**니그레스켄테스절**(*Utricularia* sect. *Nigrescentes*)은 통발과 통발속에 속하는 속씨식물 분류군의 하나이다. 3종의 육상 식충식물로 이루어져 있으며, 열대 아프리카와 아시아 그리고 오스트레일리아에서 발견된다. 1859년 올리버(Daniel Oliver)가 처음 유효한 분류군으로 이 절을 기재하고 출판했지만, 사용되지는 않았다. 그리고 1973년 코미야(Sadashi Komiya)에 의해 개정되었다. 1989년 테일러(Peter Taylor)가 통발속에 관한 분류학 논문에서 니그레스켄테스절을 통발아속(*Utricularia*)에 포함시켜 분류했다. 그리고 좀더 최근에는 계통 분류학 연구에 의해 니그레스켄테스절을 비발바리아아속(*Bivalvaria*)으로 옮겨 분류하고 있다.

하위 종
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* *Utricularia bracteata*
* 이삭귀개 (*Utricularia caerulea*)
* *Utricularia warburgii*

계통 분류
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다음은 통발속에 속하는 다양한 하위 아속(Subgenus)과 절(Section)의 관계를 표시한 계통도이다. 2003년 존슨 등(Jobson et al.)과 2004년과 2006년의 뭘러 등(Müller et al.)의 연구를 기초로 요약한 내용이다. 아라렐라절(*Aranella*)과 베시툴리나절(*Vesiculina*)은 다계통군으로 분지도에 별표(\*)를 표시했다. 연구가 이루어지지 않은 일부 단형절은 계통 분류학적 위치가 명확하지 않다. 통발아속(*Utricularia*)의 칸돌레아절(*Candollea*), 켈리돈절(*Chelidon*), 코리스토테테카이절(*Choristothecae*), 카미엔스키아절(*Kamienskia*), 마르티니아절(*Martinia*), 메이오눌라절(*Meionula*), 미라빌레스절(*Mirabiles*), 올리베리아절(*Oliveria*), 세티스카펠라절(*Setiscapella*), 스프루케아절(*Sprucea*), 스테예르마르키아절(*Steyermarkia*), 스틸로테카절(*Stylotheca*)과 비발바리아아속(*Bivalvaria*)의 미누타이절(*Minutae*), 폴리폼폴릭스아속(*Polypompholyx*)의 트리덴타타리아절(*Tridentaria*)은 아래에 포함되지 않았다.


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| 통발속 | 

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
| 통발아속 | 

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

|  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

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| --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

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| --- | --- |
|   | *Avesicaria* |
|   |
|   | *Vesiculina*\* |
|   |

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|   |
|   | 

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| --- | --- |
|   | *Utricularia olivacea* |
|   |
|   | *Vesiculina*\* |
|   |

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|   |

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|   |
|   | *Nelipus* |
|   |

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|   |
|   | 

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| --- | --- |
|   | *Lecticula* |
|   |
|   | 통발절 |
|   |

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|   |

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|   |
|   | 

|  |  |
| --- | --- |
|   | 오르키디오이데스절 |
|   |
|   | 폴리오사절 |
|   |

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|   |

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|   |
| 비발바리아아속 | 

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

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| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

|  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

|  |  |  |  |  |  |  |  |  |  |  |  |  |  |
| --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

|  |  |  |  |  |  |  |  |
| --- | --- | --- | --- | --- | --- | --- | --- |
|   | 

|  |  |
| --- | --- |
|   | 올리고키스타절 |
|   |
|   | *Avesicarioides* |
|   |

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|   |
|   | *Benjaminia* |
|   |

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|   |
|   | 

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| --- | --- |
|   | *Stomoisia* |
|   |
|   | *Enskide* |
|   |

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|   |

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|   |
|   | *Aranella*\* |
|   |

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|   |
|   | 

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| --- | --- |
|   | *Calpidisca* |
|   |
|   | *Lloydia* |
|   |
|   | *Australes* |
|   |

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|   |

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|   |
|   | 

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| --- | --- |
|   | 니그레스켄테스절 |
|   |
|   | *Phyllaria* |
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|   |

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|   |
|   | *Aranella*\* |
|   |

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|   |

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|   |
| 폴리폼폴릭스아속 | 

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| --- | --- |
|   | *Pleiochasia* |
|   |
|   | *Polypompholyx* |
|   |

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|   |

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|   |