Question: Hugh Pelham's PhD was supervised by which British biochemist and molecular physiologist, who was awarded the 2001 Nobel Prize in Physiology or Medicine?

Context articles: Ernst Boris Chain
Patrick Steptoe
Tim Hunt
Nobel Prize in Literature
List of female Nobel laureates
Hugh Pelham
Nobel Prize in Physiology or Medicine
List of Nobel laureates in Physiology or Medicine
Frederick Sanger
John Macleod (physiologist)

Sir Ernst Boris Chain, FRS (19 June 1906 – 12 August 1979) was a German-born British biochemist, and a 1945 co-recipient of the Nobel Prize for Physiology or Medicine for his work on penicillin.
Patrick Christopher Steptoe CBE FRS (9 June 1913, Oxford, England – 21 March 1988, Canterbury) was a British obstetrician and gynaecologist and a pioneer of fertility treatment.
 Steptoe was responsible with biologist and physiologist Robert Edwards for developing "in vitro" fertilization.
 Louise Joy Brown, the first test-tube baby, was born on 25 July 1978.
 Edwards was awarded the 2010 Nobel Prize in Physiology or Medicine for his work on the development of in vitro fertilization, however because the Nobel Prize is not awarded posthumously, Steptoe was not eligible for consideration.
Sir Richard Timothy Hunt, (born 19 February 1943) is a British biochemist and molecular physiologist.
 He was awarded the 2001 Nobel Prize in Physiology or Medicine with Paul Nurse and Leland H. Hartwell for their discoveries of protein molecules that control the division of cells.
 In particular, Hunt discovered what he called cyclin: a protein in fertilised sea urchin eggs which cyclically aggregates and is depleted during cell division cycles.
Since 1901, the Nobel Prize in Literature (Swedish: "Nobelpriset i litteratur" ) has been awarded annually to an author from any country who has, in the words of the will of Alfred Nobel, produced "in the field of literature the most outstanding work in an ideal direction" (original Swedish: "den som inom litteraturen har producerat det mest framstående verket i en idealisk riktning").
 Though individual works are sometimes cited as being particularly noteworthy, here "work" refers to an author's work as a whole.
 The Swedish Academy decides who, if anyone, will receive the prize in any given year.
 The academy announces the name of the chosen laureate in early October.
 It is one of the five Nobel Prizes established by the will of Alfred Nobel in 1895; the others are the Nobel Prize in Chemistry, Nobel Prize in Physics, Nobel Peace Prize, and Nobel Prize in Physiology or Medicine.
The Nobel Prizes are awarded annually by the Royal Swedish Academy of Sciences, the Swedish Academy, the Karolinska Institute, and the Norwegian Nobel Committee to individuals who make outstanding contributions in the fields of Chemistry, Physics, Literature, Peace, Physiology or Medicine and Economics.
 All but the economics prize were established by the 1895 will of Alfred Nobel, which dictates that the awards should be administered by the Nobel Foundation.
 The Nobel prize in Economics, or The Sveriges Riksbank Prize in Economic Sciences in Memory of Alfred Nobel, was established in 1968 by the Sveriges Riksbank, the central bank of Sweden, for outstanding contributions in the field of Economics.
 Each prize is awarded by a separate committee; the Royal Swedish Academy of Sciences awards the Prizes in Physics, Chemistry, and Economics, the Swedish Academy awards the Prize in Literature, the Karolinska Institute awards the Prize in Physiology or Medicine, and the Norwegian Nobel Committee awards the Prize in Peace.
 Each recipient receives a medal, a diploma and a cash prize that has varied throughout the years.
Pelham was educated at Marlborough College in Marlborough, Wiltshire and Christ's College, Cambridge.
 He graduated with a Master of Arts degree in Natural Sciences followed by a PhD for research on transcription and translation in immature blood cells (Reticulocytes).
 His PhD was supervised by Richard J. Jackson and Tim Hunt, who went on to receive the Nobel Prize in Physiology or Medicine in 2001.
The Nobel Prize in Physiology or Medicine (Swedish: "Nobelpriset i fysiologi eller medicin" ), administered by the Nobel Foundation, is awarded once a year for outstanding discoveries in the fields of life sciences and medicine.
 It is one of five Nobel Prizes established in 1895 by Swedish chemist Alfred Nobel, the inventor of dynamite, in his will.
 Nobel was personally interested in experimental physiology and wanted to establish a prize for progress through scientific discoveries in laboratories.
 The Nobel Prize is presented to the recipient(s) at an annual ceremony on 10 December, the anniversary of Nobel's death, along with a diploma and a certificate for the monetary award.
 The front side of the medal provides the same profile of Alfred Nobel as depicted on the medals for Physics, Chemistry, and Literature; its reverse side is unique to this medal.
The Nobel Prize in Physiology or Medicine (Swedish: "Nobelpriset i fysiologi eller medicin" ) is awarded annually by the Swedish Karolinska Institute to scientists and doctors in the various fields of physiology or medicine.
 It is one of the five Nobel Prizes established by the 1895 will of Alfred Nobel (who died in 1896), awarded for outstanding contributions in chemistry, physics, literature, peace, and physiology or medicine.
 As dictated by Nobel's will, the award is administered by the Nobel Foundation and awarded by a committee that consists of five members and an executive secretary elected by the Karolinska Institute.
 While commonly referred to as the Nobel Prize in Medicine, Nobel specifically stated that the prize be awarded for "physiology or medicine" in his will.
 Because of this, the prize can be awarded in a broader range of fields.
 The first Nobel Prize in Physiology or Medicine was awarded in 1901 to Emil Adolf von Behring, of Germany.
 Each recipient receives a medal, a diploma and a monetary award that has varied throughout the years.
 In 1901, von Behring received 150,782 SEK, which is equal to 7,731,004 SEK in December 2008.
 In 2013, the prize was awarded to James E. Rothman, Randy W. Schekman and Thomas C. Südhof; they were recognised "after discovering how cells precisely transport material".
 The award is presented in Stockholm at an annual ceremony on December 10, the anniversary of Nobel's death.
Frederick Sanger ( ; 13 August 1918 – 19 November 2013) was a British biochemist who twice won the Nobel Prize in Chemistry, one of only two people to have done so in the same category (the other is John Bardeen in physics), the fourth person overall with , and the third person overall with two Nobel Prizes in the sciences.
 In 1958, he was awarded a Nobel Prize in chemistry "for his work on the structure of proteins, especially that of insulin".
 In 1980, Walter Gilbert and Sanger shared half of the chemistry prize "for their contributions concerning the determination of base sequences in nucleic acids".
 The other half was awarded to Paul Berg "for his fundamental studies of the biochemistry of nucleic acids, with particular regard to recombinant DNA".
Prof John James Rickard Macleod, FRS FRSE LLD (6 September 1876 – 16 March 1935) was a Scottish biochemist and physiologist.
 He devoted his career to diverse topics in physiology and biochemistry, but was chiefly interested in carbohydrate metabolism.
 He is noted for his role in the discovery and isolation of insulin during his tenure as a lecturer at the University of Toronto, for which he and Frederick Banting received the 1923 Nobel prize in Physiology or Medicine.
 Awarding the prize to Macleod was controversial at the time, because according to Banting's version of events, Macleod's role in the discovery was negligible.
 It was not until decades after the events that an independent review acknowledged a far greater role than was attributed to him at first.