In this paper, he analyzed skeletal remains of Indian and African elephants, as well as mammoth fossils, and a fossil skeleton known at that time as the 'Ohio animal'.
在本文中，他分析了印度和非洲大象的骨骼遗骸与猛犸象化石，以及当时被称为“俄亥俄动物”的化石骨架。