He helped to bridge the divide between the foundations of microevolution developed by the population geneticists and the patterns of macroevolution observed by field biologists, with his 1937 book Genetics and the Origin of Species.
Ia membantu menjembatani perpecahan antara fonasi mikroevolusi yang dikembangkan oleh para pakar genetik populasi dan susunan makroevolusi yang diamati oleh pakar biologi lapangan, dengan buku tahun 1937 buatannya Genetics and the Origin of Species.