Question: What is the capital city of this Central American isthmus, to which the species of crab Aratus pisonii is native, bordering with Costa Rica?

Context articles: Isthmo-Colombian Area
Aratus pisonii
Mexican immigration to Costa Rica
Wildlife of Costa Rica
Grey-crowned Central American squirrel monkey
Geography of Costa Rica
Central America Volcanic Arc
Nicaragua
List of Costa Rican monkey species
Costa Rica national cricket team

The Isthmo-Colombian Area is defined as a cultural area encompassing those territories occupied predominantly by speakers of the Chibchan languages at the time of European contact.
 It includes portions of the Central American isthmus like eastern El Salvador, eastern Honduras, Caribbean Nicaragua, Costa Rica, Panama, and northern Colombia.
Aratus pisonii, commonly known as the mangrove tree crab, is a species of crab which lives in mangrove trees in tropical and subtropical parts of the Americas, from Florida to Brazil on the Atlantic coast, and from Nicaragua to Peru on the Pacific coast.
 It feeds mostly on the leaves of the mangroves, but is an omnivore, and prefers animal matter when possible.
 "A. pisonii" is the only species in the monotypic genus Aratus.
 The specific epithet "pisonii" commemorates the Dutch naturalist Willem Piso who travelled in Brazil in 1638 with Georg Marggraf.
Mexican immigration to Costa Rica comprises people who emigrated from Mexico to Costa Rica and their descendants.
 Costa Rica has the most Mexicans living in Central America, aside from Guatemala.
 The waves of migration from Mexico to Costa Rica started from 1970s, as they were attracted by a stable democracy, a mild climate and political stability that characterizes this country south of Central.
 The Mexican population in Costa Rica reaches more than 5,000 people who exercise as professionals, doctors, secretaries, among other services.
 Costa Rica is the seventh destination for Mexican immigrants in the world, and first in Hispanic America.
 Unlike other destinations, most Mexicans who immigrate to Costa Rica are mainly from Nuevo León, Tamaulipas, Chihuahua, Baja California and Mexico City.
 Despite having more cultural elements similar to the northern countries of Central America (Guatemala, El Salvador and Honduras), Mexicans in Costa Rica are quickly assimilated by the Costa Rican population.
 From 2010, a new wave of Mexican immigrants began coming to Costa Rica, and these are mainly young professionals, who find no opportunities in their country, migrate to the Central American country.
 Many others are running away from the waves of violence following the war against drug trafficking in Mexico.
The Wildlife of Costa Rica comprises all naturally occurring animals, fungi and plants that reside in this Central American country.
 Costa Rica supports an enormous variety of wildlife, due in large part to its geographic position between the North and South American continents, its neotropical climate, and its wide variety of habitats.
 Costa Rica is home to more than 500,000 species, which represents nearly 4% of the total species estimated worldwide, making Costa Rica one of the 20 countries with the highest biodiversity in the world.
 Of these 500,000 species, a little more than 300,000 are insects.
The grey-crowned Central American squirrel monkey ("Saimiri oerstedii citrinellus") is a subspecies of the Central American squirrel monkey.
 Its range is restricted to the Pacific coast of central Costa Rica.
 The northern end of its range is the Rio Tulin and the southern end of its range is the Rio Grande de Terraba.
 South of the Rio Grande de Terraba, it is replaced by the black-crowned Central American squirrel monkey, "S. oerstedii oerstedii".
 Populations are very fragmented, and the subspecies does not occur in all locations within its general range.
 It is the subspecies of Central American squirrel monkey seen in Manuel Antonio National Park in Costa Rica.
Costa Rica is located on the Central American Isthmus, surrounding the point 10° north of the equator and 84° west of the prime meridian.
 It borders both the Caribbean Sea (to the east) and the North Pacific Ocean (to the west), with a total of 1,290 km of coastline (212 km on the Caribbean coast and 1,016 km on the Pacific).
The Central American Volcanic Arc (often abbreviated to CAVA) is a chain of volcanoes which extends parallel to the Pacific coast line of the Central American Isthmus, from Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, and down to northern Panama.
 This volcanic arc, which has a length of 1500 km , is formed by an active subduction zone along the western boundary of the Caribbean Plate.
Nicaragua ( ; ] ), officially the Republic of Nicaragua (Spanish:    ), is the largest country in the Central American isthmus, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west.
 Nicaragua's capital, Managua, is the country's largest city and the third-largest city in Central America.
 The multi-ethnic population of six million includes indigenous peoples, Europeans, Africans, and Asians.
 The main language is Spanish.
 Native tribes on the eastern coast speak their own languages.
Four species of monkey are native to the forests of Costa Rica, the Central American squirrel monkey ("Saimiri oerstedii"), the white-headed capuchin ("Cebus capucinus"), the mantled howler ("Alouatta palliata") and Geoffroy's spider monkey ("Ateles geoffroyi").
 All four species are classified scientifically as New World Monkeys. Two of the species, the Central American squirrel monkey and the white-headed capuchin, belong to the family Cebidae, the family containing the squirrel monkeys and capuchins.
 The other two species belong to the family Atelidae, the family containing the howler monkeys, spider monkeys, woolly monkeys and muriquis. Each of the four species can be seen in national parks within Costa Rica, where viewing them in natural surroundings is a popular tourist attraction.
 The only park in which all four species can be seen is Corcovado National Park, on the Osa Peninsula.
The Costa Rica national cricket team represents the country of Costa Rica in international cricket matches.
 The team is organised by the Costa Rica Cricket Federation, which became an affiliate member of the International Cricket Council in 2002.
 Costa Rica's first recorded international match came in April 2002, when it toured Panama and played against the Panamanian national side.
 The team's first international tournament was the inaugural edition of the Central American Championships, played in Belize in 2006.
 It has since regularly fielded teams in that tournament, and also made its debut in ICC Americas tournaments in 2010, at the 2010 Division Four event in Mexico.