | Summer rainfall |
|  | Total: | 1,680,399 |  |  |  |  |  |
Question:
which beetle had the largest amount released?
Answer:
Onthophagus gazella Fabricius