This hypothesis was alluded to by Mayr in the closing paragraph of his 1954 paper: Rapidly evolving peripherally isolated populations may be the place of origin of many evolutionary novelties.
Mayr aludiu a esta hipótese no parágrafo final do seu artigo de 1954: As poboacións periféricas illadas en rápida evolución poden ser o lugar de orixe de moitas novidades evolutivas.