Conversation so far:
What type of work was Richard involved in?
both theoretical and experimental population genetics.
What else did he accomplish?
technology.
He was the first person to do a computer simulation of the behavior of a single gene locus (previous simulation work having been of models with multiple loci).
What else was he the first to do?
he and Ken-Ichi Kojima were the first population geneticists to give the equations for change of haplotype frequencies with interacting natural selection at two loci.

Question: What resulted from these discoveries?
Self-contained rewrite: What resulted from Richard Lewontin and Ken-Ichi Kojima's discoveries in population genetics?