Conversation so far:
What type of work was Richard Lewontin involved in?
Richard Lewontin has worked in both theoretical and experimental population genetics.
What else did Richard Lewontin accomplish?
Richard Lewontin was the first person to do a computer simulation of the behavior of a single gene locus (previous simulation work having been of models with multiple loci).

Question: What else was he the first to do?
Self-contained rewrite: What else was Richard Lewontin the first to do besides being the first person to do a computer simulation of the behavior of a single gene locus?