Abbott Handerson Thayer's 1909 book Concealing-Coloration in the Animal Kingdom, completed by his son Gerald H. Thayer, argued correctly for the widespread use of crypsis among animals, and in particular described and explained countershading for the first time.
O libro de 1909 de Abbott Handerson Thayer Concealing-Coloration in the Animal Kingdom, completado polo seu fillo Gerald H. Thayer, argumentaba correctamente o uso estendido da cripse entre os animais, e en particular describía e explicaba o contrasombreado por primeira vez.