Genetic differences between isolates in the South American and Australian populations indicate a long period of geographical separation, and the authors suggest that they "later might be regarded as independent taxa".
Perbedaan genetik antara populasi terisoasi di Amerika Selatan dan Australia mengindikasikan periode panjang dari pemisahan geografi, dan para pengarang bersugesti bahwa mereka "kemudian dianggap sebagai taxa independen".