Δ L k = L k − L k o {\displaystyle \Delta {Lk=Lk-Lk_{o}}} If the DNA is negatively supercoiled ΔLk < 0.
Δ L k = L k − L k o {\displaystyle \Delta {Lk=Lk-Lk_{o}}} Eğer DNA negatif süpersarımlı ise ΔLk < 0.