Question: After the controversies of the 2015 World Conference of Science Journalists in Seoul, David Colquhoun was critical of a British biochemist who was awarded the Nobel Prize in Physiology or Medicine in what year?

Context articles: List of Nobel laureates in Physiology or Medicine
Sir Tim Hunt controversy at WCSJ
Nobel Prize in Physiology or Medicine
Nobel Prize in Literature
Nobel Committee for Physiology or Medicine
List of female Nobel laureates
Ernst Boris Chain
David Colquhoun
Tim Hunt
Frederick Sanger

The Nobel Prize in Physiology or Medicine (Swedish: "Nobelpriset i fysiologi eller medicin" ) is awarded annually by the Swedish Karolinska Institute to scientists and doctors in the various fields of physiology or medicine.
 It is one of the five Nobel Prizes established by the 1895 will of Alfred Nobel (who died in 1896), awarded for outstanding contributions in chemistry, physics, literature, peace, and physiology or medicine.
 As dictated by Nobel's will, the award is administered by the Nobel Foundation and awarded by a committee that consists of five members and an executive secretary elected by the Karolinska Institute.
 While commonly referred to as the Nobel Prize in Medicine, Nobel specifically stated that the prize be awarded for "physiology or medicine" in his will.
 Because of this, the prize can be awarded in a broader range of fields.
 The first Nobel Prize in Physiology or Medicine was awarded in 1901 to Emil Adolf von Behring, of Germany.
 Each recipient receives a medal, a diploma and a monetary award that has varied throughout the years.
 In 1901, von Behring received 150,782 SEK, which is equal to 7,731,004 SEK in December 2008.
 In 2013, the prize was awarded to James E. Rothman, Randy W. Schekman and Thomas C. Südhof; they were recognised "after discovering how cells precisely transport material".
 The award is presented in Stockholm at an annual ceremony on December 10, the anniversary of Nobel's death.
In 2015, British scientist and Nobel Prize winner Sir Timothy Hunt was involved in a highly publicised controversy at the World Conference of Science Journalists (WCSJ) in Seoul.
The Nobel Prize in Physiology or Medicine (Swedish: "Nobelpriset i fysiologi eller medicin" ), administered by the Nobel Foundation, is awarded once a year for outstanding discoveries in the fields of life sciences and medicine.
 It is one of five Nobel Prizes established in 1895 by Swedish chemist Alfred Nobel, the inventor of dynamite, in his will.
 Nobel was personally interested in experimental physiology and wanted to establish a prize for progress through scientific discoveries in laboratories.
 The Nobel Prize is presented to the recipient(s) at an annual ceremony on 10 December, the anniversary of Nobel's death, along with a diploma and a certificate for the monetary award.
 The front side of the medal provides the same profile of Alfred Nobel as depicted on the medals for Physics, Chemistry, and Literature; its reverse side is unique to this medal.
Since 1901, the Nobel Prize in Literature (Swedish: "Nobelpriset i litteratur" ) has been awarded annually to an author from any country who has, in the words of the will of Alfred Nobel, produced "in the field of literature the most outstanding work in an ideal direction" (original Swedish: "den som inom litteraturen har producerat det mest framstående verket i en idealisk riktning").
 Though individual works are sometimes cited as being particularly noteworthy, here "work" refers to an author's work as a whole.
 The Swedish Academy decides who, if anyone, will receive the prize in any given year.
 The academy announces the name of the chosen laureate in early October.
 It is one of the five Nobel Prizes established by the will of Alfred Nobel in 1895; the others are the Nobel Prize in Chemistry, Nobel Prize in Physics, Nobel Peace Prize, and Nobel Prize in Physiology or Medicine.
The Nobel Committee for Physiology or Medicine is the Nobel Committee responsible for proposing laureates for the Nobel Prize for Physiology or Medicine.
 The Nobel Committee for Physiology or Medicine is appointed by the Nobel Assembly at Karolinska Institutet, a body of 50 members at Karolinska Institutet.
 It usually consists of Swedish professors of physiology or medical subjects active at Karolinska Institutet, although the Assembly in principle could appoint anyone to the Committee.
 Other than the five ordinary members, ten associated members are appointed each year, for that year only.
The Nobel Prizes are awarded annually by the Royal Swedish Academy of Sciences, the Swedish Academy, the Karolinska Institute, and the Norwegian Nobel Committee to individuals who make outstanding contributions in the fields of Chemistry, Physics, Literature, Peace, Physiology or Medicine and Economics.
 All but the economics prize were established by the 1895 will of Alfred Nobel, which dictates that the awards should be administered by the Nobel Foundation.
 The Nobel prize in Economics, or The Sveriges Riksbank Prize in Economic Sciences in Memory of Alfred Nobel, was established in 1968 by the Sveriges Riksbank, the central bank of Sweden, for outstanding contributions in the field of Economics.
 Each prize is awarded by a separate committee; the Royal Swedish Academy of Sciences awards the Prizes in Physics, Chemistry, and Economics, the Swedish Academy awards the Prize in Literature, the Karolinska Institute awards the Prize in Physiology or Medicine, and the Norwegian Nobel Committee awards the Prize in Peace.
 Each recipient receives a medal, a diploma and a cash prize that has varied throughout the years.
Sir Ernst Boris Chain, FRS (19 June 1906 – 12 August 1979) was a German-born British biochemist, and a 1945 co-recipient of the Nobel Prize for Physiology or Medicine for his work on penicillin.
David Colquhoun, {'1': ", '2': ", '3': ", '4': "} (born 19 July 1936) is a British pharmacologist at University College London (UCL).
 He has contributed to the general theory of receptor and synaptic mechanisms, and in particular the theory and practice of single ion channel function.
 He previously held the A.J. Clark chair of Pharmacology at UCL, and was the Hon. Director of the Wellcome Laboratory for Molecular Pharmacology.
 He was elected a Fellow of the Royal Society (FRS) in 1985 and an honorary fellow of UCL in 2004.
 Colquhoun runs the website "DC's Improbable Science", which is critical of pseudoscience, particularly alternative medicine, and managerialism.
 He was critical of Tim Hunt following the controversies of the 2015 World Conference of Science Journalists in Seoul.
Sir Richard Timothy Hunt, (born 19 February 1943) is a British biochemist and molecular physiologist.
 He was awarded the 2001 Nobel Prize in Physiology or Medicine with Paul Nurse and Leland H. Hartwell for their discoveries of protein molecules that control the division of cells.
 In particular, Hunt discovered what he called cyclin: a protein in fertilised sea urchin eggs which cyclically aggregates and is depleted during cell division cycles.
Frederick Sanger ( ; 13 August 1918 – 19 November 2013) was a British biochemist who twice won the Nobel Prize in Chemistry, one of only two people to have done so in the same category (the other is John Bardeen in physics), the fourth person overall with , and the third person overall with two Nobel Prizes in the sciences.
 In 1958, he was awarded a Nobel Prize in chemistry "for his work on the structure of proteins, especially that of insulin".
 In 1980, Walter Gilbert and Sanger shared half of the chemistry prize "for their contributions concerning the determination of base sequences in nucleic acids".
 The other half was awarded to Paul Berg "for his fundamental studies of the biochemistry of nucleic acids, with particular regard to recombinant DNA".