Question: What animal has migrated from Souteast Asia and caused the extinction of The Kauaʻ i ʻ ōʻ ō in Hawaii?

Context articles: Kauaʻi ʻōʻō
Mohoidae
ʻŌʻōkala, Hawaii
Lāna'i hookbill
David Boynton
Hawaiʻi ʻōʻō
Puʻu ʻŌʻō
Moho (genus)
Polynesian rat
Freycinetia arborea

The Kauaʻ i ʻ ōʻ ō or ʻ ōʻ ōʻ āʻ ā (Moho braccatus) is a member of the extinct genus of the ʻ ōʻ ōs ("Moho") within the extinct family Mohoidae from the islands of Hawai'i.
 It was previously regarded as member of the Australo-Pacific honeyeaters ("Meliphagidae").
 This bird was endemic to the island of Kauaʻ i.
 It was common in the subtropical forests of the island until the early twentieth century, when its decline began.
 Its song was last heard in 1987 and it is now probably extinct.
 The causes of its extinction include the introduction of the Polynesian rat, domestic pig, and mosquitoes carrying avian disease (avian malaria and avian pox), as well as habitat destruction.
Mohoidae is a family of Hawaiian species of recently extinct, nectarivorous songbirds in the genera "Moho" (ʻ ōʻ ō) and "Chaetoptila" (kioea).
 These now extinct birds form their own family, representing the only complete extinction of an entire avian family in modern times, when the disputed family Turnagridae is disregarded for being invalid.
ʻ Ōʻ ōkala (also spelled Ookala) is an unincorporated community on the island of Hawaii in Hawaii County, Hawaii, United States.
 It lies along Hawaii Route 19 north of Hilo, the county seat of Hawaii County.
 Its elevation is 371 feet (113 m), and it is located at (20.0175000, -155.2872222).
 Because the community has borne multiple names, the Board on Geographic Names officially designated it "ʻ Ōʻ ōkala" in 2000.
 Although it is unincorporated, it has a post office, with the ZIP code of 96774.
The Lānaʻ i hookbill ("Dysmorodrepanis munroi") is an extinct species of Hawaiian honeycreeper.
 It was endemic to the island of Lānaʻ i in Hawaiʻ i, and was last seen in the southwestern part of the island.
 George C. Munro collected the only known specimen of this species in 1913, which is housed in the Bernice P. Bishop Museum in Honolulu, and saw the species only twice more, once in 1916 and for a final time in 1918.
 No other sightings have been reported.
 They inhabited montane dry forests dominated by ʻ akoko ("Euphorbia" species) and ōpuhe ("Urera glabra").
 The Lānaʻ i hookbill was monotypic within the genus Dysmorodrepanis and had no known subspecies.
 Its closest relative is believed to be the ʻ ōʻ ū, and some early authors suggested that the Lānaʻ i hookbill was merely a deformed ʻ ōʻ ū.
 The Lānaʻ i hookbill was a plump, medium-sized bird with greenish olive upperparts and pale whitish yellow underparts.
 It also had a yellow or white superciliary line and a white chin and throat.
 The wings also had a distinctive and conspicuous white wing patch.
 The hookbill's distinguishing characteristic was its heavy, parrotlike bill, which had the mandibles hooking sharply towards each other, leaving a gap between them when the beak was closed.
David Boynton (1945–2007) was a leading expert on the natural history of the Hawaiian island of Kauai, especially on the Koke'e Forest and the Alakai Swamp and its wildlife.
 He was called "a voice for the Hawaiian wilderness," a "Guardian of the Koke'e Forest," and as an educator, "the window through which thousands of Hawai'i students learned about Hawaiian birds, plants, marine creatures, climate and much more."
 Boynton photographed a bird now believed extinct, the ʻ Ōʻ ōʻ āʻ ā ("Moho braccatus").
 He recorded the mating call of the single male, whose mate presumably did not survive Hurricane Iwa at the end of 1982.
 The bird, probably the last of its species, was tending an empty nest.
The Hawaiʻ i ʻ ōʻ ō ("Moho nobilis") is a member of the extinct genus of the ʻ ōʻ ōs ("Moho") within the extinct family Mohoidae.
 It was previously regarded as member of the Australo-Pacific honeyeaters ("Meliphagidae").
Puʻ u ʻ Ōʻ ō (often written Puu Oo, ] ) is a volcanic cone in the eastern rift zone of the Kīlauea volcano of the Hawaiian Islands.
 Puʻ u ʻ Ōʻ ō has been erupting continuously since January 3, 1983, making it the longest-lived rift-zone eruption of the last two centuries.
Moho is a genus of extinct birds in the Hawaiian bird family, Mohoidae, that were endemic to the Hawaiian Islands.
 Members of the genus are known as ʻ ōʻ ō in the Hawaiian language.
 Their plumage was generally striking glossy black; some species had yellowish axillary tufts and other black outer feathers. Most of these species became extinct by habitat loss and by extensive hunting because their plumage were used for the creation of precious "ʻ aʻ ahu aliʻ i" (robes) and "ʻ ahu ʻ ula" (capes) for "aliʻ i" (Hawaiian nobility).
 The Kauaʻ i ʻ ōʻ ō was the last species of this genus to become extinct, probably a victim of avian malaria.
The Polynesian rat, or Pacific rat ("Rattus exulans"), known to the Māori as kiore, is the third most widespread species of rat in the world behind the brown rat and black rat.
 The Polynesian rat originated in Southeast Asia, and like its relatives, has become widespread, migrating to most Polynesian islands, including New Zealand, Easter Island, and Hawaii.
 It shares high adaptability with other rat species extending to many environments, from grasslands to forests.
 It is also closely associated with humans, who provide easy access to food.
 It has become a major pest in most areas of its distribution.
Freycinetia arborea, ʻ Ieʻ ie, is a densely branched, brittle, woody climber in the family Pandanaceae, endemic to the Pacific Islands. "
ʻ Ieʻ ie" is found in moist forest on the Hawaiian, Marquesas, Austral, Society, and Cook Islands.
 It grows into the forest canopy, attaching itself to a host tree using aerial roots.
 It may also grow as a sprawling tangle on the forest floor.
 The shiny green leaves have pointed ends and are spiny on the lower side of the midrib and along the edges.
 Leaves measure 40 - long and 1 - wide, and are spirally arranged around the ends of branches.
 Flowers form on spike-like inflorescences at the end of branches, and are either staminate or pistillate.
 Staminate spikes are yellowish-white and up to 10 cm in length.
 Pistillate spikes are 3 - but elongate to 7.5 - once fruit are produced.
 Three to four spikes are surrounded by orange-salmon bracts. Fruit is 1 cm long and contains many 1.5 mm seeds.
 The bracts and fruit of the "ʻ ieʻ ie" were a favorite food of the "ʻ ōʻ ū" ("Psittirostra psittacea"), an extinct Hawaiian honeycreeper that was formerly a principal seed dispersal vector for plants with small seeded, fleshy fruits in low elevation forests.
 It is also a favored food of the "ʻ alalā" ("Corvus hawaiiensis"), which is currently extinct in the wild.