Conversation so far:
What type of work was Richard involved in?
both theoretical and experimental population genetics.
What else did he accomplish?
technology.
He was the first person to do a computer simulation of the behavior of a single gene locus (previous simulation work having been of models with multiple loci).

Question: What else was he the first to do?
Self-contained rewrite: What else was Richard Lewontin the first to do, in addition to being the first person to do a computer simulation of the behavior of a single gene locus?