He helped create the Wellcome Trust Centre for Cell Biology, also in Edinburgh, and served as its director from 1999 until 2011, when he was succeeded by David Tollervey.
Ia membantu pendirian Pusat Biologi Sel Wellcome Trust, juga di Edinburgh, dan mengabdi sebagai direkturnya dari 1999 hingga 2011, ketika ia digantikan oleh David Tollervey.